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<document id="83287C9599CE28D23D38B086DCF004B7" ID-DOI="10.5281/zenodo.3673403" ID-GBIF-Dataset="f01c9bd1-7635-40c7-b5ee-5fc57320b880" ID-ISSN="1942-1354" ID-Zenodo-Dep="3673403" ID-ZooBank="08CB0DA0-C83E-438F-AA97-8CD637F044A5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582027555556" checkinUser="felipe" docAuthor="Paulsen, M. J." docDate="2019" docId="0381879FFF9D6E50FF1C9EB7FA02FE55" docLanguage="en" docName="InsectaMundi.0706.1-10.pdf.imf" docOrigin="Insecta Mundi 706 (706)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Ochodaeus Dejean 1821" docType="key" docVersion="7" lastPageNumber="9" masterDocId="FFB8FFE7FF946E5AFFDC9C2EFFA2FFB4" masterDocTitle="New species of Ochodaeus Dejean from Madagascar II (Coleoptera: Scarabaeoidea: Ochodaeidae)" masterLastPageNumber="10" masterPageNumber="1" pageNumber="8" updateTime="1698718828345" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="4BC615501F8017B5F31CE2C7A3EDB4B3">New species of Ochodaeus Dejean from Madagascar II (Coleoptera: Scarabaeoidea: Ochodaeidae)</mods:title>
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<mods:namePart id="AB79EE60E42FC62D9E9E5E97643EC31E">Paulsen, M. J.</mods:namePart>
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<mods:typeOfResource id="300CAB8BF066F5D52667A5FF9D0C05AC">text</mods:typeOfResource>
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<mods:title id="BEFD098D7D5D75581207F634CA11911E">Insecta Mundi</mods:title>
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<mods:part id="117756E085C74E4A03F00A36904D6F26">
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<mods:date id="41044F2FB4D666D771AFA95A052115E9">2019</mods:date>
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<mods:number id="4BBD76E69833245A798CBAB26EE9E411">2019-05-31</mods:number>
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<mods:number id="FDCB052F76CCAD7E7DEA8C790A6997BF">706</mods:number>
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<mods:detail id="79B8CCA5358470998E0918B468C35BAF" type="issue">
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<mods:number id="C177EFD383335D5150B82E13861FC776">706</mods:number>
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<mods:classification id="922581BA613785CE0F00B610057E327A">journal article</mods:classification>
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<mods:identifier id="262C265AA3E71E8E409597E8B67A21B8" type="DOI">10.5281/zenodo.3673403</mods:identifier>
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<mods:identifier id="43BCAE135B78588E04FA6CBF20DF6877" type="ISSN">1942-1354</mods:identifier>
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</mods:mods>
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<subSection id="E2A72F62FF9D6E50FF1C9EB7FA02FE55" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8" type="multiple">
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<treatment id="0381879FFF9D6E50FF1C9EB7FA02FE55" ID-DOI="http://doi.org/10.5281/zenodo.3679989" ID-GBIF-Taxon="162161796" ID-Zenodo-Dep="3679989" LSID="urn:lsid:plazi:treatment:0381879FFF9D6E50FF1C9EB7FA02FE55" httpUri="http://treatment.plazi.org/id/0381879FFF9D6E50FF1C9EB7FA02FE55" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8">
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<subSubSection id="C3326502FF9D6E53FF1C9EB7FC1EFD05" box="[192,956,665,689]" pageId="9" pageNumber="8" type="nomenclature">
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<paragraph id="8B973689FF9D6E53FF1C9EB7FC1EFD05" blockId="9.[192,956,665,689]" box="[192,956,665,689]" pageId="9" pageNumber="8">
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<heading id="D0DF81E5FF9D6E53FF1C9EB7FC1EFD05" bold="true" box="[192,956,665,689]" fontSize="10" level="2" pageId="9" pageNumber="8" reason="3">
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<emphasis id="B95CEA9BFF9D6E53FF1C9EB7FC1EFD05" bold="true" box="[192,956,665,689]" pageId="9" pageNumber="8">
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Updated key couplets to
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<taxonomicName id="4C284D0AFF9D6E53FDC99EB7FD09FD05" ID-CoL="66FL" authorityName="Dejean" authorityYear="1821" box="[533,683,665,689]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="genus">
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<emphasis id="B95CEA9BFF9D6E53FDC99EB7FD09FD05" bold="true" box="[533,683,665,689]" italics="true" pageId="9" pageNumber="8">Ochodaeus</emphasis>
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</taxonomicName>
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spp. in
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<collectingCountry id="F33F7619FF9D6E53FCC99EB7FC1EFD05" box="[789,956,665,689]" name="Madagascar" pageId="9" pageNumber="8">Madagascar</collectingCountry>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3326502FF9D6E53FF349EE7FAF2FCB5" pageId="9" pageNumber="8">
