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<document id="2BDB7969A67EB4D260D7A9CD07E22094" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626380265891" checkinUser="felipe" docAuthor="Dietrich, C. H." docDate="2008" docId="038A213D491BFFC5FF31FF2065B73AC6" docLanguage="en" docName="zt01727p038.pdf" docOrigin="Zootaxa 1727" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Smerdalea veracruzensis Dietrich 2008, n. sp." docType="treatment" docVersion="3" lastPageNumber="32" masterDocId="FFB359454918FFC6FFA6FFB765533F6A" masterDocTitle="Some remarkable new Neotropical treehoppers (Hemiptera: Cicadomorpha: Membracidae)" masterLastPageNumber="38" masterPageNumber="29" pageNumber="32" updateTime="1699010893382" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="B6EF23A8ACC65D5671E013822FA21F00">Some remarkable new Neotropical treehoppers (Hemiptera: Cicadomorpha: Membracidae)</mods:title>
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<mods:namePart id="9199D7C966A1CF035BD81D85346AC063">Dietrich, C. H.</mods:namePart>
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<mods:date id="2A3D41DE9D5A485A4638BE3731E4AC52">2008</mods:date>
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<mods:number id="0D95B22250FC60F7B8A8DFB677B232FB">2008-03-14</mods:number>
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<mods:number id="C62450912E33A516EDE1316D04252FEA">1727</mods:number>
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<treatment id="038A213D491BFFC5FF31FF2065B73AC6" ID-DOI="http://doi.org/10.5281/zenodo.5107846" ID-Zenodo-Dep="5107846" LSID="urn:lsid:plazi:treatment:038A213D491BFFC5FF31FF2065B73AC6" httpUri="http://treatment.plazi.org/id/038A213D491BFFC5FF31FF2065B73AC6" lastPageNumber="32" pageId="3" pageNumber="32">
<subSubSection id="C339C3A0491BFFC5FF31FF20675F3FDB" box="[151,524,151,177]" pageId="3" pageNumber="32" type="nomenclature">
<paragraph id="8B9C902B491BFFC5FF31FF20675F3FDB" blockId="3.[151,524,151,212]" box="[151,524,151,177]" pageId="3" pageNumber="32">
<heading id="D0D42747491BFFC5FF31FF20675F3FDB" bold="true" box="[151,524,151,177]" fontSize="11" level="1" pageId="3" pageNumber="32" reason="1">
<emphasis id="B9574C39491BFFC5FF31FF20675F3FDB" bold="true" box="[151,524,151,177]" pageId="3" pageNumber="32">
<taxonomicName id="4C23EBA8491BFFC5FF31FF2064903FDB" authority="Dietrich, 2008" authorityName="Dietrich" authorityYear="2008" box="[151,451,151,177]" class="Insecta" family="Membracidae" genus="Smerdalea" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="veracruzensis" status="sp. nov.">
<emphasis id="B9574C39491BFFC5FF31FF2064903FDB" bold="true" box="[151,451,151,177]" italics="true" pageId="3" pageNumber="32">Smerdalea veracruzensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A264F142491BFFC5FE6CFF20675F3FDB" box="[458,524,151,177]" pageId="3" pageNumber="32" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C339C3A0491BFFC5FF31FF0D64E23FBE" box="[151,433,186,212]" pageId="3" pageNumber="32" type="description">
<paragraph id="8B9C902B491BFFC5FF31FF0D64E23FBE" blockId="3.[151,524,151,212]" box="[151,433,186,212]" pageId="3" pageNumber="32">
(
