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<document id="B1A48ECF8AC2E49763F18A5DE2982E53" ID-DOI="10.5281/zenodo.282373" ID-GBIF-Dataset="0a9d9c77-1822-4abd-9b6a-a37e0aa2ae14" ID-ISSN="1175-5326" ID-Zenodo-Dep="282373" ID-ZooBank="3C56DA61-02AD-4FB5-88E5-F98249A7EB45" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460528108206" checkinUser="plazi" docAuthor="Solís, Ángel &amp; Kohlmann, Bert" docDate="2012" docId="03EB87CB824E7820FF50FB69FD97FAAB" docLanguage="en" docName="zt03482p032.pdf" docOrigin="Zootaxa 3482" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Ateuchini" docType="treatment" docVersion="7" lastPageNumber="5" masterDocId="FFD2FFB3824D7824FFC7FF8CFFCAFFB6" masterDocTitle="Checklist and distribution atlas of the Scarabaeinae (Coleoptera: Scarabaeidae) of Costa Rica" masterLastPageNumber="32" masterPageNumber="1" pageNumber="4" updateTime="1698315427176" updateUser="plazi">
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<mods:title id="9037EF77F57FC25A0B163F5C8075F241">Checklist and distribution atlas of the Scarabaeinae (Coleoptera: Scarabaeidae) of Costa Rica</mods:title>
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<mods:namePart id="6A16F106307BBE912838B5B529EDB9A1">Solís, Ángel</mods:namePart>
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<mods:namePart id="4B0EC1C1FCC5D72D595FFCEB1772F451">Kohlmann, Bert</mods:namePart>
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<mods:date id="5CDBFC9497F93D8092592BCF966C41C0">2012</mods:date>
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<treatment id="03EB87CB824E7820FF50FB69FD97FAAB" ID-DOI="http://doi.org/10.5281/zenodo.5691361" ID-GBIF-Taxon="119526506" ID-Zenodo-Dep="5691361" LSID="urn:lsid:plazi:treatment:03EB87CB824E7820FF50FB69FD97FAAB" httpUri="http://treatment.plazi.org/id/03EB87CB824E7820FF50FB69FD97FAAB" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<subSubSection id="C3586556824E7827FF50FB69FE0AFA13" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="8BFD36DD824E7827FF50FB69FEF4FB49" blockId="3.[151,318,1253,1279]" box="[151,318,1253,1279]" pageId="3" pageNumber="4">
<heading id="D0B581B1824E7827FF50FB69FEF4FB49" bold="true" box="[151,318,1253,1279]" fontSize="11" level="1" pageId="3" pageNumber="4" reason="1">
<taxonomicName id="4C424D5E824E7827FF50FB69FEF4FB49" ID-CoL="KLZ" box="[151,318,1253,1279]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="tribe" tribe="Ateuchini">
<emphasis id="B936EACF824E7827FF50FB69FEF4FB49" bold="true" box="[151,318,1253,1279]" pageId="3" pageNumber="4">ATEUCHINI</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BFD36DD824E7827FF50FAA5FE56FAD4" blockId="3.[151,445,1321,1379]" pageId="3" pageNumber="4">
<taxonomicName id="4C424D5E824E7827FF50FAA5FE77FAF7" authority="Bates, 1887" authorityName="Bates" authorityYear="1887" box="[151,445,1321,1346]" class="Insecta" family="Scarabaeidae" genus="Agamopus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B936EACF824E7827FF50FAA5FEF9FAF4" bold="true" box="[151,307,1321,1346]" italics="true" pageId="3" pageNumber="4">AGAMOPUS</emphasis>
Bates, 1887
</taxonomicName>
<taxonomicName id="4C424D5E824E7827FF50FAC7FE56FAD4" authority="Bates, 1887" authorityName="Bates" authorityYear="1887" box="[151,412,1354,1379]" class="Insecta" family="Scarabaeidae" genus="Agamopus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="lampros">
<emphasis id="B936EACF824E7827FF50FAC7FEDBFAD5" box="[151,273,1355,1379]" italics="true" pageId="3" pageNumber="4">A. lampros</emphasis>
Bates, 1887
</taxonomicName>
</paragraph>
<paragraph id="8BFD36DD824E7827FF50FA01FE0AFA13" blockId="3.[151,647,1421,1912]" box="[151,448,1421,1446]" pageId="3" pageNumber="4">
<taxonomicName id="4C424D5E824E7827FF50FA01FE0AFA13" authority="Weber, 1801" authorityName="Weber" authorityYear="1801" box="[151,448,1421,1446]" class="Insecta" family="Scarabaeidae" genus="Ateuchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B936EACF824E7827FF50FA01FEE1FA10" bold="true" box="[151,299,1421,1446]" italics="true" pageId="3" pageNumber="4">ATEUCHUS</emphasis>
Weber, 1801
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C3586556824E7820FF00FA22FD97FAAB" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="reference_group">
<paragraph id="8BFD36DD824E7827FF00FA22FE3FFA51" blockId="3.[151,647,1421,1912]" pageId="3" pageNumber="4">
<taxonomicName id="4C424D5E824E7827FF00FA22FD4DFA70" authority="LePeletier &amp; Serville, 1828" authorityName="LePeletier &amp; Serville" authorityYear="1828" box="[199,647,1454,1478]" class="Insecta" family="Scarabaeidae" genus="Choeridium" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B936EACF824E7827FF00FA22FE87FA70" box="[199,333,1454,1478]" italics="true" pageId="3" pageNumber="4">Choeridium</emphasis>