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<paragraph id="8B973689FF9D6E53FF349EE7FAF2FCB5" blockId="9.[192,1440,713,769]" pageId="9" pageNumber="8">
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The following couplet changes to the species key in
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<bibRefCitation id="EFB94B78FF9D6E53FCAC9EE7FB8BFD56" author="Paulsen, M. J." box="[880,1065,713,738]" pageId="9" pageNumber="8" pagination="1 - 14" refId="ref6680" refString="Paulsen, M. J. 2019. Three new Madagascan species of Ochodaeus Dejean (Coleoptera: Scarabaeoidea: Ochodaeidae). Insecta Mundi 0684: 1 - 14." type="book chapter" year="2019">Paulsen (2019)</bibRefCitation>
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allow for determination of all Madagascan species having a granulate pronotum, including the three newly described species.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3326502FF9D6E50FF1C9F34FA02FE55" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8" type="key">
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<key id="FFB961B8FF9D6E50FF1C9F34FA02FE55" lastPageId="10" lastPageNumber="9" pageId="9" pageNumber="8">
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<paragraph id="8B973689FF9D6E53FF1C9F34FA02FCC5" blockId="9.[192,1440,793,881]" pageId="9" pageNumber="8">
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<keyStep id="30DC2E2CFF9D6E53FF1C9F34FA02FCC5" pageId="9" pageNumber="8">
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<keyLead id="30D995BCFF9D6E53FF1C9F34FA02FCC5" pageId="9" pageNumber="8">
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2(1). Elytral intervals with setose tubercles of 2 sizes, with setae of larger tubercles flattened, scale-like........................................................
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<taxonomicName id="4C284D0AFF9D6E53FB5C9F17FA02FCE5" ID-CoL="6SF2R" authority="Paulian" authorityName="Paulian" authorityYear="1976" box="[1152,1440,825,849]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="pygmaeus">
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<emphasis id="B95CEA9BFF9D6E53FB5C9F17FA02FCE5" bold="true" box="[1152,1440,825,849]" pageId="9" pageNumber="8">
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<emphasis id="B95CEA9BFF9D6E53FB5C9F17FA8FFCE5" bold="true" box="[1152,1325,825,849]" italics="true" pageId="9" pageNumber="8">O. pygmaeus</emphasis>
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Paulian
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</emphasis>
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</taxonomicName>
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— Elytral intervals with tubercles either absent or of uniform size and without flattened setae.
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<emphasis id="B95CEA9BFF9D6E53FA4D9F77FA02FCC5" bold="true" box="[1425,1440,857,881]" pageId="9" pageNumber="8">3</emphasis>
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</keyLead>
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</keyStep>
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</paragraph>
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<paragraph id="8B973689FF9D6E53FF1C9FA4FA02FC55" blockId="9.[192,1440,905,994]" pageId="9" pageNumber="8">
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<keyStep id="30DC2E2CFF9D6E53FF1C9FA4FA02FC55" pageId="9" pageNumber="8">
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<keyLead id="30D995BCFF9D6E53FF1C9FA4FA02FC55" pageId="9" pageNumber="8">
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3(2). Clypeus with median tubercle or small horn that is distinct from anterior margin (
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<figureCitation id="13132A0CFF9D6E53FAEC9FA7FA3AFC15" box="[1328,1432,905,929]" captionStart="Figures 4–8" captionStartId="9.[192,287,1790,1812]" captionTargetBox="[192,1440,1171,1757]" captionTargetId="figure@9.[192,1440,1161,1759]" captionTargetPageId="9" captionText="Figures 4–8. Key characters of Madagascar Ochodaeus species. 4) Head of O. meridialis Paulsen, n. sp., showing central tubercle of clypeus. 5) Head of O. umbonulus Paulsen, n. sp., showing tubercle anterior to straight frontoclypeal suture. 6) Head of O. iniquipes Paulsen, n. sp., showing minute horn on posteriorly curved frontoclypeal suture. 7) Head of O. isoanalensis Paulian showing broadly thickened clypeal margin. 8) Head of O. miliaris (Klug), showing narrowly thickened clypeal margin." figureDoi="http://doi.org/10.5281/zenodo.3673411" httpUri="https://zenodo.org/record/3673411/files/figure.png" pageId="9" pageNumber="8">Fig. 4–6</figureCitation>
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)...........................................................................