<figureCitation id="13188CAE491BFFC5FF39FF0D645A3FBE" box="[159,265,186,212]" captionStart="FIGURES 18" captionStartId="2.[151,269,1834,1858]" captionTargetBox="[179,1409,197,1754]" captionTargetId="figure-80@2.[169,1417,197,1817]" captionTargetPageId="2" captionText="FIGURES 18. 15, habitus. 1, Smerdalea veracruzensis, holotype, lateral view; 2, Smerdalea imminens, male (Onkonegare, Orellana, Ecuador), lateral view; 34, Smergotomia clairae, holotype, lateral and dorsal views; 5, Braxtonota enigmata, holotype, lateral view. 6, S. veracruzensis, second and third valvulae, lateral view. 7, S. clairae, second valvulae, lateral view. 8, B. enigmata, second and third valvulae, lateral view." pageId="3" pageNumber="32">Figs 1, 6</figureCitation>
,
<figureCitation id="13188CAE491BFFC5FEB3FF0D64023FBE" box="[277,337,186,212]" captionStart="FIGURES 916" captionStartId="5.[151,269,1779,1803]" captionTargetBox="[188,1400,382,1749]" captionTargetId="figure-183@5.[169,1417,376,1775]" captionTargetPageId="5" captionText="FIGURES 916. 911, head, anterior view. 9, S. veracruzensis, holotype; 10, S. clairae, holotype; 11, B. enigmata, holotype. 1216, wings. 12, S. veracruzensis, forewing; 1314, S. clairae, fore- and hind wing; 1516, B. enigmata, fore- and hind wing." pageId="3" pageNumber="32">9, 12</figureCitation>
,
<figureCitation id="13188CAE491BFFC5FEF8FF0D64FA3FBE" box="[350,425,186,212]" captionStart="FIGURES 1724" captionStartId="8.[151,269,1699,1723]" captionTargetBox="[248,1331,303,1679]" captionTargetId="figure-159@8.[238,1358,298,1690]" captionTargetPageId="8" captionText="FIGURES 1724. male genitalia. 1720, S. veracruzensis: 17, pygofer and subgenital plate, lateral view; 18, sternite IX and subgenital plate, ventral view; 1920, aedeagus, style, and connective, lateral and ventral views. 2124, S. clairae: 21, pygofer and segment X, dorsal view; 22, sternite IX and subgenital plate, ventral view; 2324, aedeagus, style, and connective, lateral and ventral views." pageId="3" pageNumber="32">1722</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C339C3A0491BFFC5FF31FEBD601F3E1E" pageId="3" pageNumber="32" type="diagnosis">
<paragraph id="8B9C902B491BFFC5FF31FEBD601F3E1E" blockId="3.[151,1436,266,1452]" pageId="3" pageNumber="32">
<emphasis id="B9574C39491BFFC5FF31FEBD64443E4E" bold="true" box="[151,279,266,292]" pageId="3" pageNumber="32">Diagnosis.</emphasis>
This species is readily distinguished from other species in the genus by the structure of the pronotum, which has the suprahumeral horns depressed and spinelike, the posterior process lacking a dorsal node basally and the terminal node clublike rather than strongly compressed and lacking a dorsal projection.
</paragraph>
</subSubSection>
<subSubSection id="C339C3A0491BFFC5FF60FE35649D3B06" pageId="3" pageNumber="32" type="description">
<paragraph id="8B9C902B491BFFC5FF60FE35649D3B06" blockId="3.[151,1436,266,1452]" pageId="3" pageNumber="32">
<emphasis id="B9574C39491BFFC5FF60FE35640D3EF6" bold="true" box="[198,350,386,412]" pageId="3" pageNumber="32">Description.</emphasis>
<emphasis id="B9574C39491BFFC5FEC0FE3567063EF1" box="[358,597,386,411]" italics="true" pageId="3" pageNumber="32">Measurements (mm)</emphasis>
. Length, male 9.8, female 10.811.1; head width, male 4.2, female 4.2; pronotum width, male 4.7, female 4.95.2; forewing length, male 8.5, female 9.49.5;
<emphasis id="B9574C39491BFFC5FB21FE1D605B3EA9" box="[1159,1288,426,451]" italics="true" pageId="3" pageNumber="32">Coloration</emphasis>
. Nearly uniform brown dorsally with face, thoracic pleuron and abdominal sternites dull yellow; forewing with broad dark brown macula extended across midlength; tibiae dark brown with broad yellow area near midlength.