LePeletier &amp; Serville, 1828
</taxonomicName>
<taxonomicName id="4C424D5E824E7827FF50FA5CFE3FFA51" ID-CoL="7ZLWQ" authority="Harold, 1868" authorityName="Harold" authorityYear="1868" box="[151,501,1487,1512]" class="Insecta" family="Scarabaeidae" genus="Ateuchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="aeneomicans">
<emphasis id="B936EACF824E7827FF50FA5CFE82FA5E" box="[151,328,1488,1512]" italics="true" pageId="3" pageNumber="4">A. aeneomicans</emphasis>
(
<bibRefCitation id="EFD34B2C824E7827FE90FA43FE27FA51" author="Harold" box="[343,493,1487,1511]" pageId="3" pageNumber="4" refString="Harold, E. von (1868) Monographie der Gattung Canthon. Berliner Entomologische Zeitung, 12, 1 - 141." type="journal article" year="1868">Harold, 1868</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="8BFD36DD824E7827FF50FA7DFE0FF99C" blockId="3.[151,647,1421,1912]" pageId="3" pageNumber="4">
<taxonomicName id="4C424D5E824E7827FF50FA7DFDF6F9BF" ID-CoL="7ZLWS" authority="Kohlmann &amp; Solis, 2012" authorityName="Kohlmann &amp; Solis" authorityYear="2012" box="[151,572,1521,1545]" class="Insecta" family="Scarabaeidae" genus="Ateuchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="alutacius">
<emphasis id="B936EACF824E7827FF50FA7DFED7F9BF" box="[151,285,1521,1545]" italics="true" pageId="3" pageNumber="4">A. alutacius</emphasis>
<bibRefCitation id="EFD34B2C824E7827FEE3FA7DFDF6F9BF" author="Kohlmann" box="[292,572,1521,1545]" pageId="3" pageNumber="4" refString="Kohlmann, B. &amp; Solis, A. (2012) New species and revalidations of scarab beetles (Coleoptera: Geotrupidae: Athyreini and Coleoptera: Scarabaeidae: Scarabaeinae) from Costa Rica and Panama. Zootaxa, 3193, 28 - 52." type="journal article" year="2012">Kohlmann &amp; Solís, 2012</bibRefCitation>
</taxonomicName>
<taxonomicName id="4C424D5E824E7827FF50F99FFE0FF99C" ID-CoL="3YZQX" authority="Harold, 1868" authorityName="Harold" authorityYear="1868" box="[151,453,1554,1579]" class="Insecta" family="Scarabaeidae" genus="Megathoposoma" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="candezei">
<emphasis id="B936EACF824E7827FF50F99FFED2F99D" box="[151,280,1555,1579]" italics="true" pageId="3" pageNumber="4">A. candezei</emphasis>
(
<bibRefCitation id="EFD34B2C824E7827FEE1F99EFE77F99C" author="Harold" box="[294,445,1554,1578]" pageId="3" pageNumber="4" refString="Harold, E. von (1868) Monographie der Gattung Canthon. Berliner Entomologische Zeitung, 12, 1 - 141." type="journal article" year="1868">Harold, 1868</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="8BFD36DD824E7827FF00F9B8FE76F938" blockId="3.[151,647,1421,1912]" pageId="3" pageNumber="4">
<taxonomicName id="4C424D5E824E7827FF00F9B8FD8DF9FA" authority="Bates, 1887" authorityName="Bates" authorityYear="1887" box="[199,583,1588,1612]" class="Insecta" family="Scarabaeidae" genus="Choeridium" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="poropyge">
<emphasis id="B936EACF824E7827FF00F9B8FE77F9FA" box="[199,445,1588,1612]" italics="true" pageId="3" pageNumber="4">Choeridium poropyge</emphasis>
Bates, 1887
</taxonomicName>
<taxonomicName id="4C424D5E824E7827FF50F9D9FD81F9DB" ID-CoL="9X5PQ" authority="Kohlmann &amp; Solis, 2009" authorityName="Kohlmann &amp; Solis" authorityYear="2009" box="[151,587,1621,1645]" class="Insecta" family="Scarabaeidae" genus="Ateuchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="earthorum">
<emphasis id="B936EACF824E7827FF50F9D9FEE6F9DB" box="[151,300,1621,1645]" italics="true" pageId="3" pageNumber="4">A. earthorum</emphasis>
Kohlmann &amp; Solís, 2009
</taxonomicName>
<taxonomicName id="4C424D5E824E7827FF50F9FAFE76F938" ID-CoL="7ZLXN" authority="Kohlmann, 1997" authorityName="Kohlmann" authorityYear="1997" box="[151,444,1654,1678]" class="Insecta" family="Scarabaeidae" genus="Ateuchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="fetteri">
<emphasis id="B936EACF824E7827FF50F9FAFF33F938" box="[151,249,1654,1678]" italics="true" pageId="3" pageNumber="4">A. fetteri</emphasis>
<bibRefCitation id="EFD34B2C824E7827FEC7F9FAFE76F938" author="Kohlmann" box="[256,444,1654,1678]" pageId="3" pageNumber="4" refString="Kohlmann, B. &amp; Solis, A. (1997) El genero Dichotomius (Coleoptera: Scarabaeidae) en Costa Rica. Giornale Italiano di Entomologia, 8, 343 - 382." type="journal article" year="1997">Kohlmann, 1997</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BFD36DD824E7827FF50F914FE7DF919" blockId="3.[151,647,1421,1912]" box="[151,439,1687,1712]" pageId="3" pageNumber="4">