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<emphasis id="B95CEA9BFF9D6E53FA4D9F87FA02FC75" bold="true" box="[1425,1440,937,961]" pageId="9" pageNumber="8">4</emphasis>
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— Clypeus lacking free tubercle or horn, anterior margin usually thickened medially (
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<figureCitation id="13132A0CFF9D6E53FAD79FE7FACCFC56" box="[1291,1390,969,994]" captionStart="Figures 4–8" captionStartId="9.[192,287,1790,1812]" captionTargetBox="[192,1440,1171,1757]" captionTargetId="figure@9.[192,1440,1161,1759]" captionTargetPageId="9" captionText="Figures 4–8. Key characters of Madagascar Ochodaeus species. 4) Head of O. meridialis Paulsen, n. sp., showing central tubercle of clypeus. 5) Head of O. umbonulus Paulsen, n. sp., showing tubercle anterior to straight frontoclypeal suture. 6) Head of O. iniquipes Paulsen, n. sp., showing minute horn on posteriorly curved frontoclypeal suture. 7) Head of O. isoanalensis Paulian showing broadly thickened clypeal margin. 8) Head of O. miliaris (Klug), showing narrowly thickened clypeal margin." figureDoi="http://doi.org/10.5281/zenodo.3673411" httpUri="https://zenodo.org/record/3673411/files/figure.png" pageId="9" pageNumber="8">Fig. 7–8</figureCitation>
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).
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<emphasis id="B95CEA9BFF9D6E53FA4D9FE7FA02FC55" bold="true" box="[1425,1440,969,993]" pageId="9" pageNumber="8">6</emphasis>
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</keyLead>
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</keyStep>
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</paragraph>
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<paragraph id="8B973689FF9D6E53FF1C9FD4FA02FB85" blockId="9.[192,1440,1017,1073]" pageId="9" pageNumber="8">
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<keyStep id="30DC2E2CFF9D6E53FF1C9FD4FA02FB85" pageId="9" pageNumber="8">
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<keyLead id="30D995BCFF9D6E53FF1C9FD4FA02FB85" pageId="9" pageNumber="8">
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4(3). Mentum with a knobby protuberance on each side (
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<figureCitation id="13132A0CFF9D6E53FC589FD7FC7AFBA5" box="[900,984,1017,1041]" captionStart="Figure 1" captionStartId="4.[192,275,1448,1470]" captionTargetBox="[242,1418,281,1404]" captionTargetId="figure@4.[192,1440,263,1414]" captionTargetPageId="4" captionText="Figure 1. Ochodaeus meridialis Paulsen, new species. a) Dorsal habitus, male holotype. b) Right front leg, dorsal view.c) Right stridulatory peg, dorsal view (elytra lifted). d) Head, female paratype, ventral view. Arrows indicating knob-like protuberances of mentum. e) Holotype labels." figureDoi="http://doi.org/10.5281/zenodo.3673405" httpUri="https://zenodo.org/record/3673405/files/figure.png" pageId="9" pageNumber="8">Fig. 1d</figureCitation>
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)....
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<emphasis id="B95CEA9BFF9D6E53FBFC9FD7FA02FBA5" bold="true" box="[1056,1440,1017,1041]" pageId="9" pageNumber="8">
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<taxonomicName id="4C284D0AFF9D6E53FBFC9FD7FAE9FBA5" ID-CoL="48DDC" authority="Paulsen" authorityName="Paulsen" authorityYear="2019" box="[1056,1355,1017,1041]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="meridialis" status="sp. nov.">
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<emphasis id="B95CEA9BFF9D6E53FBFC9FD7FB76FBA5" bold="true" box="[1056,1236,1017,1041]" italics="true" pageId="9" pageNumber="8">O. meridialis</emphasis>
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Paulsen
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</taxonomicName>
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,
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<taxonomicNameLabel id="A26F57E0FF9D6E53FA859FD7FA02FBA5" box="[1369,1440,1017,1041]" pageId="9" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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— Mentum lacking protuberances laterally............................................
