168 lines
17 KiB
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168 lines
17 KiB
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<document id="ABBE074150678FD798308334D4BD9344" ID-DOI="10.11646/zootaxa.4104.1.1" ID-GBIF-Dataset="9b3e7716-1099-48ca-803b-620c25d788f1" ID-ISSN="1175-5326" ID-Zenodo-Dep="265809" ID-ZooBank="D598E724-C9E4-4BBA-B25D-511300A47B1D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1461050759050" checkinUser="plazi" docAuthor="Duellman, William E., Marion, Angela B. & Hedges, Blair" docDate="2016" docId="03EA87A5FFB61236F3988FA2317AF578" docLanguage="en" docName="zootaxa.4104.pdf.imf" docOrigin="Zootaxa 4104 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Phyllomedusidae Gunther 1859" docType="treatment" docVersion="11" lastPageNumber="32" masterDocId="FFD3FFDDFFA91229F30F8D5E333EF35E" masterDocTitle="Phylogenetics, classification, and biogeography of the treefrogs (Amphibia: Anura: Arboranae)" masterLastPageNumber="109" masterPageNumber="1" pageNumber="32" updateTime="1698579563523" updateUser="plazi">
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<mods:title id="7C489B9143C15D8B10B424857377EA3F">Phylogenetics, classification, and biogeography of the treefrogs (Amphibia: Anura: Arboranae)</mods:title>
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<mods:namePart id="EA41392E958D44D4B337241925458DEA">Duellman, William E.</mods:namePart>
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<mods:namePart id="994E6E89555004AAC0B277CAF1A23794">Marion, Angela B.</mods:namePart>
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<mods:name id="C04D017F3639FAFE58B028D5ACB14F15" type="personal">
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<mods:namePart id="E115A2F871F7A13605D7EE0D242DABC6">Hedges, Blair</mods:namePart>
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<mods:typeOfResource id="28DF344387AC7D4D1FA87537D8E0EA31">text</mods:typeOfResource>
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<mods:title id="0071B8EA24100CF23AF9A7517C436D01">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="015F41C7383BFA05A6F739EBAECC773B">
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<mods:date id="69FB5821F8AB3355637711E86D267C1A">2016</mods:date>
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<mods:detail id="1B0E7B87120DE75A4A366070B9FAC71C" type="volume">
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<mods:number id="C697D0C7F400EB55D5D9E4FA1CDBB78D">4104</mods:number>
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<mods:number id="C2DCB44AAACF38461F5F41297BFDD324">1</mods:number>
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<mods:classification id="456FFC8470496C5032F53EF48FA883F8">journal article</mods:classification>
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<mods:identifier id="E4812BBE7983B40107C6895DD407DC5B" type="DOI">10.11646/zootaxa.4104.1.1</mods:identifier>
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<mods:identifier id="608EE556246D81F382A2BEE1E1CF6FED" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03EA87A5FFB61236F3988FA2317AF578" ID-DOI="http://doi.org/10.5281/zenodo.5458532" ID-GBIF-Taxon="119612009" ID-Zenodo-Dep="5458532" LSID="urn:lsid:plazi:treatment:03EA87A5FFB61236F3988FA2317AF578" httpUri="http://treatment.plazi.org/id/03EA87A5FFB61236F3988FA2317AF578" lastPageNumber="32" pageId="31" pageNumber="32">
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<subSubSection id="C3596538FFB61236F3988FA231BBF048" box="[151,645,764,790]" pageId="31" pageNumber="32" type="nomenclature">
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<paragraph id="8BFC36B3FFB61236F3988FA231BBF048" blockId="31.[151,645,764,790]" box="[151,645,764,790]" pageId="31" pageNumber="32">
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<heading id="D0B481DFFFB61236F3988FA231BBF048" bold="true" box="[151,645,764,790]" fontSize="11" level="1" pageId="31" pageNumber="32" reason="1">
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<emphasis id="B937EAA1FFB61236F3988FA231BBF048" bold="true" box="[151,645,764,790]" pageId="31" pageNumber="32">
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Family
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<taxonomicName id="4C434D30FFB61236F3F98FA231BBF048" ID-CoL="625JR" authority="Gunther, 1859" authorityName="Gunther" authorityYear="1859" box="[246,645,764,790]" class="Amphibia" family="Phyllomedusidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="31" pageNumber="32" phylum="Chordata" rank="family">Phyllomedusidae Günther, 1859</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3596538FFB61236F3988E1C3173F770" pageId="31" pageNumber="32" type="reference_group">
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<paragraph id="8BFC36B3FFB61236F3988E1C37A2F006" blockId="31.[151,1180,834,856]" box="[151,1180,834,856]" pageId="31" pageNumber="32">
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<taxonomicName id="4C434D30FFB61236F3988E1C32D7F006" ID-CoL="625JR" authority="Gunther" authorityName="Gunther" authorityYear="1858" box="[151,489,834,856]" class="Amphibia" family="Phyllomedusidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="31" pageNumber="32" phylum="Chordata" rank="family">
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Phyllomedusidae
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<bibRefCitation id="EFD24B42FFB61236F2448E1C32D7F006" author="Gunther" box="[331,489,834,856]" pageId="31" pageNumber="32" refString="Gunther, A. C. L. G. (" 1858 " 1859) Catalogue of the Batrachia Salientia in the Collection of the British Museum. Taylor and Francis, London, xvi + 160 pp." type="book" year="1858">Günther, “1858</bibRefCitation>
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</taxonomicName>
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” 1859:346.
