treatments-xml/data/03/8B/1D/038B1D1D6447FF81FCA1EDFBD20AFD67.xml

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<mods:title id="0C8E29C16CA24786B1DE98606CF5842C">A Unitary Association-based conodont biozonation of the Smithian-Spathian boundary (Early Triassic) and associated biotic crisis from South China</mods:title>
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<mods:namePart id="FDDFB5A9AA7C8751C68F677A9A192408">Leu, Marc</mods:namePart>
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<paragraph id="8B9DAC0B6447FF9FFCA1EDFBD4F4FEE7" blockId="46.[813,1221,235,291]" box="[813,1221,235,259]" pageId="46" pageNumber="47">
<heading id="D0D51B676447FF9FFCA1EDFBD4F4FEE7" box="[813,1221,235,259]" fontSize="9" level="2" pageId="46" pageNumber="47" reason="8">
<taxonomicName id="4C22D7886447FF9FFCA1EDFBD4F4FEE7" authority="(Muller, 1956)" baseAuthorityName="MUller" baseAuthorityYear="1956" box="[813,1221,235,259]" family="Gondolellidae" genus="Guangxidella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ozarkodinida" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="bransoni">
<emphasis id="B95670196447FF9FFCA1EDFBD414FEE7" box="[813,1061,235,259]" italics="true" pageId="46" pageNumber="47">Guangxidella bransoni</emphasis>
(
<bibRefCitation id="EFB3D1FA6447FF9FFBBEEDFBD48AFEE7" author="Muller" box="[1074,1211,235,259]" firstAuthor="Muller" pageId="46" pageNumber="47" pagination="818 - 830" refId="ref45368" refString="MUller, K. J. (1956). Triassic conodonts from Nevada. Journal of Paleontology, 30, 818 - 830." type="journal article" year="1956">Müller, 1956</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C338FF806447FF81FCA1EC1BD20AFD67" lastPageId="48" lastPageNumber="49" pageId="46" pageNumber="47" type="description">
<paragraph id="8B9DAC0B6447FF9FFCA1EC1BD3FBFEC7" blockId="46.[813,1221,235,291]" box="[813,970,267,291]" pageId="46" pageNumber="47">
<figureCitation id="1319B08E6447FF9FFCA1EC1BD3A1FEC7" box="[813,912,267,291]" captionStart="Fig" captionStartId="23.[169,200,1776,1796]" captionTargetBox="[152,1438,238,1940]" captionTargetId="figure-11@23.[228,1362,254,1764]" captionTargetPageId="23" captionText="Fig. 15 Neogondolellinae and Mullerinae from Qiakong, Laren and Shanggang. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. A, D Neospathodus bevelledi n. sp.; A SHA333,PIMUZ 39176; D, SHA332, PIMUZ 39177. B, J Discretella aff. discreta (MÜller); B SHA304, PIMUZ 39104; J SHA334,PIMUZ 39105. C, L Discretella discreta (MÜller); C SHA334, PIMUZ 39107; L SHA332, PIMUZ 39108. E Guangxidella bransoni (MÜller); SHA304, PIMUZ 39139.F, H, K, M, P Discretella pseudodieneri n. sp.; F QIA124, PIMUZ 39120; H QIA124,PIMUZ SQL54990; K QIA121, PIMUZ 39121; M QIA120, PIMUZ 39122; P LAR232, PIMUZ 39123. G Discretella cf. discreta (MÜller); QIA120, PIMUZ 39109.I sp. indet.; QIA120, PIMUZ 39294. N Discretella? n. sp. B; LAR232, PIMUZ 39112.O Discretella? n. sp. C; SHA342,PIMUZ 39114" figureDoi="http://doi.org/10.5281/zenodo.12000286" httpUri="https://zenodo.org/record/12000286/files/figure.png" pageId="46" pageNumber="47">Figs. 15E</figureCitation>
; 
<figureCitation id="1319B08E6447FF9FFC16EC1BD3F7FEC7" box="[922,966,267,291]" captionStart="Fig" captionStartId="24.[169,200,1740,1760]" captionTargetBox="[153,1439,238,1800]" captionTargetId="figure-11@25.[229,1363,254,1763]" captionTargetPageId="25" captionText="Fig. 16 Neogondolellinae, Novispathodinae and Mullerinae from Shanggang and Youping cascade. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P1 elements if not specifically identified otherwise. A, F, M Discretella? n. sp. C; A YC16,PIMUZ 39115; F YC16, PIMUZ 39116; M SHA342C, PIMUZ 39117. B Discretella? n. sp. B;YC16, PIMUZ 39113. C Discretella aff.discreta (MÜller);YC22,PIMUZ 39106. D, S Discretella discreta (MÜller); D YC16, PIMUZ 39110; S SHA304C, PIMUZ 39111. E Novispathodus ex gr. abruptus (Orchard);YC41, PIMUZ 39225. G–I, K, L, N, O Discretella pseudodieneri n. sp.; G YC16, PIMUZ 39124; H YC25, PIMUZ 39125; I SHA304C,PIMUZ 39126; K YC22, PIMUZ 39127,L YC25, PIMUZ 39128; N SHA304C,PIMUZ 39129; O SHA343C,PIMUZ SQL54979. J Guangxidella bransoni (MÜller);YC16, PIMUZ 39140. P–R Neospathodus bevelledi n. sp.; P SHA 304C,PIMUZ 39178; Q SHA333C, PIMUZ 39179; R YC12, PIMUZ 39180" figureDoi="http://doi.org/10.5281/zenodo.13127506" httpUri="https://zenodo.org/record/13127506/files/figure.png" pageId="46" pageNumber="47">16E</figureCitation>
.