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<emphasis id="B95CEA9BFF9D6E53FA4D9837FA02FB85" bold="true" box="[1425,1440,1049,1073]" pageId="9" pageNumber="8">5</emphasis>
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</keyLead>
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</keyStep>
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</paragraph>
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<caption id="DF576601FF9D6E53FF1C9AD0FD08F83C" ID-DOI="http://doi.org/10.5281/zenodo.3673411" ID-Zenodo-Dep="3673411" httpUri="https://zenodo.org/record/3673411/files/figure.png" pageId="9" pageNumber="8" startId="9.[192,287,1790,1812]" targetBox="[192,1440,1171,1757]" targetPageId="9">
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<paragraph id="8B973689FF9D6E53FF1C9AD0FD08F83C" blockId="9.[192,1439,1790,1928]" pageId="9" pageNumber="8">
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<emphasis id="B95CEA9BFF9D6E53FF1C9AD0FEF6F8A7" bold="true" box="[192,340,1790,1812]" pageId="9" pageNumber="8">Figures 4–8.</emphasis>
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Key characters of Madagascar
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<taxonomicName id="4C284D0AFF9D6E53FD719AD0FC87F8A7" ID-CoL="66FL" authorityName="Dejean" authorityYear="1821" box="[685,805,1790,1811]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="genus">
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<emphasis id="B95CEA9BFF9D6E53FD719AD0FC87F8A7" box="[685,805,1790,1811]" italics="true" pageId="9" pageNumber="8">Ochodaeus</emphasis>
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</taxonomicName>
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species.
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<emphasis id="B95CEA9BFF9D6E53FC5A9AD0FC3FF8A0" bold="true" box="[902,925,1790,1812]" pageId="9" pageNumber="8">4)</emphasis>
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Head of
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<taxonomicName id="4C284D0AFF9D6E53FC239AD1FB53F8A1" ID-CoL="48DDC" authority="Paulsen" authorityName="Paulsen" authorityYear="2019" box="[1023,1265,1790,1813]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="meridialis" status="sp. nov.">
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<emphasis id="B95CEA9BFF9D6E53FC239AD1FB32F8A7" box="[1023,1168,1790,1812]" italics="true" pageId="9" pageNumber="8">O. meridialis</emphasis>
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Paulsen
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</taxonomicName>
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,
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<taxonomicNameLabel id="A26F57E0FF9D6E53FB209AD1FA9AF8A0" box="[1276,1336,1791,1812]" pageId="9" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
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, showing central tubercle of clypeus.
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<emphasis id="B95CEA9BFF9D6E53FDDF9B35FDB9F885" bold="true" box="[515,539,1819,1841]" pageId="9" pageNumber="8">5)</emphasis>
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Head of
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<taxonomicName id="4C284D0AFF9D6E53FD569B32FC30F886" ID-CoL="74F4J" authority="Paulsen" authorityName="Paulsen" authorityYear="2019" box="[650,914,1819,1842]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="umbonulus" status="sp. nov.">
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<emphasis id="B95CEA9BFF9D6E53FD569B32FC8EF884" box="[650,812,1819,1841]" italics="true" pageId="9" pageNumber="8">O. umbonulus</emphasis>
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Paulsen
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</taxonomicName>
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,
|
|||
|
|
<taxonomicNameLabel id="A26F57E0FF9D6E53FC7D9B32FC41F885" box="[929,995,1820,1841]" pageId="9" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
|
|||
|
|
, showing tubercle anterior to straight frontoclypeal suture.