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<typeStatus id="54F88811FFB61236F1698E1D31A6F006" box="[614,664,835,856]" pageId="31" pageNumber="32">Type</typeStatus>
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genus:
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<taxonomicName id="4C434D30FFB61236F1E88E1C3738F006" authority="Wagler, 1830" authorityName="Wagler" authorityYear="1830" box="[743,1030,834,856]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="31" pageNumber="32" phylum="Chordata" rank="genus">
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<emphasis id="B937EAA1FFB61236F1E88E1C3048F006" box="[743,886,834,856]" italics="true" pageId="31" pageNumber="32">Phyllomedusa</emphasis>
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<bibRefCitation id="EFD24B42FFB61236F0738E1C3738F006" author="Wagler" box="[892,1030,834,856]" pageId="31" pageNumber="32" refString="Wagler, J. (1830) Naturliches System der Amphibien, mit vorangehender Classification der Saugthiere und Vogel. Ein Beitrag zur vergleichenden Zoologie. J. G. Cotta, Munchen, Stuttgart and Tubingen, 354 pp." type="book" year="1830">Wagler, 1830</bibRefCitation>
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</taxonomicName>
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, by monotypy.
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</paragraph>
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<paragraph id="8BFC36B3FFB61236F3988EDB3789F754" blockId="31.[151,1436,901,1574]" pageId="31" pageNumber="32">
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<emphasis id="B937EAA1FFB61236F3988EDB3228F0C0" bold="true" box="[151,278,901,926]" pageId="31" pageNumber="32">Definition.</emphasis>
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<taxonomicName id="4C434D30FFB61236F22A8EDB3274F0C0" box="[293,330,901,926]" class="Amphibia" family="Hylidae" genus="Litoria" kingdom="Animalia" order="Anura" pageId="31" pageNumber="104" phylum="Chordata" rank="species" species="iris">Iris</taxonomicName>
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vertically elliptical; arciferal pectoral girdle, eight procoelous presacral vertebrae, separate calcaneum and astragalus, intercalary cartilages between terminal and penultimate phalanges, claw-shaped terminal phalanges, SVL from
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<quantity id="4CBB9B56FFB61236F11D8E933157F0BB" box="[530,617,973,998]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" pageId="31" pageNumber="32" unit="mm" value="45.0">45 mm</quantity>
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in
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<taxonomicName id="4C434D30FFB61236F1938E9337DEF0B8" ID-CoL="PQD4" authority="Duellman et al. 1988" authorityName="Duellman et al." authorityYear="1988" box="[668,1248,973,998]" class="Amphibia" family="Phyllomedusidae" genus="Callimedusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="atelopoides">
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<emphasis id="B937EAA1FFB61236F1938E933084F0B8" box="[668,954,973,998]" italics="true" pageId="31" pageNumber="32">Callimedusa atelopoides</emphasis>
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(
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<bibRefCitation id="EFD24B42FFB61236F0DF8E9337E6F0B8" author="Duellman" box="[976,1240,973,998]" pageId="31" pageNumber="32" refString="Duellman, W. E., Cadle, J. E. & Cannatella, D. C. (1988) A new species of terrestrial Phyllomedusa (Anura: Hylidae) from southern Peru. Herpetologica, 44, 91 - 95." type="journal article" year="1988">
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Duellman
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<emphasis id="B937EAA1FFB61236F75F8E9037AEF0B8" box="[1104,1168,973,998]" italics="true" pageId="31" pageNumber="32">et al.</emphasis>
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1988
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</bibRefCitation>
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)
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</taxonomicName>
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to
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<quantity id="4CBB9B56FFB61236F61B8E903646F0BB" box="[1300,1400,974,998]" metricMagnitude="-1" metricUnit="m" metricValue="1.19" pageId="31" pageNumber="32" unit="mm" value="119.0">119 mm</quantity>
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in
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<taxonomicName id="4C434D30FFB61236F3988EAF3023F754" authority="Venancio & Melo-Sampiro, 2010" authorityName="Venancio & Melo-Sampiro" authorityYear="2010" box="[151,797,1009,1034]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="bicolor">
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<emphasis id="B937EAA1FFB61236F3988EAF32B0F754" box="[151,398,1009,1034]" italics="true" pageId="31" pageNumber="32">Phyllomedusa bicolor</emphasis>
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(
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<bibRefCitation id="EFD24B42FFB61236F2948EAF302BF754" author="Venancio" box="[411,789,1009,1034]" pageId="31" pageNumber="32" refString="Venancio, N. M. & Melo-Sampaio, P. R. (2010) Reproductive behavior of the giant leaf-frog Phyllomedusa bicolor (Anura: Hylidae) in the western Amazon. Phyllomedusa, 9, 63 - 67. http: // dx. doi. org / 10.11606 / issn. 2316 - 9079. v 9 i 1 p 63 - 67" type="journal article" year="2010">Venâncio & Melo-Sampiro, 2010</bibRefCitation>
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)
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</taxonomicName>
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. Chromosome complement 2n = 26.