</paragraph>
<paragraph id="8B9DAC0B6447FF9FFCA1EC5BD3E1FE67" blockId="46.[813,1437,331,867]" pageId="46" pageNumber="47">
*1956 
<taxonomicName id="4C22D7886447FF9FFCF9EC5BD44CFE87" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[885,1149,331,355]" genus="Neoprioniodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="bransoni" status="sp. nov.">
<emphasis id="B95670196447FF9FFCF9EC5BD44CFE87" box="[885,1149,331,355]" italics="true" pageId="46" pageNumber="47">Neoprioniodus bransoni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626447FF9FFB0BEC5CD4F4FE87" box="[1159,1221,332,355]" pageId="46" pageNumber="47" rank="species">n. sp.</taxonomicNameLabel>
; Müller, p. 829, pl. 95, figs. 19–21.
</paragraph>
<paragraph id="8B9DAC0B6447FF9FFCA1EC9BD3FDFE27" blockId="46.[813,1437,331,867]" pageId="46" pageNumber="47">
1956 
<taxonomicName id="4C22D7886447FF9FFCEBEC9BD4A7FE47" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[871,1174,395,419]" genus="Neoprioniodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="bicuspidatus" status="sp. nov.">
<emphasis id="B95670196447FF9FFCEBEC9BD4A7FE47" box="[871,1174,395,419]" italics="true" pageId="46" pageNumber="47">Neoprioniodus bicuspidatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626447FF9FFB10EC9CD4E7FE47" box="[1180,1238,396,419]" pageId="46" pageNumber="47" rank="species">n. sp.</taxonomicNameLabel>
; Müller, p. 828, pl. 95, figs. 16, 17.
</paragraph>
<paragraph id="8B9DAC0B6447FF9FFCA1ECDBD3F4FDE7" blockId="46.[813,1437,331,867]" pageId="46" pageNumber="47">
1979 
<taxonomicName id="4C22D7886447FF9FFCE4ECDBD4C6FE07" authority="(Muller)" baseAuthorityName="Muller" baseAuthorityYear="1956" box="[872,1271,459,483]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="bicuspidatus">
<emphasis id="B95670196447FF9FFCE4ECDBD4A2FE07" box="[872,1171,459,483]" italics="true" pageId="46" pageNumber="47">Neospathodus bicuspidatus</emphasis>
(Müller)
</taxonomicName>
; Solien, p. 302, pl. 3, figs. 2, 3.
</paragraph>
<paragraph id="8B9DAC0B6447FF9FFCA1EF1BD394FDA7" blockId="46.[813,1437,331,867]" pageId="46" pageNumber="47">
1989 
<taxonomicName id="4C22D7886447FF9FFCE0EF1BD462FDC7" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[876,1107,523,547]" family="Prioniodontidae" genus="Ozarkodina" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="gigantea" status="sp. nov.">
<emphasis id="B95670196447FF9FFCE0EF1BD462FDC7" box="[876,1107,523,547]" italics="true" pageId="46" pageNumber="47">Ozarkodina gigantea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626447FF9FFBD1EF1CD4ADFDC7" box="[1117,1180,524,547]" pageId="46" pageNumber="47" rank="species">n. sp.</taxonomicNameLabel>
; Tang, p. 409, pl. 31, figs. 10, 14.
</paragraph>
<paragraph id="8B9DAC0B6447FF9FFCA1EF5BD3C0FD67" blockId="46.[813,1437,331,867]" pageId="46" pageNumber="47">
1991 
<taxonomicName id="4C22D7886447FF9FFCE4EF5BD477FD87" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[872,1094,587,611]" family="Gondolellidae" genus="Guangxidella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="typica" status="sp. nov.">
<emphasis id="B95670196447FF9FFCE4EF5BD477FD87" box="[872,1094,587,611]" italics="true" pageId="46" pageNumber="47">Guangxidella typica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626447FF9FFBC1EF5CD4B9FD87" box="[1101,1160,588,611]" pageId="46" pageNumber="47" rank="species">n. sp.</taxonomicNameLabel>
; Zhang &amp; Yang, p. 33, pl. 1, figs. 1a, b, 2a, b.
</paragraph>
<paragraph id="8B9DAC0B6447FF9FFCA1EF9BD4A7FD27" blockId="46.[813,1437,331,867]" pageId="46" pageNumber="47">
2009 
<taxonomicName id="4C22D7886447FF9FFCE6EF9BD4F8FD47" authority="(Muller)" baseAuthorityName="MUller" baseAuthorityYear="1956" box="[874,1225,651,675]" family="Gondolellidae" genus="Guangxidella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="bransoni">
<emphasis id="B95670196447FF9FFCE6EF9BD455FD47" box="[874,1124,651,675]" italics="true" pageId="46" pageNumber="47">Guangxidella bransoni</emphasis>
(Müller)
</taxonomicName>
; Orchard &amp; Zonneveld, p. 780, fig. 15, parts 26–28.