<taxonomicName id="4C424D5E824E7827FF50F914FE7DF919" ID-CoL="7ZLXW" authority="Kohlmann, 1997" authorityName="Kohlmann" authorityYear="1997" box="[151,439,1687,1712]" class="Insecta" family="Scarabaeidae" genus="Ateuchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="ginae">
<emphasis id="B936EACF824E7827FF50F914FF3EF906" box="[151,244,1688,1712]" italics="true" pageId="3" pageNumber="4">A. ginae</emphasis>
<bibRefCitation id="EFD34B2C824E7827FF3CF91BFE7DF919" author="Kohlmann" box="[251,439,1687,1711]" pageId="3" pageNumber="4" refString="Kohlmann, B. &amp; Solis, A. (1997) El genero Dichotomius (Coleoptera: Scarabaeidae) en Costa Rica. Giornale Italiano di Entomologia, 8, 343 - 382." type="journal article" year="1997">Kohlmann, 1997</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BFD36DD824E7827FF50F935FE25F967" blockId="3.[151,647,1421,1912]" box="[151,495,1721,1745]" pageId="3" pageNumber="4">
<taxonomicName id="4C424D5E824E7827FF50F935FE25F967" ID-CoL="7ZLY4" authority="Kohlmann, 1997" authorityName="Kohlmann" authorityYear="1997" box="[151,495,1721,1745]" class="Insecta" family="Scarabaeidae" genus="Ateuchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="hendrichsi">
<emphasis id="B936EACF824E7827FF50F935FEE6F967" box="[151,300,1721,1745]" italics="true" pageId="3" pageNumber="4">A. hendrichsi</emphasis>
<bibRefCitation id="EFD34B2C824E7827FEF4F935FE25F967" author="Kohlmann" box="[307,495,1721,1745]" pageId="3" pageNumber="4" refString="Kohlmann, B. &amp; Solis, A. (1997) El genero Dichotomius (Coleoptera: Scarabaeidae) en Costa Rica. Giornale Italiano di Entomologia, 8, 343 - 382." type="journal article" year="1997">Kohlmann, 1997</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BFD36DD824E7827FF50F957FE12F944" blockId="3.[151,647,1421,1912]" box="[151,472,1754,1779]" pageId="3" pageNumber="4">
<taxonomicName id="4C424D5E824E7827FF50F957FE12F944" ID-CoL="7ZLY9" authority="Kohlmann, 1997" authorityName="Kohlmann" authorityYear="1997" box="[151,472,1754,1779]" class="Insecta" family="Scarabaeidae" genus="Ateuchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="howdeni">
<emphasis id="B936EACF824E7827FF50F957FEDFF945" box="[151,277,1755,1779]" italics="true" pageId="3" pageNumber="4">A. howdeni</emphasis>
<bibRefCitation id="EFD34B2C824E7827FEDBF956FE12F944" author="Kohlmann" box="[284,472,1754,1778]" pageId="3" pageNumber="4" refString="Kohlmann, B. &amp; Solis, A. (1997) El genero Dichotomius (Coleoptera: Scarabaeidae) en Costa Rica. Giornale Italiano di Entomologia, 8, 343 - 382." type="journal article" year="1997">Kohlmann, 1997</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BFD36DD824E7827FF50F970FD4DF8A2" blockId="3.[151,647,1421,1912]" box="[151,647,1788,1812]" pageId="3" pageNumber="4">
<taxonomicName id="4C424D5E824E7827FF50F970FEE0F8A2" ID-CoL="7ZLZF" box="[151,298,1788,1812]" class="Insecta" family="Scarabaeidae" genus="Ateuchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="rodriguezi">
<emphasis id="B936EACF824E7827FF50F970FEE0F8A2" box="[151,298,1788,1812]" italics="true" pageId="3" pageNumber="4">A. rodriguezi</emphasis>
</taxonomicName>
(Preudhomme de Borre, 1886)
</paragraph>
<paragraph id="8BFD36DD824E7827FF00F891FE7EF8E0" blockId="3.[151,647,1421,1912]" pageId="3" pageNumber="4">
<taxonomicName id="4C424D5E824E7827FF00F891FD9CF883" authority="Bates, 1887" authorityName="Bates" authorityYear="1887" box="[199,598,1821,1845]" class="Insecta" family="Scarabaeidae" genus="Choeridium" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="ampliatum">
<emphasis id="B936EACF824E7827FF00F891FE06F883" box="[199,460,1821,1845]" italics="true" pageId="3" pageNumber="4">Choeridium ampliatum</emphasis>
Bates, 1887
</taxonomicName>
<taxonomicName id="4C424D5E824E7827FF50F8B2FE7EF8E0" ID-CoL="ZV65" authority="Kohlmann, 1997" authorityName="Kohlmann" authorityYear="1997" box="[151,436,1854,1878]" class="Insecta" family="Scarabaeidae" genus="Cryptocanthon" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="solisi">
<emphasis id="B936EACF824E7827FF50F8B2FF3BF8E0" box="[151,241,1854,1878]" italics="true" pageId="3" pageNumber="4">A. solisi</emphasis>
<bibRefCitation id="EFD34B2C824E7827FF3FF8B2FE7EF8E0" author="Kohlmann" box="[248,436,1854,1878]" pageId="3" pageNumber="4" refString="Kohlmann, B. &amp; Solis, A. (1997) El genero Dichotomius (Coleoptera: Scarabaeidae) en Costa Rica. Giornale Italiano di Entomologia, 8, 343 - 382." type="journal article" year="1997">Kohlmann, 1997</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BFD36DD824E7827FF50F8ECFE2BF8C1" blockId="3.[151,647,1421,1912]" box="[151,481,1887,1912]" pageId="3" pageNumber="4">