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</paragraph>
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<paragraph id="8BFC36B3FFB61236F3C8894B3173F770" blockId="31.[151,1436,901,1574]" box="[199,589,1045,1070]" pageId="31" pageNumber="32">
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<emphasis id="B937EAA1FFB61236F3C8894B3210F770" bold="true" box="[199,302,1045,1070]" pageId="31" pageNumber="32">Content.</emphasis>
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Eight genera, 59 species.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3596538FFB61236F3C8896730BAF70C" box="[199,900,1081,1106]" pageId="31" pageNumber="32" type="distribution">
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<paragraph id="8BFC36B3FFB61236F3C8896730BAF70C" blockId="31.[151,1436,901,1574]" box="[199,900,1081,1106]" pageId="31" pageNumber="32">
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<emphasis id="B937EAA1FFB61236F3C88967325EF70C" bold="true" box="[199,352,1081,1106]" pageId="31" pageNumber="32">Distribution.</emphasis>
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Neotropics, from
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<collectingCountry id="F3547623FFB61236F122896731BDF70C" box="[557,643,1081,1106]" name="Mexico" pageId="31" pageNumber="32">Mexico</collectingCountry>
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to northern
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<collectingCountry id="F3547623FFB61236F002896730BEF70C" box="[781,896,1081,1106]" name="Argentina" pageId="31" pageNumber="32">Argentina</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3596538FFB61236F3C88903308AF7C7" pageId="31" pageNumber="32" type="etymology">
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<paragraph id="8BFC36B3FFB61236F3C88903308AF7C7" blockId="31.[151,1436,901,1574]" pageId="31" pageNumber="32">
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<emphasis id="B937EAA1FFB61236F3C889033270F728" bold="true" box="[199,334,1117,1142]" pageId="31" pageNumber="32">Etymology.</emphasis>
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The familial name is derived from the Greek
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<emphasis id="B937EAA1FFB61236F06C8903309CF728" box="[867,930,1117,1142]" italics="true" pageId="31" pageNumber="32">phyll,</emphasis>
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meaning leaf, and the Greek
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<emphasis id="B937EAA1FFB61236F7F589033656F728" box="[1274,1384,1117,1142]" italics="true" pageId="31" pageNumber="32">Medousa.</emphasis>
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The name alludes to the gelatinous egg masses deposited on leaves of trees.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3596538FFB61236F3C889FB317AF578" pageId="31" pageNumber="32" type="discussion">
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<paragraph id="8BFC36B3FFB61236F3C889FB317AF578" blockId="31.[151,1436,901,1574]" pageId="31" pageNumber="32">
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<emphasis id="B937EAA1FFB61236F3C889FB3205F7E0" bold="true" box="[199,315,1189,1214]" pageId="31" pageNumber="32">Remarks.</emphasis>
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Herein we include three genera that have not been recognized recently by previous authors. Two of these names are resurrected from the synonomies of
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<taxonomicName id="4C434D30FFB61236F1E389973056F7BC" box="[748,872,1225,1250]" class="Amphibia" family="Hylidae" genus="Agalychnis" kingdom="Animalia" order="Anura" pageId="31" pageNumber="32" phylum="Chordata" rank="genus">
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<emphasis id="B937EAA1FFB61236F1E389973056F7BC" box="[748,872,1225,1250]" italics="true" pageId="31" pageNumber="32">Agalychnis</emphasis>
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</taxonomicName>
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and
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<emphasis id="B937EAA1FFB61236F0AE89973779F7BC" box="[929,1095,1225,1250]" italics="true" pageId="31" pageNumber="32">
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<taxonomicName id="4C434D30FFB61236F0AE8997377CF7BC" box="[929,1090,1225,1250]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="31" pageNumber="32" phylum="Chordata" rank="genus">Phyllomedusa</taxonomicName>
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,
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</emphasis>
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whereas the third is new. Our maximum likelihood analysis includes 46 species, 78% of the known members of the family. In the parsimony analysis of 45 species by