</paragraph>
<paragraph id="8B9DAC0B6447FF9FFCA1EFDBD380FCC7" blockId="46.[813,1437,331,867]" pageId="46" pageNumber="47">
2014 
<taxonomicName id="4C22D7886447FF9FFCE2EFDBD4E4FD07" authority="(Muller)" baseAuthorityName="MUller" baseAuthorityYear="1956" box="[878,1237,715,739]" family="Gondolellidae" genus="Guangxidella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="bransoni">
<emphasis id="B95670196447FF9FFCE2EFDBD45DFD07" box="[878,1132,715,739]" italics="true" pageId="46" pageNumber="47">Guangxidella bransoni</emphasis>
(Müller)
</taxonomicName>
; Maekawa &amp; Igo in Shigeta et al., p. 211, figs. 152.16–152.18, 153–159, 160.1–160.4
</paragraph>
<paragraph id="8B9DAC0B6447FF9FFCA1EE3BD4FAFC87" blockId="46.[813,1437,331,867]" pageId="46" pageNumber="47">
2018 
<taxonomicName id="4C22D7886447FF9FFCFBEE3BD4C1FCA7" authority="(Muller)" baseAuthorityName="MUller" baseAuthorityYear="1956" box="[887,1264,811,835]" family="Gondolellidae" genus="Guangxidella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="bransoni">
<emphasis id="B95670196447FF9FFCFBEE3BD44FFCA7" box="[887,1150,811,835]" italics="true" pageId="46" pageNumber="47">Guangxidella bransoni</emphasis>
(Müller)
</taxonomicName>
; Maekawa in Maekawa et al., p. 24, figs. 14.7–14.10.
</paragraph>
<paragraph id="8B9DAC0B6447FF9FFCA1EE9BD460FC47" blockId="46.[813,1437,907,1539]" box="[813,1105,907,931]" pageId="46" pageNumber="47">Number of specimens:&gt;30</paragraph>
<paragraph id="8B9DAC0B6447FF9FFCA1EEBCD5BAFBA7" blockId="46.[813,1437,907,1539]" pageId="46" pageNumber="47">
<emphasis id="B95670196447FF9FFCA1EEBCD381FC27" box="[813,944,940,963]" italics="true" pageId="46" pageNumber="47">Description.</emphasis>
Te segminate P1 element has variably discrete, laterally compressed denticles whose height decreases to the anterior and a very large, variably reclined, terminal cusp. Te basal margin is up-arched and the basal cavity is flat, cordiform and asymmetrical.
</paragraph>
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<emphasis id="B95670196447FF9FFCA1E95BD469FB87" box="[813,1112,1099,1123]" italics="true" pageId="46" pageNumber="47">
Remarks. 
<taxonomicName id="4C22D7886447FF9FFC17E95CD469FB87" baseAuthorityName="Muller" baseAuthorityYear="1956" box="[923,1112,1099,1123]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="bicuspidatus">Gu. bicuspidatus</taxonomicName>
</emphasis>
was differentiated from 
<taxonomicName id="4C22D7886447FF9FFAF9E95CD3BDFB67" baseAuthorityName="MUller" baseAuthorityYear="1956" family="Gondolellidae" genus="Guangxidella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="bransoni">
<emphasis id="B95670196447FF9FFAF9E95CD3BDFB67" italics="true" pageId="46" pageNumber="47">Gu. bransoni</emphasis>
</taxonomicName>
on the basis of its closer spaced denticles, the one in front of the cusp being almost as high as the latter (
<bibRefCitation id="EFB3D1FA6447FF9FFCD5E9BBD3D2FB27" author="Muller" box="[857,995,1195,1219]" firstAuthor="Muller" pageId="46" pageNumber="47" pagination="818 - 830" refId="ref45368" refString="MUller, K. J. (1956). Triassic conodonts from Nevada. Journal of Paleontology, 30, 818 - 830." type="journal article" year="1956">Müller, 1956</bibRefCitation>
). Yet, given the broad range of intraspecific variation present in large collections such as those illustrated by 
<bibRefCitation id="EFB3D1FA6447FF9FFC4FE9FBD4ACFAE7" author="Shigeta" box="[963,1181,1259,1283]" etAl="et al." firstAuthor="Shigeta" pageId="46" pageNumber="47" refId="ref46406" refString="Shigeta, Y., Komatsu, T., Maekawa, T., &amp; Huyen, D. T. (Eds.). (2014). Olenekian (Early Triassic) stratigraphy and fossil assemblages in northeastern Vietnam (No. 45). Tokyo: National Museum of Nature and Science." type="book" year="2014">Shigeta et al. (2014)</bibRefCitation>
, it is in our opinion no longer tenable to keep both species separated. Although both taxa are defined in the same paper by 
<bibRefCitation id="EFB3D1FA6447FF9FFA8AE83BD5ACFAA7" author="Muller" box="[1286,1437,1323,1347]" firstAuthor="Muller" pageId="46" pageNumber="47" pagination="818 - 830" refId="ref45368" refString="MUller, K. J. (1956). Triassic conodonts from Nevada. Journal of Paleontology, 30, 818 - 830." type="journal article" year="1956">Müller (1956)</bibRefCitation>
and the definition of 
<taxonomicName id="4C22D7886447FF9FFBA7E85CD4D8FA87" baseAuthorityName="Muller" baseAuthorityYear="1956" box="[1067,1257,1355,1379]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="bicuspidatus">