<taxonomicName id="4C424D5E824E7827FF50F8ECFE2BF8C1" ID-CoL="7ZM25" authority="Kohlmann, 1997" authorityName="Kohlmann" authorityYear="1997" box="[151,481,1887,1912]" class="Insecta" family="Scarabaeidae" genus="Ateuchus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="zoebischi">
<emphasis id="B936EACF824E7827FF50F8ECFED5F8CE" box="[151,287,1888,1912]" italics="true" pageId="3" pageNumber="4">A. zoebischi</emphasis>
<bibRefCitation id="EFD34B2C824E7827FEE2F8D3FE2BF8C1" author="Kohlmann" box="[293,481,1887,1911]" pageId="3" pageNumber="4" refString="Kohlmann, B. &amp; Solis, A. (1997) El genero Dichotomius (Coleoptera: Scarabaeidae) en Costa Rica. Giornale Italiano di Entomologia, 8, 343 - 382." type="journal article" year="1997">Kohlmann, 1997</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BFD36DD824E7820FF50F82EFDE9FF67" blockId="3.[151,451,1954,2012]" lastBlockId="4.[151,620,151,209]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<taxonomicName id="4C424D5E824E7827FF50F82EFE09F80C" ID-CoL="38ZW" authority="Harold, 1869" authorityName="Harold" authorityYear="1869" box="[151,451,1954,1979]" class="Insecta" family="Scarabaeidae" genus="Bdelyrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B936EACF824E7827FF50F82EFEE2F80D" bold="true" box="[151,296,1954,1979]" italics="true" pageId="3" pageNumber="4">BDELYRUS</emphasis>
Harold, 1869
</taxonomicName>
<taxonomicName id="4C424D5E824E7827FF50F849FE7DF86A" ID-CoL="5WFLW" authority="Bates, 1887" authorityName="Bates" authorityYear="1887" box="[151,439,1988,2012]" class="Insecta" family="Scarabaeidae" genus="Bdelyrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="seminudus">
<emphasis id="B936EACF824E7827FF50F849FEE7F86A" box="[151,301,1988,2012]" italics="true" pageId="3" pageNumber="4">B. seminudus</emphasis>
Bates, 1887
</taxonomicName>
<taxonomicName id="4C424D5E82497820FF50FF1BFE4FFF06" ID-CoL="3CQT" box="[151,389,151,176]" class="Insecta" family="Scarabaeidae" genus="Bradypodidium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B936EACF82497820FF50FF1BFE4FFF06" bold="true" box="[151,389,151,176]" italics="true" pageId="4" pageNumber="5">BRADYPODIDIUM</emphasis>
</taxonomicName>
Vaz-de-Mello, 2008
<taxonomicName id="4C424D5E82497820FF50FF36FDE9FF67" ID-CoL="5X3QX" authority="Boucomont, 1928" authorityName="Boucomont" authorityYear="1928" box="[151,547,185,209]" class="Insecta" family="Scarabaeidae" genus="Bradypodidium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bradyporum">
<emphasis id="B936EACF82497820FF50FF36FE8BFF67" box="[151,321,185,209]" italics="true" pageId="4" pageNumber="5">B. bradyporum</emphasis>
(Boucomont, 1928)
</taxonomicName>
</paragraph>
<paragraph id="8BFD36DD82497820FF50FF77FDEFFE83" blockId="4.[151,549,251,309]" pageId="4" pageNumber="5">
<taxonomicName id="4C424D5E82497820FF50FF77FDEBFEA2" ID-CoL="4HFG" authority="Martinez, 1969" authorityName="Martinez" authorityYear="1969" box="[151,545,251,276]" class="Insecta" family="Scarabaeidae" genus="Eutrichillum" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B936EACF82497820FF50FF77FEA5FEA2" bold="true" box="[151,367,251,276]" italics="true" pageId="4" pageNumber="5">EUTRICHILLUM</emphasis>
Martínez, 1969
</taxonomicName>
<taxonomicName id="4C424D5E82497820FF50FE92FDEFFE83" ID-CoL="6HKR3" authority="Solis &amp; Kohlmann, 2003" authorityName="Solis &amp; Kohlmann" authorityYear="2003" box="[151,549,285,309]" class="Insecta" family="Scarabaeidae" genus="Eutrichillum" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="arcus">
<emphasis id="B936EACF82497820FF50FE92FF3EFE83" box="[151,244,286,309]" italics="true" pageId="4" pageNumber="5">E. arcus</emphasis>
(
<bibRefCitation id="EFD34B2C82497820FEC5FE91FDD7FE83" author="Solis" box="[258,541,285,309]" pageId="4" pageNumber="5" refString="Solis, A. &amp; Kohlmann, B. (2003) New species of dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) from Costa Rica and Panama. Zootaxa, 139, 1 - 14." type="journal article" year="2003">Solís &amp; Kohlmann, 2003</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="8BFD36DD82497820FF50FED3FDFDFEC1" blockId="4.[151,587,351,410]" box="[151,567,351,376]" pageId="4" pageNumber="5">
<taxonomicName id="4C424D5E82497820FF50FED3FE9AFECE" ID-CoL="68D6" box="[151,336,351,376]" class="Insecta" family="Scarabaeidae" genus="Onoreidium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B936EACF82497820FF50FED3FE9AFECE" bold="true" box="[151,336,351,376]" italics="true" pageId="4" pageNumber="5">ONOREIDIUM</emphasis>