|
|||
|
|
<emphasis id="B95CEA9BFF9D6E53FE789B16FE18F8FA" bold="true" box="[420,442,1848,1870]" pageId="9" pageNumber="8">6)</emphasis>
|
|||
|
|
Head of
|
|||
|
|
<taxonomicName id="4C284D0AFF9D6E53FDCA9B16FD54F8FA" ID-CoL="48DCQ" authority="Paulsen" authorityName="Paulsen" authorityYear="2019" box="[534,758,1848,1870]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="iniquipes" status="sp. nov.">
|
|||
|
|
<emphasis id="B95CEA9BFF9D6E53FDCA9B16FD35F8FA" box="[534,663,1848,1870]" italics="true" pageId="9" pageNumber="8">O. iniquipes</emphasis>
|
|||
|
|
Paulsen
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicNameLabel id="A26F57E0FF9D6E53FD239B17FC9BF8FA" box="[767,825,1849,1870]" pageId="9" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
|
|||
|
|
, showing minute horn on posteriorly curved frontoclypeal suture.
|
|||
|
|
<emphasis id="B95CEA9BFF9D6E53FECF9B7BFE88F8DF" bold="true" box="[275,298,1877,1899]" pageId="9" pageNumber="8">7)</emphasis>
|
|||
|
|
Head of
|
|||
|
|
<taxonomicName id="4C284D0AFF9D6E53FE549B7BFD2AF8DF" ID-CoL="74F35" authority="Paulian" authorityName="Paulian" authorityYear="1959" box="[392,648,1877,1899]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="isoanalensis">
|
|||
|
|
<emphasis id="B95CEA9BFF9D6E53FE549B7BFD8FF8DE" box="[392,557,1877,1898]" italics="true" pageId="9" pageNumber="8">O. isoanalensis</emphasis>
|
|||
|
|
Paulian
|
|||
|
|
</taxonomicName>
|
|||
|
|
showing broadly thickened clypeal margin.
|
|||
|
|
<emphasis id="B95CEA9BFF9D6E53FBB89B7BFBD8F8DF" bold="true" box="[1124,1146,1877,1899]" pageId="9" pageNumber="8">8)</emphasis>
|
|||
|
|
Head of
|
|||
|
|
<taxonomicName id="4C284D0AFF9D6E53FB049B7BFA39F8DF" ID-CoL="48DDD" authority="(Klug)" baseAuthorityName="Klug" box="[1240,1435,1877,1899]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="miliaris">
|
|||
|
|
<emphasis id="B95CEA9BFF9D6E53FB049B7BFAECF8DE" box="[1240,1358,1877,1898]" italics="true" pageId="9" pageNumber="8">O. miliaris</emphasis>
|
|||
|
|
(Klug)
|
|||
|
|
</taxonomicName>
|
|||
|
|
, showing narrowly thickened clypeal margin.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<keyStep id="30DC2E2CFF9E6E50FF1C9CF7FA02FEE5" pageId="10" pageNumber="9">
|
|||
|
|
<paragraph id="8B973689FF9E6E50FF1C9CF7FA02FEA5" blockId="10.[192,1440,217,337]" pageId="10" pageNumber="9">
|
|||
|
|
<keyLead id="30D995BCFF9E6E50FF1C9CF7FA02FEA5" pageId="10" pageNumber="9">
|
|||
|
|
5(4). Clypeus (
|
|||
|
|
<figureCitation id="13132A0CFF9E6E50FE5D9CF7FE66FF45" box="[385,452,217,241]" captionStart="Figures 4–8" captionStartId="9.[192,287,1790,1812]" captionTargetBox="[192,1440,1171,1757]" captionTargetId="figure@9.[192,1440,1161,1759]" captionTargetPageId="9" captionText="Figures 4–8. Key characters of Madagascar Ochodaeus species. 4) Head of O. meridialis Paulsen, n. sp., showing central tubercle of clypeus. 5) Head of O. umbonulus Paulsen, n. sp., showing tubercle anterior to straight frontoclypeal suture. 6) Head of O. iniquipes Paulsen, n. sp., showing minute horn on posteriorly curved frontoclypeal suture. 7) Head of O. isoanalensis Paulian showing broadly thickened clypeal margin. 8) Head of O. miliaris (Klug), showing narrowly thickened clypeal margin." figureDoi="http://doi.org/10.5281/zenodo.3673411" httpUri="https://zenodo.org/record/3673411/files/figure.png" pageId="10" pageNumber="9">Fig. 5</figureCitation>
|
|||
|
|
) with median tubercle anterior to frontoclypeal suture; anterior margin of clypeus indistinct and eroded medially........................