<emphasis id="B9574C39491BFFC5FACDFE4D65863D51" italics="true" pageId="3" pageNumber="32">Vestiture</emphasis>
. Head, pronotum, scutellum, and basal two-thirds of forewing conspicuously pubescent.
<emphasis id="B9574C39491BFFC5FB43FD9560033D51" box="[1253,1360,546,571]" italics="true" pageId="3" pageNumber="32">Structure</emphasis>
. Head (
<figureCitation id="13188CAE491BFFC5FF39FDFD65B83D0E" box="[159,235,586,612]" captionStart="FIGURES 916" captionStartId="5.[151,269,1779,1803]" captionTargetBox="[188,1400,382,1749]" captionTargetId="figure-183@5.[169,1417,376,1775]" captionTargetPageId="5" captionText="FIGURES 916. 911, head, anterior view. 9, S. veracruzensis, holotype; 10, S. clairae, holotype; 11, B. enigmata, holotype. 1216, wings. 12, S. veracruzensis, forewing; 1314, S. clairae, fore- and hind wing; 1516, B. enigmata, fore- and hind wing." pageId="3" pageNumber="32">Fig. 9</figureCitation>
) without dorsal projections; ocelli sessile; dorsomedian concavity shallow; frontoclypeus without median carina. Pronotum (
<figureCitation id="13188CAE491BFFC5FE72FDC5674C3DE6" box="[468,543,626,652]" captionStart="FIGURES 18" captionStartId="2.[151,269,1834,1858]" captionTargetBox="[179,1409,197,1754]" captionTargetId="figure-80@2.[169,1417,197,1817]" captionTargetPageId="2" captionText="FIGURES 18. 15, habitus. 1, Smerdalea veracruzensis, holotype, lateral view; 2, Smerdalea imminens, male (Onkonegare, Orellana, Ecuador), lateral view; 34, Smergotomia clairae, holotype, lateral and dorsal views; 5, Braxtonota enigmata, holotype, lateral view. 6, S. veracruzensis, second and third valvulae, lateral view. 7, S. clairae, second valvulae, lateral view. 8, B. enigmata, second and third valvulae, lateral view." pageId="3" pageNumber="32">Figs 1</figureCitation>
,
<figureCitation id="13188CAE491BFFC5FD96FDC5676C3DE6" box="[560,575,626,652]" captionStart="FIGURES 916" captionStartId="5.[151,269,1779,1803]" captionTargetBox="[188,1400,382,1749]" captionTargetId="figure-183@5.[169,1417,376,1775]" captionTargetPageId="5" captionText="FIGURES 916. 911, head, anterior view. 9, S. veracruzensis, holotype; 10, S. clairae, holotype; 11, B. enigmata, holotype. 1216, wings. 12, S. veracruzensis, forewing; 1314, S. clairae, fore- and hind wing; 1516, B. enigmata, fore- and hind wing." pageId="3" pageNumber="32">9</figureCitation>
) depressed anteriorly; suprahumeral horns short, spinelike, not elevated above midline, projecting laterally slightly beyond humeri; median carina not elevated anterad of horns; posterior process without basal hump, slightly arcuate in lateral view, lateral carina absent; posterior node resting in posterior emargination of scutellum, flattened dorsally with weak median carina, with pair of short posterolateral spines, posterodorsal spines and angulate dorsomedial projection absent. Scutellum humped basally, depressed distally, apex emarginate with lateral lobes rounded. Hind femur with few scattered preapical cucullate setae dorsally; tibia with cucullate setal rows IIII well developed, row I double. Forewing (