</taxonomicName>
Vaz-de-Mello, 2008
</paragraph>
<paragraph id="8BFD36DD82497820FF50FE0EFD81FE2F" blockId="4.[151,587,351,410]" box="[151,587,385,410]" pageId="4" pageNumber="5">
<taxonomicName id="4C424D5E82497820FF50FE0EFD81FE2F" ID-CoL="49LLB" authority="Howden &amp; Young, 1981" authorityName="Howden &amp; Young" authorityYear="1981" box="[151,587,385,410]" class="Insecta" family="Scarabaeidae" genus="Onoreidium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bottimeri">
<emphasis id="B936EACF82497820FF50FE0EFEEAFE2F" box="[151,288,385,409]" italics="true" pageId="4" pageNumber="5">
<collectingCountry id="F355764D82497820FF50FE0EFF64FE2F" box="[151,174,386,409]" name="Iceland" pageId="4" pageNumber="5">O</collectingCountry>
. bottimeri
</emphasis>
(
<bibRefCitation id="EFD34B2C82497820FEE9FE0DFD89FE2F" author="Howden" box="[302,579,385,410]" pageId="4" pageNumber="5" refString="Howden, H. F. &amp; Young, O. (1981) Panamanian Scarabaeinae: taxonomy, distribution, and habits (Coleoptera, Scarabaeidae). Contributions of the American Entomological Institute, 18, 1 - 204." type="journal article" year="1981">Howden &amp; Young, 1981</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="8BFD36DD82497820FF50FE4FFE19FE6A" blockId="4.[151,657,451,609]" box="[151,467,451,476]" pageId="4" pageNumber="5">
<taxonomicName id="4C424D5E82497820FF50FE4FFE19FE6A" ID-CoL="7D7C" authority="Erichson, 1847" authorityName="Erichson" authorityYear="1847" box="[151,467,451,476]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B936EACF82497820FF50FE4FFEE9FE6A" bold="true" box="[151,291,451,476]" italics="true" pageId="4" pageNumber="5">SCATIMUS</emphasis>
Erichson, 1847
</taxonomicName>
</paragraph>
<paragraph id="8BFD36DD82497820FF50FE6AFDF8FE4B" blockId="4.[151,657,451,609]" box="[151,562,485,509]" pageId="4" pageNumber="5">
<taxonomicName id="4C424D5E82497820FF50FE6AFDF8FE4B" ID-CoL="4V3Y7" authority="Kohlmann &amp; Solis, 1996" authorityName="Kohlmann &amp; Solis" authorityYear="1996" box="[151,562,485,509]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="erinnyos">
<emphasis id="B936EACF82497820FF50FE6AFED9FE4B" box="[151,275,486,509]" italics="true" pageId="4" pageNumber="5">S. erinnyos</emphasis>
Kohlmann &amp; Solís, 1996
</taxonomicName>
</paragraph>
<paragraph id="8BFD36DD82497820FF50FD8BFE5DFDA8" blockId="4.[151,657,451,609]" box="[151,407,518,543]" pageId="4" pageNumber="5">
<taxonomicName id="4C424D5E82497820FF50FD8BFE5DFDA8" ID-CoL="4V3YC" authority="Harold, 1862" authorityName="Harold" authorityYear="1862" box="[151,407,518,543]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="ovatus">
<emphasis id="B936EACF82497820FF50FD8BFF36FDA9" box="[151,252,519,543]" italics="true" pageId="4" pageNumber="5">S. ovatus</emphasis>
Harold, 1862
</taxonomicName>
</paragraph>
<paragraph id="8BFD36DD82497820FF00FDA4FD88FDD7" blockId="4.[151,657,451,609]" pageId="4" pageNumber="5">
<taxonomicName id="4C424D5E82497820FF00FDA4FE28FD89" ID-CoL="79VWH" authority="Preudhomme" authorityName="Preudhomme" box="[199,482,551,576]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="patruelis">
<emphasis id="B936EACF82497820FF00FDA4FE8CFDF6" box="[199,326,552,576]" italics="true" pageId="4" pageNumber="5">S. patruelis</emphasis>
Preudhomme
</taxonomicName>
de Borre, 1886
<taxonomicName id="4C424D5E82497820FF00FDC6FD88FDD7" ID-CoL="4V3YG" authority="Balthasar, 1939" authorityName="Balthasar" authorityYear="1939" box="[199,578,585,609]" class="Insecta" family="Scarabaeidae" genus="Scatimus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="quadridentatus">
<emphasis id="B936EACF82497820FF00FDC6FE47FDD7" box="[199,397,585,609]" italics="true" pageId="4" pageNumber="5">S. quadridentatus</emphasis>
Balthasar, 1939
</taxonomicName>
</paragraph>
<paragraph id="8BFD36DD82497820FF50FD07FE1CFD73" blockId="4.[151,601,651,709]" pageId="4" pageNumber="5">
<taxonomicName id="4C424D5E82497820FF50FD07FEB8FD12" ID-CoL="7Z64" box="[151,370,651,676]" class="Insecta" family="Scarabaeidae" genus="Trichillidium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B936EACF82497820FF50FD07FEB8FD12" bold="true" box="[151,370,651,676]" italics="true" pageId="4" pageNumber="5">TRICHILLIDIUM</emphasis>
</taxonomicName>
Vaz-de-Mello, 2008
<taxonomicName id="4C424D5E82497820FF50FD22FE1CFD73" ID-CoL="58927" authority="Robinson, 1948" authorityName="Robinson" authorityYear="1948" box="[151,470,685,709]" class="Insecta" family="Scarabaeidae" genus="Trichillidium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="pilosum">