|
|||
|
|
<emphasis id="B95CEA9BFF9E6E50FBCA9CD7FA02FEA5" bold="true" box="[1046,1440,249,273]" pageId="10" pageNumber="9">
|
|||
|
|
<taxonomicName id="4C284D0AFF9E6E50FBCA9CD7FAE9FEA5" ID-CoL="74F4J" authority="Paulsen" authorityName="Paulsen" authorityYear="2019" box="[1046,1355,249,273]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="umbonulus" status="sp. nov.">
|
|||
|
|
<emphasis id="B95CEA9BFF9E6E50FBCA9CD7FB71FEA5" bold="true" box="[1046,1235,249,273]" italics="true" pageId="10" pageNumber="9">O. umbonulus</emphasis>
|
|||
|
|
Paulsen
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicNameLabel id="A26F57E0FF9E6E50FA859CD7FA02FEA5" box="[1369,1440,249,273]" pageId="10" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B973689FF9E6E50FF1C9D34FA02FEE5" blockId="10.[192,1440,217,337]" pageId="10" pageNumber="9">
|
|||
|
|
<keyLead id="30D995BCFF9E6E50FF1C9D34FA02FEE5" pageId="10" pageNumber="9">
|
|||
|
|
— Clypeus (
|
|||
|
|
<figureCitation id="13132A0CFF9E6E50FE5B9D37FE72FE85" box="[391,464,281,305]" captionStart="Figures 4–8" captionStartId="9.[192,287,1790,1812]" captionTargetBox="[192,1440,1171,1757]" captionTargetId="figure@9.[192,1440,1161,1759]" captionTargetPageId="9" captionText="Figures 4–8. Key characters of Madagascar Ochodaeus species. 4) Head of O. meridialis Paulsen, n. sp., showing central tubercle of clypeus. 5) Head of O. umbonulus Paulsen, n. sp., showing tubercle anterior to straight frontoclypeal suture. 6) Head of O. iniquipes Paulsen, n. sp., showing minute horn on posteriorly curved frontoclypeal suture. 7) Head of O. isoanalensis Paulian showing broadly thickened clypeal margin. 8) Head of O. miliaris (Klug), showing narrowly thickened clypeal margin." figureDoi="http://doi.org/10.5281/zenodo.3673411" httpUri="https://zenodo.org/record/3673411/files/figure.png" pageId="10" pageNumber="9">Fig. 6</figureCitation>
|
|||
|
|
) with small horn located on frontoclypeal suture; anterior margin of clypeus distinct and entire throughout..........................
|
|||
|
|
<emphasis id="B95CEA9BFF9E6E50FBF39D17FA02FEE5" bold="true" box="[1071,1440,313,337]" pageId="10" pageNumber="9">
|
|||
|
|
<taxonomicName id="4C284D0AFF9E6E50FBF39D17FAE9FEE5" ID-CoL="48DCQ" authority="Paulsen" authorityName="Paulsen" authorityYear="2019" box="[1071,1355,313,337]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="iniquipes" status="sp. nov.">
|
|||
|
|
<emphasis id="B95CEA9BFF9E6E50FBF39D17FB76FEE5" bold="true" box="[1071,1236,313,337]" italics="true" pageId="10" pageNumber="9">O. iniquipes</emphasis>
|
|||
|
|
Paulsen
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicNameLabel id="A26F57E0FF9E6E50FA859D17FA02FEE5" box="[1369,1440,313,337]" pageId="10" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
</keyStep>
|
|||
|
|
<keyStep id="30DC2E2CFF9E6E50FF1C9D47FA02FE55" pageId="10" pageNumber="9">
|
|||
|
|
<paragraph id="8B973689FF9E6E50FF1C9D47FA02FE15" blockId="10.[192,1440,361,481]" pageId="10" pageNumber="9">
|
|||
|
|
<keyLead id="30D995BCFF9E6E50FF1C9D47FA02FE15" pageId="10" pageNumber="9">
|
|||
|
|
6(3). Median thickening of clypeal bead variable, lateral thickening always evident (
|
|||
|
|
<figureCitation id="13132A0CFF9E6E50FB609D47FAA2FE36" box="[1212,1280,361,386]" captionStart="Figures 4–8" captionStartId="9.[192,287,1790,1812]" captionTargetBox="[192,1440,1171,1757]" captionTargetId="figure@9.[192,1440,1161,1759]" captionTargetPageId="9" captionText="Figures 4–8. Key characters of Madagascar Ochodaeus species. 4) Head of O. meridialis Paulsen, n. sp., showing central tubercle of clypeus. 5) Head of O. umbonulus Paulsen, n. sp., showing tubercle anterior to straight frontoclypeal suture. 6) Head of O. iniquipes Paulsen, n. sp., showing minute horn on posteriorly curved frontoclypeal suture. 7) Head of O. isoanalensis Paulian showing broadly thickened clypeal margin. 8) Head of O. miliaris (Klug), showing narrowly thickened clypeal margin." figureDoi="http://doi.org/10.5281/zenodo.3673411" httpUri="https://zenodo.org/record/3673411/files/figure.png" pageId="10" pageNumber="9">Fig. 7</figureCitation>
|
|||
|
|
); body length usually> 9.0 mm........................................