<figureCitation id="13188CAE491BFFC5FABFFCD560363C16" box="[1305,1381,866,892]" captionStart="FIGURES 18" captionStartId="2.[151,269,1834,1858]" captionTargetBox="[179,1409,197,1754]" captionTargetId="figure-80@2.[169,1417,197,1817]" captionTargetPageId="2" captionText="FIGURES 18. 15, habitus. 1, Smerdalea veracruzensis, holotype, lateral view; 2, Smerdalea imminens, male (Onkonegare, Orellana, Ecuador), lateral view; 34, Smergotomia clairae, holotype, lateral and dorsal views; 5, Braxtonota enigmata, holotype, lateral view. 6, S. veracruzensis, second and third valvulae, lateral view. 7, S. clairae, second valvulae, lateral view. 8, B. enigmata, second and third valvulae, lateral view." pageId="3" pageNumber="32">Figs 1</figureCitation>
,
<figureCitation id="13188CAE491BFFC5FAD3FCD560C03C16" box="[1397,1427,866,892]" captionStart="FIGURES 916" captionStartId="5.[151,269,1779,1803]" captionTargetBox="[188,1400,382,1749]" captionTargetId="figure-183@5.[169,1417,376,1775]" captionTargetPageId="5" captionText="FIGURES 916. 911, head, anterior view. 9, S. veracruzensis, holotype; 10, S. clairae, holotype; 11, B. enigmata, holotype. 1216, wings. 12, S. veracruzensis, forewing; 1314, S. clairae, fore- and hind wing; 1516, B. enigmata, fore- and hind wing." pageId="3" pageNumber="32">12</figureCitation>
) with opaque sclerotization in basal two-thirds, darkly pigmented and pubescent except in hyaline area between vein M and costal margin in distal third. Male pygofer (
<figureCitation id="13188CAE491BFFC5FC30FC0566BF3CA6" box="[918,1004,946,972]" captionStart="FIGURES 1724" captionStartId="8.[151,269,1699,1723]" captionTargetBox="[248,1331,303,1679]" captionTargetId="figure-159@8.[238,1358,298,1690]" captionTargetPageId="8" captionText="FIGURES 1724. male genitalia. 1720, S. veracruzensis: 17, pygofer and subgenital plate, lateral view; 18, sternite IX and subgenital plate, ventral view; 1920, aedeagus, style, and connective, lateral and ventral views. 2124, S. clairae: 21, pygofer and segment X, dorsal view; 22, sternite IX and subgenital plate, ventral view; 2324, aedeagus, style, and connective, lateral and ventral views." pageId="3" pageNumber="32">Fig. 17</figureCitation>
) with prominent posterolateral lobe; subgenital plate (
<figureCitation id="13188CAE491BFFC5FEC4FC6D64EB3C9E" box="[354,440,986,1012]" captionStart="FIGURES 1724" captionStartId="8.[151,269,1699,1723]" captionTargetBox="[248,1331,303,1679]" captionTargetId="figure-159@8.[238,1358,298,1690]" captionTargetPageId="8" captionText="FIGURES 1724. male genitalia. 1720, S. veracruzensis: 17, pygofer and subgenital plate, lateral view; 18, sternite IX and subgenital plate, ventral view; 1920, aedeagus, style, and connective, lateral and ventral views. 2124, S. clairae: 21, pygofer and segment X, dorsal view; 22, sternite IX and subgenital plate, ventral view; 2324, aedeagus, style, and connective, lateral and ventral views." pageId="3" pageNumber="32">Fig. 18</figureCitation>
) narrower than sternite IX, tapered distally; aedeagus in ventral view (
<figureCitation id="13188CAE491BFFC5FAA6FC6D60043C9E" box="[1280,1367,986,1012]" captionStart="FIGURES 1724" captionStartId="8.[151,269,1699,1723]" captionTargetBox="[248,1331,303,1679]" captionTargetId="figure-159@8.[238,1358,298,1690]" captionTargetPageId="8" captionText="FIGURES 1724. male genitalia. 1720, S. veracruzensis: 17, pygofer and subgenital plate, lateral view; 18, sternite IX and subgenital plate, ventral view; 1920, aedeagus, style, and connective, lateral and ventral views. 2124, S. clairae: 21, pygofer and segment X, dorsal view; 22, sternite IX and subgenital plate, ventral view; 2324, aedeagus, style, and connective, lateral and ventral views." pageId="3" pageNumber="32">Fig. 22</figureCitation>