<emphasis id="B936EACF82497820FF50FD22FEC3FD73" box="[151,265,685,709]" italics="true" pageId="4" pageNumber="5">T. pilosum</emphasis>
(Robinson, 1948)
</taxonomicName>
</paragraph>
<paragraph id="8BFD36DD82497820FF50FD63FE02FCB1" blockId="4.[151,832,751,1309]" box="[151,456,751,776]" pageId="4" pageNumber="5">
<taxonomicName id="4C424D5E82497820FF50FD63FE02FCB1" ID-CoL="85CV" authority="Westwood, 1842" authorityName="Westwood" authorityYear="1842" box="[151,456,751,776]" class="Insecta" family="Scarabaeidae" genus="Uroxys" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B936EACF82497820FF50FD63FECCFCBE" bold="true" box="[151,262,751,776]" italics="true" pageId="4" pageNumber="5">UROXYS</emphasis>
Westwood, 1842
</taxonomicName>
</paragraph>
<paragraph id="8BFD36DD82497820FF00FC9DFDC1FC9F" blockId="4.[151,832,751,1309]" box="[199,523,785,809]" pageId="4" pageNumber="5">
<taxonomicName id="4C424D5E82497820FF00FC9DFDC1FC9F" ID-CoL="745R" authority="Balthasar, 1938" authorityName="Balthasar" authorityYear="1938" box="[199,523,785,809]" class="Insecta" family="Scarabaeidae" genus="Pseuduroxys" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B936EACF82497820FF00FC9DFE9FFC9F" box="[199,341,785,809]" italics="true" pageId="4" pageNumber="5">Pseuduroxys</emphasis>
Balthasar, 1938
</taxonomicName>
</paragraph>
<paragraph id="8BFD36DD82497820FF50FCB8FDDCFCFC" blockId="4.[151,832,751,1309]" box="[151,534,818,843]" pageId="4" pageNumber="5">
<taxonomicName id="4C424D5E82497820FF50FCB8FDDCFCFC" ID-CoL="7DZFY" authority="Pereira &amp; Halffter, 1961" authorityName="Pereira &amp; Halffter" authorityYear="1961" box="[151,534,818,843]" class="Insecta" family="Scarabaeidae" genus="Uroxys" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="boneti">
<emphasis id="B936EACF82497820FF50FCB8FF35FCFD" box="[151,255,819,843]" italics="true" pageId="4" pageNumber="5">U. boneti</emphasis>
Pereira &amp; Halffter, 1961
</taxonomicName>
</paragraph>
<paragraph id="8BFD36DD82497820FF00FCD9FD91FCDA" blockId="4.[151,832,751,1309]" box="[199,603,852,877]" pageId="4" pageNumber="5">
<taxonomicName id="4C424D5E82497820FF00FCD9FD91FCDA" ID-CoL="7DZFZ" authority="Howden &amp; Young, 1981" authorityName="Howden &amp; Young" authorityYear="1981" box="[199,603,852,877]" class="Insecta" family="Scarabaeidae" genus="Uroxys" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bidentis">
<emphasis id="B936EACF82497820FF00FCD9FE88FCDA" box="[199,322,852,876]" italics="true" pageId="4" pageNumber="5">U. bidentis</emphasis>
<bibRefCitation id="EFD34B2C82497820FE8EFCD8FD91FCDA" author="Howden" box="[329,603,852,877]" pageId="4" pageNumber="5" refString="Howden, H. F. &amp; Young, O. (1981) Panamanian Scarabaeinae: taxonomy, distribution, and habits (Coleoptera, Scarabaeidae). Contributions of the American Entomological Institute, 18, 1 - 204." type="journal article" year="1981">Howden &amp; Young, 1981</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BFD36DD82497820FF00FCFAFDD1FC79" blockId="4.[151,832,751,1309]" pageId="4" pageNumber="5">
<taxonomicName id="4C424D5E82497820FF00FCFAFD5CFC3B" ID-CoL="7DZF9" authority="Howden &amp; Young, 1981" authorityName="Howden &amp; Young" authorityYear="1981" box="[199,662,885,910]" class="Insecta" family="Scarabaeidae" genus="Uroxys" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="macrocularis">
<emphasis id="B936EACF82497820FF00FCFAFEB7FC3B" box="[199,381,885,909]" italics="true" pageId="4" pageNumber="5">U. macrocularis</emphasis>
<bibRefCitation id="EFD34B2C82497820FE43FCF9FD5CFC3B" author="Howden" box="[388,662,885,910]" pageId="4" pageNumber="5" refString="Howden, H. F. &amp; Young, O. (1981) Panamanian Scarabaeinae: taxonomy, distribution, and habits (Coleoptera, Scarabaeidae). Contributions of the American Entomological Institute, 18, 1 - 204." type="journal article" year="1981">Howden &amp; Young, 1981</bibRefCitation>
</taxonomicName>
<emphasis id="B936EACF82497820FD5AFCF9FCF5FC3B" bold="true" box="[669,831,885,909]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="A20557B482497820FD5AFCF9FCF5FC3B" box="[669,831,885,909]" pageId="4" pageNumber="5">new synonym</taxonomicNameLabel>
</emphasis>
<taxonomicName id="4C424D5E82497820FF50FC14FD93FC18" ID-CoL="7DZGM" authority="Delgado &amp; Kohlmann, 2006" authorityName="Delgado &amp; Kohlmann" authorityYear="2006" box="[151,601,918,943]" class="Insecta" family="Scarabaeidae" genus="Uroxys" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="deavilai">
<emphasis id="B936EACF82497820FF50FC14FED9FC18" box="[151,275,918,943]" italics="true" pageId="4" pageNumber="5">U. deavilai</emphasis>