<emphasis id="B95670196447FF9FFBA7E85CD4D8FA87" box="[1067,1257,1355,1379]" italics="true" pageId="46" pageNumber="47">Gu. bicuspidatus</emphasis>
</taxonomicName>
appears before that of 
<taxonomicName id="4C22D7886447FF9FFC0BE87CD42BFA67" baseAuthorityName="MUller" baseAuthorityYear="1956" box="[903,1050,1387,1411]" family="Gondolellidae" genus="Guangxidella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="bransoni">
<emphasis id="B95670196447FF9FFC0BE87CD42BFA67" box="[903,1050,1387,1411]" italics="true" pageId="46" pageNumber="47">Gu. bransoni</emphasis>
</taxonomicName>
, we have decided to name them 
<taxonomicName id="4C22D7886447FF9FFCA1E89BD3BDFA47" baseAuthorityName="MUller" baseAuthorityYear="1956" box="[813,908,1419,1443]" family="Gondolellidae" genus="Guangxidella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="bransoni">
<emphasis id="B95670196447FF9FFCA1E89BD3BDFA47" box="[813,908,1419,1443]" italics="true" pageId="46" pageNumber="47">bransoni</emphasis>
</taxonomicName>
because the etymology of 
<taxonomicName id="4C22D7886447FF9FFB3CE89BD50BFA47" baseAuthorityName="Muller" baseAuthorityYear="1956" box="[1200,1338,1419,1443]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="bicuspidatus">
<emphasis id="B95670196447FF9FFB3CE89BD50BFA47" box="[1200,1338,1419,1443]" italics="true" pageId="46" pageNumber="47">bicuspidatus</emphasis>
</taxonomicName>
refers to a 2-cusp element, which is not typical of the revised species. Furthermore, Müller’s 
<typeStatus id="549912A96447FF9FFBD2E8DBD48FFA07" box="[1118,1214,1483,1507]" pageId="46" pageNumber="47" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C22D7886447FF9FFB6FE8DCD5ADFA07" baseAuthorityName="Muller" baseAuthorityYear="1956" box="[1251,1436,1483,1507]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="bicuspidatus">
<emphasis id="B95670196447FF9FFB6FE8DCD5ADFA07" box="[1251,1436,1483,1507]" italics="true" pageId="46" pageNumber="47">Gu. bicuspidatus</emphasis>
</taxonomicName>
is broken, whereas his 
<typeStatus id="549912A96447FF9FFBA4E8FBD4B9F9E7" box="[1064,1160,1515,1539]" pageId="46" pageNumber="47" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="4C22D7886447FF9FFB21E8FCD50AF9E7" baseAuthorityName="MUller" baseAuthorityYear="1956" box="[1197,1339,1515,1539]" family="Gondolellidae" genus="Guangxidella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="bransoni">
<emphasis id="B95670196447FF9FFB21E8FCD50AF9E7" box="[1197,1339,1515,1539]" italics="true" pageId="46" pageNumber="47">Gu. bransoni</emphasis>
</taxonomicName>
is better preserved (not broken), better illustrated (also in aboral view) and corresponds better to the revised type.
</paragraph>
<subSection id="E2ADB5E06447FF9FFF26EB8AD1B4F94A" box="[170,389,1690,1710]" pageId="46" pageNumber="47" type="multiple">
<paragraph id="8B9DAC0B6447FF9FFF26EB8AD1B4F94A" blockId="46.[170,389,1690,1710]" box="[170,389,1690,1710]" pageId="46" pageNumber="47">(See figure on next page.)</paragraph>
</subSection>
<caption id="DF5DFC836447FF9FFF25EBA5D41AF860" ID-DOI="http://doi.org/10.5281/zenodo.13127516" ID-Zenodo-Dep="13127516" httpUri="https://zenodo.org/record/13127516/files/figure.png" pageId="46" pageNumber="47" startId="46.[169,200,1717,1737]" targetBox="[152,1438,238,1801]" targetPageId="47">
<paragraph id="8B9DAC0B6447FF9FFF25EBA5D41AF860" blockId="46.[169,1414,1716,1924]" pageId="46" pageNumber="47">
<figureCitation id="1319B08E6447FF9FFF25EBA5D0D9F92D" box="[169,232,1717,1737]" captionStart="Fig" captionStartId="47.[169,200,1776,1796]" captionTargetBox="[152,1438,238,1801]" captionTargetId="figure-11@47.[228,1362,254,1763]" captionTargetPageId="47" captionText="Fig. 26 (See legend on previous page.)" figureDoi="http://doi.org/10.5281/zenodo.12000454" httpUri="https://zenodo.org/record/12000454/files/figure.png" pageId="46" pageNumber="47">
<emphasis id="B95670196447FF9FFF25EBA5D0D9F92D" bold="true" box="[169,232,1717,1737]" pageId="46" pageNumber="47">Fig. 26</emphasis>
</figureCitation>
<taxonomicName id="4C22D7886447FF9FFF7EEBA5D157F92D" authorityName="Orchard" authorityYear="2005" box="[242,358,1717,1737]" pageId="46" pageNumber="50" rank="subFamily" subFamily="Cornudininae">Cornudininae</taxonomicName>
from Qiakong, Laren and Shanggang. Magnification is ×80.The scale bar is 400 μm. All elements are considered to be P 
<subScript id="17A6AE4E6447FF9FFAECEBAFD556F929" attach="left" box="[1376,1383,1727,1741]" fontSize="6" pageId="46" pageNumber="47">1</subScript>
elements if not specifically identified otherwise. 