|
|||
|
|
<taxonomicName id="4C284D0AFF9E6E50FB819DA7FA02FE15" ID-CoL="74F35" authority="Paulian" authorityName="Paulian" authorityYear="1959" box="[1117,1440,393,417]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="isoanalensis">
|
|||
|
|
<emphasis id="B95CEA9BFF9E6E50FB819DA7FA02FE15" bold="true" box="[1117,1440,393,417]" pageId="10" pageNumber="9">
|
|||
|
|
<emphasis id="B95CEA9BFF9E6E50FB819DA7FA8EFE15" bold="true" box="[1117,1324,393,417]" italics="true" pageId="10" pageNumber="9">O. isoanalensis</emphasis>
|
|||
|
|
Paulian
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B973689FF9E6E50FF1C9D84FA02FE55" blockId="10.[192,1440,361,481]" pageId="10" pageNumber="9">
|
|||
|
|
<keyLead id="30D995BCFF9E6E50FF1C9D84FA02FE55" pageId="10" pageNumber="9">
|
|||
|
|
— Median thickening of clypeal bead simply convex, bead slender laterally (
|
|||
|
|
<figureCitation id="13132A0CFF9E6E50FB779D87FB56FE76" box="[1195,1268,425,450]" captionStart="Figures 4–8" captionStartId="9.[192,287,1790,1812]" captionTargetBox="[192,1440,1171,1757]" captionTargetId="figure@9.[192,1440,1161,1759]" captionTargetPageId="9" captionText="Figures 4–8. Key characters of Madagascar Ochodaeus species. 4) Head of O. meridialis Paulsen, n. sp., showing central tubercle of clypeus. 5) Head of O. umbonulus Paulsen, n. sp., showing tubercle anterior to straight frontoclypeal suture. 6) Head of O. iniquipes Paulsen, n. sp., showing minute horn on posteriorly curved frontoclypeal suture. 7) Head of O. isoanalensis Paulian showing broadly thickened clypeal margin. 8) Head of O. miliaris (Klug), showing narrowly thickened clypeal margin." figureDoi="http://doi.org/10.5281/zenodo.3673411" httpUri="https://zenodo.org/record/3673411/files/figure.png" pageId="10" pageNumber="9">Fig. 8</figureCitation>
|
|||
|
|
); body length usually 4.5–8.0 mm............................................
|
|||
|
|
<taxonomicName id="4C284D0AFF9E6E50FB1F9DE7FA02FE55" ID-CoL="48DDD" authority="Klug" authorityName="Klug" authorityYear="1832" box="[1219,1440,457,481]" class="Insecta" family="Ochodaeidae" genus="Ochodaeus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="miliaris">
|
|||
|
|
<emphasis id="B95CEA9BFF9E6E50FB1F9DE7FA02FE55" bold="true" box="[1219,1440,457,481]" pageId="10" pageNumber="9">
|
|||
|
|
<emphasis id="B95CEA9BFF9E6E50FB1F9DE7FAF4FE55" bold="true" box="[1219,1366,457,481]" italics="true" pageId="10" pageNumber="9">O. miliaris</emphasis>
|
|||
|
|
Klug
|
|||
|
|
</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
</keyLead>
|
|||
|
|
</paragraph>
|
|||
|
|
</keyStep>
|
|||
|
|
</key>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</subSection>
|
|||
|
|
</document>
|