Delgado &amp; Kohlmann, 2006
</taxonomicName>
<taxonomicName id="4C424D5E82497820FF50FC35FDD1FC79" ID-CoL="7DZGY" authority="Howden &amp; Young, 1981" authorityName="Howden &amp; Young" authorityYear="1981" box="[151,539,951,976]" class="Insecta" family="Scarabaeidae" genus="Uroxys" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="dybasi">
<emphasis id="B936EACF82497820FF50FC35FEC8FC66" box="[151,258,952,976]" italics="true" pageId="4" pageNumber="5">U. dybasi</emphasis>
<bibRefCitation id="EFD34B2C82497820FECEFC3BFDD1FC79" author="Howden" box="[265,539,951,976]" pageId="4" pageNumber="5" refString="Howden, H. F. &amp; Young, O. (1981) Panamanian Scarabaeinae: taxonomy, distribution, and habits (Coleoptera, Scarabaeidae). Contributions of the American Entomological Institute, 18, 1 - 204." type="journal article" year="1981">Howden &amp; Young, 1981</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BFD36DD82497820FF50FC56FE57FBA4" blockId="4.[151,832,751,1309]" pageId="4" pageNumber="5">
<taxonomicName id="4C424D5E82497820FF50FC56FD80FC47" ID-CoL="7DZH8" authority="Howden &amp; Young, 1981" authorityName="Howden &amp; Young" authorityYear="1981" box="[151,586,985,1010]" class="Insecta" family="Scarabaeidae" genus="Uroxys" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="gatunensis">
<emphasis id="B936EACF82497820FF50FC56FEFBFC47" box="[151,305,986,1009]" italics="true" pageId="4" pageNumber="5">U. gatunensis</emphasis>
<bibRefCitation id="EFD34B2C82497820FEFFFC55FD80FC47" author="Howden" box="[312,586,985,1010]" pageId="4" pageNumber="5" refString="Howden, H. F. &amp; Young, O. (1981) Panamanian Scarabaeinae: taxonomy, distribution, and habits (Coleoptera, Scarabaeidae). Contributions of the American Entomological Institute, 18, 1 - 204." type="journal article" year="1981">Howden &amp; Young, 1981</bibRefCitation>
</taxonomicName>
<taxonomicName id="4C424D5E82497820FF50FC70FE57FBA4" ID-CoL="7DZGV" authority="Arrow, 1933" authorityName="Arrow" authorityYear="1933" box="[151,413,1018,1043]" class="Insecta" family="Scarabaeidae" genus="Uroxys" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="gorgon">
<emphasis id="B936EACF82497820FF50FC70FEC3FBA5" box="[151,265,1020,1043]" italics="true" pageId="4" pageNumber="5">U. gorgon</emphasis>
Arrow, 1933
</taxonomicName>
</paragraph>
<paragraph id="8BFD36DD82497820FF50FB91FE45FBC0" blockId="4.[151,832,751,1309]" pageId="4" pageNumber="5">
<taxonomicName id="4C424D5E82497820FF50FB91FD92FB82" ID-CoL="7DZFG" authority="Howden &amp; Young, 1981" authorityName="Howden &amp; Young" authorityYear="1981" box="[151,600,1052,1077]" class="Insecta" family="Scarabaeidae" genus="Uroxys" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="metagorgon">
<emphasis id="B936EACF82497820FF50FB91FEF5FB82" box="[151,319,1053,1076]" italics="true" pageId="4" pageNumber="5">U. metagorgon</emphasis>
<bibRefCitation id="EFD34B2C82497820FE81FB90FD92FB82" author="Howden" box="[326,600,1052,1077]" pageId="4" pageNumber="5" refString="Howden, H. F. &amp; Young, O. (1981) Panamanian Scarabaeinae: taxonomy, distribution, and habits (Coleoptera, Scarabaeidae). Contributions of the American Entomological Institute, 18, 1 - 204." type="journal article" year="1981">Howden &amp; Young, 1981</bibRefCitation>
</taxonomicName>
<taxonomicName id="4C424D5E82497820FF50FBB2FDAAFBE3" ID-CoL="7DZFD" authority="Howden &amp; Young, 1981" authorityName="Howden &amp; Young" authorityYear="1981" box="[151,608,1085,1110]" class="Insecta" family="Scarabaeidae" genus="Uroxys" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="microcularis">
<emphasis id="B936EACF82497820FF50FBB2FE8DFBE3" box="[151,327,1085,1109]" italics="true" pageId="4" pageNumber="5">U. microcularis</emphasis>
<bibRefCitation id="EFD34B2C82497820FE89FBB1FDAAFBE3" author="Howden" box="[334,608,1085,1110]" pageId="4" pageNumber="5" refString="Howden, H. F. &amp; Young, O. (1981) Panamanian Scarabaeinae: taxonomy, distribution, and habits (Coleoptera, Scarabaeidae). Contributions of the American Entomological Institute, 18, 1 - 204." type="journal article" year="1981">Howden &amp; Young, 1981</bibRefCitation>
</taxonomicName>
<taxonomicName id="4C424D5E82497820FF50FBECFE45FBC0" ID-CoL="7DZFC" authority="Bates, 1887" authorityName="Bates" authorityYear="1887" box="[151,399,1118,1143]" class="Insecta" family="Scarabaeidae" genus="Uroxys" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="micros">
<emphasis id="B936EACF82497820FF50FBECFECEFBC0" box="[151,260,1119,1143]" italics="true" pageId="4" pageNumber="5">U. micros</emphasis>
Bates, 1887
</taxonomicName>
</paragraph>