<emphasis id="B95670196447FF9FFDCDEBC0D25DF900" bold="true" box="[577,620,1744,1764]" pageId="46" pageNumber="47">A–W</emphasis>
<taxonomicName id="4C22D7886447FF9FFDFDEBC0D331F900" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[625,768,1744,1764]" family="Gondolellidae" genus="Urdyella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="unicorna" status="n. gen.">
<emphasis id="B95670196447FF9FFDFDEBC0D331F900" box="[625,768,1744,1764]" italics="true" pageId="46" pageNumber="47">Urdyella unicorna</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626447FF9FFC89EBC0D371F900" box="[773,832,1744,1764]" pageId="46" pageNumber="47" rank="genus">n. gen.</taxonomicNameLabel>
n. sp.; 
<emphasis id="B95670196447FF9FFCFAEBC0D3B5F900" bold="true" box="[886,900,1744,1764]" pageId="46" pageNumber="47">A</emphasis>
LAR225, PIMUZ 39345; 
<emphasis id="B95670196447FF9FFBC6EBC0D467F900" bold="true" box="[1098,1110,1744,1764]" pageId="46" pageNumber="47">B</emphasis>
LAR225, PIMUZ 39346; 
<emphasis id="B95670196447FF9FFA91EBC0D51BF900" bold="true" box="[1309,1322,1744,1764]" pageId="46" pageNumber="47">C</emphasis>
LAR229, PIMUZ 39347; 
<emphasis id="B95670196447FF9FFEAFEBFAD103F91A" bold="true" box="[291,306,1770,1790]" pageId="46" pageNumber="47">D</emphasis>
LAR227, PIMUZ 39348; 
<emphasis id="B95670196447FF9FFE74EBFAD232F91A" bold="true" box="[504,515,1770,1790]" pageId="46" pageNumber="47">E</emphasis>
LAR229,PIMUZ 39349; 
<emphasis id="B95670196447FF9FFD46EBFAD2E4F91A" bold="true" box="[714,725,1770,1790]" pageId="46" pageNumber="47">F</emphasis>
LAR229,PIMUZ 39350; 
<emphasis id="B95670196447FF9FFC10EBFAD39BF91A" bold="true" box="[924,938,1770,1790]" pageId="46" pageNumber="47">G</emphasis>
LAR225,PIMUZ 39351; 
<emphasis id="B95670196447FF9FFBFDEBFAD44EF91A" bold="true" box="[1137,1151,1770,1790]" pageId="46" pageNumber="47">H</emphasis>
LAR229,PIMUZ 39352; 
<emphasis id="B95670196447FF9FFACAEBFAD57DF91A" bold="true" box="[1350,1356,1770,1790]" pageId="46" pageNumber="47">I</emphasis>
QIA123, PIMUZ 39353; 
<emphasis id="B95670196447FF9FFEE6EA15D143F8FD" bold="true" box="[362,370,1797,1817]" pageId="46" pageNumber="47">J</emphasis>
LAR227,PIMUZ 39354; 
<emphasis id="B95670196447FF9FFDB5EA15D274F8FD" bold="true" box="[569,581,1797,1817]" pageId="46" pageNumber="47">K</emphasis>
QIA123, PIMUZ 39355; 
<emphasis id="B95670196447FF9FFC87EA15D327F8FD" bold="true" box="[779,790,1797,1817]" pageId="46" pageNumber="47">L</emphasis>
QIA123, PIMUZ 39356; 
<emphasis id="B95670196447FF9FFC57EA15D3DCF8FD" bold="true" box="[987,1005,1797,1817]" pageId="46" pageNumber="47">M</emphasis>
LAR227,PIMUZ 39357; 
<emphasis id="B95670196447FF9FFB3FEA15D4F0F8FD" bold="true" box="[1203,1217,1797,1817]" pageId="46" pageNumber="47">N</emphasis>
LAR225, PIMUZ 39358; 
<emphasis id="B95670196447FF9FFF25EA30D089F8D0" bold="true" box="[169,184,1824,1844]" pageId="46" pageNumber="47">O</emphasis>
QIA120, PIMUZ 39359; 
<emphasis id="B95670196447FF9FFEF2EA30D1BBF8D0" bold="true" box="[382,394,1824,1844]" pageId="46" pageNumber="47">P</emphasis>
LAR225, PIMUZ 39360; 
<emphasis id="B95670196447FF9FFDDCEA30D26EF8D0" bold="true" box="[592,607,1824,1844]" pageId="46" pageNumber="47">Q</emphasis>
QIA121, PIMUZ 39361; 
<emphasis id="B95670196447FF9FFCA9EA30D300F8D0" bold="true" box="[805,817,1824,1844]" pageId="46" pageNumber="47">R</emphasis>
LAR232, PIMUZ 39362; 
<emphasis id="B95670196447FF9FFC74EA30D432F8D0" bold="true" box="[1016,1027,1824,1844]" pageId="46" pageNumber="47">S</emphasis>
LAR232, PIMUZ 39363; 
<emphasis id="B95670196447FF9FFB46EA30D4E4F8D0" bold="true" box="[1226,1237,1824,1844]" pageId="46" pageNumber="47">T</emphasis>
QIA124,PIMUZ 39364; 
<emphasis id="B95670196447FF9FFF6AEA2AD0C5F8AA" bold="true" box="[230,244,1850,1870]" pageId="46" pageNumber="47">U</emphasis>
LAR232, PIMUZ 39365; 
<emphasis id="B95670196447FF9FFE37EA2AD1F9F8AA" bold="true" box="[443,456,1850,1870]" pageId="46" pageNumber="47">
<collectionCode id="ED3334CE6447FF9FFE37EA2AD1F9F8AA" box="[443,456,1850,1870]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="46" pageNumber="47" type="Museum">V</collectionCode>
</emphasis>
QIA123, PIMUZ 39366; 
<emphasis id="B95670196447FF9FFD01EA2AD291F8AA" bold="true" box="[653,672,1850,1870]" pageId="46" pageNumber="47">W</emphasis>
SHA341, PIMUZ 39367. 