) with expanded distal section occupying ca. one-third length of shaft; ejaculatory duct internal, not extended beyond base of gonopore; gonopore margin with numerous microtrichia. Female with second valvulae slender throughout length (
<figureCitation id="13188CAE491BFFC5FEDEFBE564933B06" box="[376,448,1106,1132]" captionStart="FIGURES 18" captionStartId="2.[151,269,1834,1858]" captionTargetBox="[179,1409,197,1754]" captionTargetId="figure-80@2.[169,1417,197,1817]" captionTargetPageId="2" captionText="FIGURES 18. 15, habitus. 1, Smerdalea veracruzensis, holotype, lateral view; 2, Smerdalea imminens, male (Onkonegare, Orellana, Ecuador), lateral view; 34, Smergotomia clairae, holotype, lateral and dorsal views; 5, Braxtonota enigmata, holotype, lateral view. 6, S. veracruzensis, second and third valvulae, lateral view. 7, S. clairae, second valvulae, lateral view. 8, B. enigmata, second and third valvulae, lateral view." pageId="3" pageNumber="32">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C339C3A0491BFFC5FF60FBCD65B73AC6" pageId="3" pageNumber="32" type="materials_examined">
<paragraph id="8B9C902B491BFFC5FF60FBCD61403B8E" blockId="3.[151,1436,266,1452]" pageId="3" pageNumber="32">
<emphasis id="B9574C39491BFFC5FF60FBCD64EC3BFE" bold="true" box="[198,447,1146,1172]" pageId="3" pageNumber="32">Material examined.</emphasis>
<materialsCitation id="3B4B9A76491BFFC5FE6DFBCD61403B8E" collectingDate="1989-09-20" collectingDateMax="1998-12-03" collectingDateMin="1989-09-20" collectionCode="UNAM, INHS" collectorName="H. Rojas &amp; J. L. Colin &amp; E. Mejorada" country="Mexico" location="Estacion Biologica Los Tuxtlas" pageId="3" pageNumber="32" specimenCount="3" specimenCount-female="2" stateProvince="Veracruz de Ignacio de la Llave" typeStatus="holotype">
<typeStatus id="54982E89491BFFC5FE6DFBCD676B3BFE" box="[459,568,1146,1172]" pageId="3" pageNumber="32" type="holotype">Holotype</typeStatus>
male,
<collectingCountry id="F334D0BB491BFFC5FD36FBCD66543BFE" box="[656,775,1146,1172]" name="Mexico" pageId="3" pageNumber="32">MEXICO</collectingCountry>
,
<collectingRegion id="49E75EC9491BFFC5FCB1FBCD66D13BFE" box="[791,898,1146,1172]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="3" pageNumber="32">Veracruz</collectingRegion>
,
<location id="8EFCC6F0491BFFC5FC34FBCD60403BFE" LSID="urn:lsid:plazi:treatment:038A213D491BFFC5FF31FF2065B73AC6:8EFCC6F0491BFFC5FC34FBCD60403BFE" box="[914,1299,1146,1172]" country="Mexico" name="Estacion Biologica Los Tuxtlas" pageId="3" pageNumber="32" stateProvince="Veracruz de Ignacio de la Llave">Estacion Biologica Los Tuxtlas</location>
,
<date id="FF9DB6EB491BFFC5FA85FBCD65823BD6" pageId="3" pageNumber="32" value="1989-10-09">
<collectingDate id="EFD94F03491BFFC5FA85FBCD65823BD6" pageId="3" pageNumber="32" value="1989-10-09">9 October 1989</collectingDate>
</date>
(
<collectorName id="26D6F5FD491BFFC5FF45FB15641C3BD6" box="[227,335,1186,1212]" pageId="3" pageNumber="32">H. Rojas</collectorName>
,