<paragraph id="8BFD36DD82497820FF50FB0DFDEBFB21" blockId="4.[151,832,751,1309]" box="[151,545,1151,1176]" pageId="4" pageNumber="5">
<taxonomicName id="4C424D5E82497820FF50FB0DFDEBFB21" ID-CoL="7DZFN" authority="Howden &amp; Gill, 1987" authorityName="Howden &amp; Gill" authorityYear="1987" box="[151,545,1151,1176]" class="Insecta" family="Scarabaeidae" genus="Uroxys" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="nebulinus">
<emphasis id="B936EACF82497820FF50FB0DFEECFB2E" box="[151,294,1152,1176]" italics="true" pageId="4" pageNumber="5">U. nebulinus</emphasis>
<bibRefCitation id="EFD34B2C82497820FEEAFBF3FDEBFB21" author="Howden" box="[301,545,1151,1175]" pageId="4" pageNumber="5" refString="Howden, H. F. &amp; Gill, B. D. (1987) New species and records of Panamanian and Costa Rican Scarabaeinae (Coleoptera: Scarabaeidae). Coleopterists Bulletin, 41, 201 - 224." type="journal article" year="1987">Howden &amp; Gill, 1987</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BFD36DD82497820FF50FB2EFE06FB0F" blockId="4.[151,832,751,1309]" box="[151,460,1185,1209]" pageId="4" pageNumber="5">
<taxonomicName id="4C424D5E82497820FF50FB2EFE06FB0F" ID-CoL="7DZFL" authority="Balthasar, 1940" authorityName="Balthasar" authorityYear="1940" box="[151,460,1185,1209]" class="Insecta" family="Scarabaeidae" genus="Uroxys" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="pauliani">
<emphasis id="B936EACF82497820FF50FB2EFEDCFB0F" box="[151,278,1185,1209]" italics="true" pageId="4" pageNumber="5">U. pauliani</emphasis>
Balthasar, 1940
</taxonomicName>
</paragraph>
<paragraph id="8BFD36DD82497820FF00FB48FD97FAAB" blockId="4.[151,832,751,1309]" pageId="4" pageNumber="5">
<taxonomicName id="4C424D5E82497820FF00FB48FD5DFB6C" ID-CoL="7DZGL" authority="Howden &amp; Young, 1981" authorityName="Howden &amp; Young" authorityYear="1981" box="[199,663,1218,1243]" class="Insecta" family="Scarabaeidae" genus="Uroxys" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="depressifrons">
<emphasis id="B936EACF82497820FF00FB48FEB4FB6D" box="[199,382,1219,1243]" italics="true" pageId="4" pageNumber="5">U. depressifrons</emphasis>
<bibRefCitation id="EFD34B2C82497820FE42FB4EFD5DFB6C" author="Howden" box="[389,663,1218,1243]" pageId="4" pageNumber="5" refString="Howden, H. F. &amp; Young, O. (1981) Panamanian Scarabaeinae: taxonomy, distribution, and habits (Coleoptera, Scarabaeidae). Contributions of the American Entomological Institute, 18, 1 - 204." type="journal article" year="1981">Howden &amp; Young, 1981</bibRefCitation>
</taxonomicName>
<emphasis id="B936EACF82497820FD59FB4FFC8AFB6D" bold="true" box="[670,832,1219,1243]" pageId="4" pageNumber="5">
<taxonomicNameLabel id="A20557B482497820FD59FB4FFC8AFB6D" box="[670,832,1219,1243]" pageId="4" pageNumber="5">new synonym</taxonomicNameLabel>
</emphasis>
<taxonomicName id="4C424D5E82497820FF50FB69FD8AFB4A" ID-CoL="7DZFJ" authority="Howden &amp; Young, 1981" authorityName="Howden &amp; Young" authorityYear="1981" box="[151,576,1252,1277]" class="Insecta" family="Scarabaeidae" genus="Uroxys" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="platypyga">
<emphasis id="B936EACF82497820FF50FB69FEEDFB4A" box="[151,295,1252,1276]" italics="true" pageId="4" pageNumber="5">U. platypyga</emphasis>
<bibRefCitation id="EFD34B2C82497820FEE9FB68FD8AFB4A" author="Howden" box="[302,576,1252,1277]" pageId="4" pageNumber="5" refString="Howden, H. F. &amp; Young, O. (1981) Panamanian Scarabaeinae: taxonomy, distribution, and habits (Coleoptera, Scarabaeidae). Contributions of the American Entomological Institute, 18, 1 - 204." type="journal article" year="1981">Howden &amp; Young, 1981</bibRefCitation>
</taxonomicName>
<taxonomicName id="4C424D5E82497820FF50FA8AFD97FAAB" ID-CoL="7DZG3" authority="Howden &amp; Gill, 1987" authorityName="Howden &amp; Gill" authorityYear="1987" box="[151,605,1285,1309]" class="Insecta" family="Scarabaeidae" genus="Uroxys" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="transversifrons">
<emphasis id="B936EACF82497820FF50FA8AFEA8FAAB" box="[151,354,1285,1309]" italics="true" pageId="4" pageNumber="5">U. transversifrons</emphasis>
<bibRefCitation id="EFD34B2C82497820FEAEFA89FD97FAAB" author="Howden" box="[361,605,1285,1309]" pageId="4" pageNumber="5" refString="Howden, H. F. &amp; Gill, B. D. (1987) New species and records of Panamanian and Costa Rican Scarabaeinae (Coleoptera: Scarabaeidae). Coleopterists Bulletin, 41, 201 - 224." type="journal article" year="1987">Howden &amp; Gill, 1987</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>
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