<emphasis id="B95670196447FF9FFCE5EA2AD3A6F8AA" bold="true" box="[873,919,1850,1870]" pageId="46" pageNumber="47">X-AA</emphasis>
<taxonomicName id="4C22D7886447FF9FFC10EA2AD413F8AA" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[924,1058,1850,1870]" family="Gondolellidae" genus="Urdyella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="46" pageNumber="47" phylum="Chordata" rank="species" species="tridenta" status="n. gen.">
<emphasis id="B95670196447FF9FFC10EA2AD413F8AA" box="[924,1058,1850,1870]" italics="true" pageId="46" pageNumber="47">Urdyella tridenta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626447FF9FFBABEA2AD453F8AA" box="[1063,1122,1850,1870]" pageId="46" pageNumber="47" rank="genus">n. gen.</taxonomicNameLabel>
n. sp.; 
<emphasis id="B95670196447FF9FFB14EA2AD494F8AA" bold="true" box="[1176,1189,1850,1870]" pageId="46" pageNumber="47">X</emphasis>
LAR227, PIMUZ 39341; 
<emphasis id="B95670196447FF9FFAE0EA2AD549F8AA" bold="true" box="[1388,1400,1850,1870]" pageId="46" pageNumber="47">Y</emphasis>
LAR227, PIMUZ 39342; 
<emphasis id="B95670196447FF9FFEE7EA45D146F88D" bold="true" box="[363,375,1877,1897]" pageId="46" pageNumber="47">Z</emphasis>
LAR227, PIMUZ 39343; 
<emphasis id="B95670196447FF9FFDB2EA45D268F88D" bold="true" box="[574,601,1877,1897]" pageId="46" pageNumber="47">AA</emphasis>
LAR227, PIMUZ 39344. 
<emphasis id="B95670196447FF9FFCACEA45D30BF88D" bold="true" box="[800,826,1877,1897]" pageId="46" pageNumber="47">AB</emphasis>
, 
<emphasis id="B95670196447FF9FFCCFEA45D36CF88D" bold="true" box="[835,861,1877,1897]" pageId="46" pageNumber="47">AC</emphasis>
<taxonomicName id="4C22D7886447FF9FFCEDEA45D384F88D" authorityName="Orchard" authorityYear="2005" box="[865,949,1877,1897]" family="Gondolellidae" genus="Discretella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="46" pageNumber="47" phylum="Chordata" rank="genus">
<emphasis id="B95670196447FF9FFCEDEA45D384F88D" box="[865,949,1877,1897]" italics="true" pageId="46" pageNumber="47">Discretella</emphasis>
</taxonomicName>
? n. sp. D; 
<emphasis id="B95670196447FF9FFB8BEA45D410F88D" bold="true" box="[1031,1057,1877,1897]" pageId="46" pageNumber="47">AB</emphasis>
QIA124, PIMUZ 39118; 
<emphasis id="B95670196447FF9FFB6BEA45D530F88D" bold="true" box="[1255,1281,1877,1897]" pageId="46" pageNumber="47">AC</emphasis>
QIA123, PIMUZ 39119. 
<emphasis id="B95670196447FF9FFF6AEA60D133F860" bold="true" box="[230,258,1904,1924]" pageId="46" pageNumber="47">AD</emphasis>
<taxonomicName id="4C22D7886447FF9FFE8AEA60D18CF860" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[262,445,1904,1924]" genus="Neostrachanognathus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="46" pageNumber="47" phylum="Chordata" rank="genus" status="n. sp.">
<emphasis id="B95670196447FF9FFE8AEA60D18CF860" box="[262,445,1904,1924]" italics="true" pageId="46" pageNumber="47">Neostrachanognathus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626447FF9FFE4EEA60D1DEF860" box="[450,495,1904,1924]" pageId="46" pageNumber="47" rank="species">n. sp.</taxonomicNameLabel>
A; LAR232, PIMUZ 39183. 