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<bibRefCitation id="EFD24B42FFB61236F2BA884F3187F674" box="[437,697,1297,1322]" pageId="31" pageNumber="32" refString="Faivovich, J., Haddad, C. F. B., Baeta, D., Jungfer, K. - H., Alvares, G. F. R., Brandao, R. A., Sheil, C., Barrientos, L. S., Barrio- Amoros, C. L., Cruz, C. A. G. & Wheeler, W. C. (2010) The phylogenetic relationships of the charismatic poster frogs, Phyllomedusinae (Anura, Hylidae). Cladistics, 26, 227 - 261. http: // dx. doi. org / 10.1111 / j. 1096 - 0031.2009.00287. x" type="journal article">
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Faivovich
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<emphasis id="B937EAA1FFB61236F123884C315BF674" box="[556,613,1297,1322]" italics="true" pageId="31" pageNumber="32">et al.</emphasis>
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(2010)
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</bibRefCitation>
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, some of the same clades emerge as in our analysis. The principal difference is in the proposed classifications. Our proposal of three additional genera is a reflection on some of the species groups recognized by
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<bibRefCitation id="EFD24B42FFB61236F2E2880731CAF62C" box="[493,756,1369,1394]" pageId="31" pageNumber="32" refString="Faivovich, J., Haddad, C. F. B., Baeta, D., Jungfer, K. - H., Alvares, G. F. R., Brandao, R. A., Sheil, C., Barrientos, L. S., Barrio- Amoros, C. L., Cruz, C. A. G. & Wheeler, W. C. (2010) The phylogenetic relationships of the charismatic poster frogs, Phyllomedusinae (Anura, Hylidae). Cladistics, 26, 227 - 261. http: // dx. doi. org / 10.1111 / j. 1096 - 0031.2009.00287. x" type="journal article">
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Faivovich
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<emphasis id="B937EAA1FFB61236F16A880431A1F62C" box="[613,671,1369,1394]" italics="true" pageId="31" pageNumber="32">et al.</emphasis>
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(2010)
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</bibRefCitation>
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. We recovered a major clade of phyllomedusids with 100% support. Within this clade are large frogs, genus
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<taxonomicName id="4C434D30FFB61236F1CC8823305FF6C8" box="[707,865,1405,1430]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" kingdom="Animalia" order="Anura" pageId="31" pageNumber="32" phylum="Chordata" rank="genus">
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<emphasis id="B937EAA1FFB61236F1CC8823305FF6C8" box="[707,865,1405,1430]" italics="true" pageId="31" pageNumber="32">Phyllomedusa</emphasis>
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</taxonomicName>
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(100% support), with vomerine teeth and another clade with 100% support of smaller frogs lacking vomerine teeth (100% support). Within the latter are two clades with significant support. One clade,
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<taxonomicName id="4C434D30FFB61236F13F889B3192F680" box="[560,684,1477,1502]" class="Amphibia" family="Phyllomedusidae" genus="Pithecopus" higherTaxonomySource="CoL" kingdom="Animalia" order="Anura" pageId="31" pageNumber="32" phylum="Chordata" rank="genus">
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<emphasis id="B937EAA1FFB61236F13F889B3192F680" box="[560,684,1477,1502]" italics="true" pageId="31" pageNumber="32">Pithecopus</emphasis>
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</taxonomicName>
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(100% support), is characterized by having opposable thumbs and no bright flash colors. In the second clade,
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<taxonomicName id="4C434D30FFB61236F18E88B73031F55C" box="[641,783,1513,1538]" class="Amphibia" family="Phyllomedusidae" genus="Callimedusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="31" pageNumber="33" phylum="Chordata" rank="genus" status="gen. nov.">
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<emphasis id="B937EAA1FFB61236F18E88B73031F55C" box="[641,783,1513,1538]" italics="true" pageId="31" pageNumber="32">Callimedusa</emphasis>
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</taxonomicName>
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(78% support), the thumb is not opposable, and with one exception, all have bright flash colors.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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