<collectorName id="26D6F5FD491BFFC5FEFDFB1564883BD6" box="[347,475,1186,1212]" pageId="3" pageNumber="32">J. L. Colin</collectorName>
) [
<collectionCode id="ED3208EE491BFFC5FE55FB1567053BD6" box="[499,598,1186,1212]" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="3" pageNumber="32">UNAM</collectionCode>
];
<specimenCount id="9D255BA2491BFFC5FDCCFB1567823BD6" box="[618,721,1186,1212]" pageId="3" pageNumber="32" type="female">1 female</specimenCount>
<typeStatus id="54982E89491BFFC5FD7FFB1566123BD6" box="[729,833,1186,1212]" pageId="3" pageNumber="32" type="paratype">paratype</typeStatus>
, same data except
<date id="FF9DB6EB491BFFC5FB87FB1560553BD6" box="[1057,1286,1186,1212]" pageId="3" pageNumber="32" value="1989-09-20">
<collectingDate id="EFD94F03491BFFC5FB87FB1560553BD6" box="[1057,1286,1186,1212]" pageId="3" pageNumber="32" value="1989-09-20">20 September 1989</collectingDate>
</date>
[
<collectionCode id="ED3208EE491BFFC5FAB0FB15602A3BD6" box="[1302,1401,1186,1212]" country="Mexico" httpUri="http://grbio.org/cool/10ei-x9nj" name="Universidad Nacional Autonoma de Mexico" pageId="3" pageNumber="32">UNAM</collectionCode>
];
<specimenCount id="9D255BA2491BFFC5FA2BFB1565B53B8E" pageId="3" pageNumber="32" type="female">1 female</specimenCount>
<typeStatus id="54982E89491BFFC5FF48FB7D64053B8E" box="[238,342,1226,1252]" pageId="3" pageNumber="32" type="paratype">paratype</typeStatus>
, same data except
<date id="FF9DB6EB491BFFC5FD97FB7D66533B8E" box="[561,768,1226,1252]" pageId="3" pageNumber="32" value="1998-12-03">
<collectingDate id="EFD94F03491BFFC5FD97FB7D66533B8E" box="[561,768,1226,1252]" pageId="3" pageNumber="32" value="1998-12-03">3 December 1998</collectingDate>
</date>
(
<collectorName id="26D6F5FD491BFFC5FCB7FB7D66F63B8E" box="[785,933,1226,1252]" pageId="3" pageNumber="32">E. Mejorada</collectorName>
) [
<collectionCode id="ED3208EE491BFFC5FC1DFB7D615B3B8E" box="[955,1032,1226,1252]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34797" lsid="urn:lsid:biocol.org:col:34797" name="Illinois Natural History Survey" pageId="3" pageNumber="32" type="Other type of research institution/biorepository">INHS</collectionCode>
].
</materialsCitation>
</paragraph>
<paragraph id="8B9C902B491BFFC5FF60FB4565B73AC6" blockId="3.[151,1436,266,1452]" pageId="3" pageNumber="32">
<emphasis id="B9574C39491BFFC5FF60FB4564403A66" bold="true" box="[198,275,1266,1292]" pageId="3" pageNumber="32">Notes.</emphasis>
Because this species has the least ornate pronotum of all known
<taxonomicName id="4C23EBA8491BFFC5FBBEFB4461A63A66" box="[1048,1269,1266,1292]" class="Insecta" family="Membracidae" genus="Smerdalea" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="32" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B9574C39491BFFC5FBBEFB4461C73A66" box="[1048,1172,1267,1292]" italics="true" pageId="3" pageNumber="32">Smerdalea</emphasis>
species
</taxonomicName>
, it may be the most plesiomorphic member of the genus. It is also unusual in having the female second valvulae narrow throughout their length; the other species of the genus have the second valvulae abruptly broadened preapically. The species name, an adjective, refers to the Mexican state in which all known specimens were collected.
</paragraph>
</subSubSection>
</treatment>
</document>