<emphasis id="B95670196447FF9FFD45EA60D2D3F860" bold="true" box="[713,738,1904,1924]" pageId="46" pageNumber="47">AE</emphasis>
<taxonomicName id="4C22D7886447FF9FFD6AEA60D318F860" authority="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand, 2022" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[742,809,1904,1924]" family="Gondolellidae" genus="Urdyella" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="46" pageNumber="47" phylum="Chordata" rank="genus" status="n. sp.">
<emphasis id="B95670196447FF9FFD6AEA60D318F860" box="[742,809,1904,1924]" italics="true" pageId="46" pageNumber="47">Urdyella</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A265CD626447FF9FFCA1EA60D36BF860" box="[813,858,1904,1924]" pageId="46" pageNumber="47" rank="species">n. sp.</taxonomicNameLabel>
A; LAR232, PIMUZ 39340
</paragraph>
</caption>
<caption id="DF5DFC836446FF9EFF25EBE0D1C8F8E0" ID-DOI="http://doi.org/10.5281/zenodo.12000454" ID-Zenodo-Dep="12000454" box="[169,505,1776,1796]" httpUri="https://zenodo.org/record/12000454/files/figure.png" pageId="47" pageNumber="48" startId="47.[169,200,1776,1796]" targetBox="[152,1438,238,1801]" targetPageId="47" targetType="figure">
<paragraph id="8B9DAC0B6446FF9EFF25EBE0D1C8F8E0" blockId="47.[169,505,1776,1796]" box="[169,505,1776,1796]" pageId="47" pageNumber="48">
<emphasis id="B95670196446FF9EFF25EBE0D0D6F8E0" bold="true" box="[169,231,1776,1796]" pageId="47" pageNumber="48">Fig. 26</emphasis>
(See legend on previous page.)
</paragraph>
</caption>
<paragraph id="8B9DAC0B6459FF81FF21EC3CD20AFD67" blockId="48.[151,776,235,644]" pageId="48" pageNumber="49">
<emphasis id="B95670196459FF81FF21EC3CD119FEA7" box="[173,296,300,323]" italics="true" pageId="48" pageNumber="49">Occurrence</emphasis>
. 
<collectingCountry id="F335EC9B6459FF81FEBAEC3BD15DFEA7" box="[310,364,299,323]" name="United States of America" pageId="48" pageNumber="49">USA</collectingCountry>
: Dinner Springs Canyon, Lower Triassic 
<taxonomicName id="4C22D7886459FF81FF59EC5BD163FE87" box="[213,338,331,355]" class="Cephalopoda" family="Prionitidae" genus="Meekoceras" kingdom="Animalia" pageId="48" pageNumber="49" phylum="Mollusca" rank="genus">
<emphasis id="B95670196459FF81FF59EC5BD163FE87" box="[213,338,331,355]" italics="true" pageId="48" pageNumber="49">Meekoceras</emphasis>
</taxonomicName>
beds, Crittenden Ranch, Elko County, 
<collectingRegion id="49E662E96459FF81FF1BEC7BD0DBFE67" box="[151,234,363,387]" country="United States of America" name="Nevada" pageId="48" pageNumber="49">Nevada</collectingRegion>
(
<bibRefCitation id="EFB3D1FA6459FF81FE8EEC7BD1A4FE67" author="Muller" box="[258,405,363,387]" firstAuthor="Muller" pageId="48" pageNumber="49" pagination="818 - 830" refId="ref45368" refString="MUller, K. J. (1956). Triassic conodonts from Nevada. Journal of Paleontology, 30, 818 - 830." type="journal article" year="1956">Müller, 1956</bibRefCitation>
); Taynes Formation (
<taxonomicName id="4C22D7886459FF81FD26EC7BD178FE47" authority="Zone" authorityName="Zone" genus="Parachirognathus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="49" phylum="Chordata" rank="genus">
<emphasis id="B95670196459FF81FD26EC7BD135FE47" italics="true" pageId="48" pageNumber="49">Parachirognathus</emphasis>
Zone
</taxonomicName>
, 
<collectingRegion id="49E662E96459FF81FED4EC9BD1A1FE47" box="[344,400,395,419]" country="United States of America" name="Utah" pageId="48" pageNumber="49">Utah</collectingRegion>
, (
<bibRefCitation id="EFB3D1FA6459FF81FE29EC9BD219FE47" author="Solien" box="[421,552,395,419]" firstAuthor="Solien" pageId="48" pageNumber="49" refId="ref46622" refString="Solien, M. A. (1979). Conodont biostratigraphy of the Lower Triassic Thaynes Formation, Utah. Journal of Paleontology, 276 - 306. Retrieved from https: // www. jstor. org / stable / 1303871" type="url" year="1979">Solien 1979</bibRefCitation>
). 
<collectingRegion id="49E662E96459FF81FDCDEC9BD332FE47" box="[577,771,395,419]" country="Canada" name="British Columbia" pageId="48" pageNumber="49">British Columbia</collectingRegion>
, 
<collectingCountry id="F335EC9B6459FF81FF1BECBBD0DBFE27" box="[151,234,427,451]" name="Canada" pageId="48" pageNumber="49">Canada</collectingCountry>
(
<bibRefCitation id="EFB3D1FA6459FF81FF76ECBBD207FE27" author="Orchard &amp; Zonneveld" box="[250,566,427,451]" firstAuthor="Orchard" pageId="48" pageNumber="49" refId="ref45906" refString="Orchard, M. J., &amp; Zonneveld, J. P. (2009). The Lower Triassic Sulphur Moun-" type="book" year="2009">Orchard &amp; Zonneveld, 2009</bibRefCitation>
). Taho Formation, 
<taxonomicName id="4C22D7886459FF81FF1BECDBD108FE07" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[151,313,459,483]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="49" phylum="Chordata" rank="genus">
<emphasis id="B95670196459FF81FF1BECDBD108FE07" box="[151,313,459,483]" italics="true" pageId="48" pageNumber="49">Novispathodus</emphasis>
</taxonomicName>
ex. gr. 
<taxonomicName id="4C22D7886459FF81FE14ECDCD209FE07" authority="Zone" authorityName="Zone" box="[408,568,459,483]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B95670196459FF81FE14ECDCD1C0FE07" box="[408,497,460,483]" italics="true" pageId="48" pageNumber="49">waageni</emphasis>
Zone
</taxonomicName>
) Southwest 
<collectingCountry id="F335EC9B6459FF81FD47ECDCD336FE07" box="[715,775,460,483]" name="Japan" pageId="48" pageNumber="49">Japan</collectingCountry>
(
<bibRefCitation id="EFB3D1FA6459FF81FF12ECFBD1A6FDE7" author="Maekawa" box="[158,407,491,515]" etAl="et al." firstAuthor="Maekawa" pageId="48" pageNumber="49" pagination="1 - 62" refId="ref44774" refString="Maekawa, T., Komatsu, T., &amp; Koike, T. (2018). Early Triassic conodonts from the Tahogawa Member of the Taho Formation, Ehime Prefecture, Southwest Japan. Paleontological Research, 22 (s 1), 1 - 62. https: // doi. org / 10. 2517 / 2018 PR 001" type="journal article" year="2018">Maekawa et al., 2018</bibRefCitation>
). Nanpanjiang basin: Bac Tuy Formation, 
<taxonomicName id="4C22D7886459FF81FE95EF1CD14CFDC7" authorityName="Hyatt &amp; Smith" authorityYear="1905" box="[281,381,524,547]" class="Cephalopoda" family="Melagathiceratidae" genus="Owenites" kingdom="Animalia" pageId="48" pageNumber="49" phylum="Mollusca" rank="genus">
<emphasis id="B95670196459FF81FE95EF1CD14CFDC7" box="[281,381,524,547]" italics="true" pageId="48" pageNumber="49">Owenites</emphasis>
</taxonomicName>
beds within the 
<taxonomicName id="4C22D7886459FF81FDB1EF1BD2EEFDC7" authorityName="Leu &amp; Bucher &amp; Vennemann &amp; Bagherpour &amp; Ji &amp; Brosse &amp; Goudemand" authorityYear="2022" box="[573,735,523,547]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="49" phylum="Chordata" rank="genus">
<emphasis id="B95670196459FF81FDB1EF1BD2EEFDC7" box="[573,735,523,547]" italics="true" pageId="48" pageNumber="49">Novispathodus</emphasis>
</taxonomicName>
ex. gr. 
<taxonomicName id="4C22D7886459FF81FF36EF3CD167FDA7" authority="Zone" authorityName="Zone" box="[186,342,556,579]" family="Gondolellidae" genus="Novispathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B95670196459FF81FF36EF3CD122FDA7" box="[186,275,556,579]" italics="true" pageId="48" pageNumber="49">waageni</emphasis>
Zone
</taxonomicName>
, North-eastern 
<collectingCountry id="F335EC9B6459FF81FD8AEF3BD252FDA7" box="[518,611,555,579]" name="Vietnam" pageId="48" pageNumber="49">Vietnam</collectingCountry>
(
<bibRefCitation id="EFB3D1FA6459FF81FDFEEF3BD0FAFD87" author="Shigeta" etAl="et al." firstAuthor="Shigeta" pageId="48" pageNumber="49" refId="ref46406" refString="Shigeta, Y., Komatsu, T., Maekawa, T., &amp; Huyen, D. T. (Eds.). (2014). Olenekian (Early Triassic) stratigraphy and fossil assemblages in northeastern Vietnam (No. 45). Tokyo: National Museum of Nature and Science." type="book" year="2014">Shigeta et al., 2014</bibRefCitation>
). Luolou Formation, 
<taxonomicName id="4C22D7886459FF81FE4CEF5BD332FD87" authority="Zone" authorityName="Zone" box="[448,771,587,611]" family="Gondolellidae" genus="Neospathodus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="48" pageNumber="49" phylum="Chordata" rank="species" species="waageni">
<emphasis id="B95670196459FF81FE4CEF5BD28CFD87" box="[448,701,587,611]" italics="true" pageId="48" pageNumber="49">Neospathodus waageni</emphasis>
Zone
</taxonomicName>
, 
<collectingRegion id="49E662E96459FF81FF1BEF7BD0C7FD67" box="[151,246,619,643]" country="China" name="Guangxi" pageId="48" pageNumber="49">Guangxi</collectingRegion>
, 
<collectingCountry id="F335EC9B6459FF81FE8CEF7BD173FD67" box="[256,322,619,643]" name="China" pageId="48" pageNumber="49">China</collectingCountry>
(
<bibRefCitation id="EFB3D1FA6459FF81FEDCEF7BD21CFD67" author="Zhang &amp; Yang" box="[336,557,619,644]" firstAuthor="Zhang" pageId="48" pageNumber="49" pagination="17 - 47" refId="ref48867" refString="Zhang, S., &amp; Yang, Z. (1991). On multielement taxonomy of the Early Triassic conodonts. Stratigraphy and Paleontology of China, 1, 17 - 47." type="journal article" year="1991">Zhang &amp; Yang, 1991</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>