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206 lines
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<mods:title id="F866A03DF0B4947C1CD72FA8CF371093">Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)</mods:title>
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<mods:namePart id="558948AD3373C1AF97BB4D15D32BE61B">Bistroem, Olof</mods:namePart>
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<mods:namePart id="2C03201A7055956914FD09050FCF0AE8">Nilsson, Anders N.</mods:namePart>
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<mods:namePart id="F57ED60B15069BBA0FCB44A56927DB61">Bergsten, Johannes</mods:namePart>
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<mods:title id="F1F0D72E54B8DBF8CC90B8141FC8B411">ZooKeys</mods:title>
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<mods:date id="414F951947D47EA39621C51E45F3BB93">2015</mods:date>
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<mods:number id="228C925C89668DDAD1023E09C4399E63">542</mods:number>
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<mods:url id="1547EB9CEDA455109B8CEBC2D7E9F5B0">http://dx.doi.org/10.3897/zookeys.542.5975</mods:url>
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<mods:classification id="598C11D1DC55247698C53876321E7AA4">journal article</mods:classification>
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<treatment id="821CC969B567729DEB78B72A1A96ECD4" ID-GBIF-Taxon="127862733" LSID="urn:lsid:plazi:treatment:821CC969B567729DEB78B72A1A96ECD4" httpUri="http://treatment.plazi.org/id/821CC969B567729DEB78B72A1A96ECD4" lastPageId="172" lastPageNumber="173" pageId="171" pageNumber="172">
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<subSubSection id="84FB5707F9B9297FAEB7200744E72659" pageId="171" pageNumber="172" type="multiple">
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<paragraph id="CC97213B6B127AC2E9EE167B648E7222" pageId="171" pageNumber="172">
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<pageBreakToken id="370714129398D4ACDEF8D31260B1FAE5" pageId="171" pageNumber="172" start="start">Taxon</pageBreakToken>
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classification Animalia Coleoptera Dytiscidae
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</paragraph>
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</subSubSection>
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<subSubSection id="812A575FDBE72BAFCF58FFCAF560CFA6" pageId="171" pageNumber="172" type="nomenclature">
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<paragraph id="6DF00FA710CBB8103A2968C5D08D92BB" pageId="171" pageNumber="172">
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<taxonomicName id="F8E3A90B31C883007706C757D1E99065" ID-CoL="3RM7X" authority="Gschwendtner, 1933" authorityName="Gschwendtner" authorityYear="1933" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus incrassatus" order="Coleoptera" pageId="171" pageNumber="172" phylum="Arthropoda" rank="species" species="incrassatus">Laccophilus incrassatus Gschwendtner, 1933</taxonomicName>
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Figs 141-142, 327, 472-473, 564
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</paragraph>
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</subSubSection>
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<subSubSection id="B85AB502567CE0C8763D1EE0AE9A4790" pageId="171" pageNumber="172" type="reference_group">
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<paragraph id="7481E2DF00BB767B42AACBDC2F94E10D" pageId="171" pageNumber="172">
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<taxonomicName id="28F4AF8867CC7ACBCC3981F2A8CC88EA" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus incrassatus" order="Coleoptera" pageId="171" pageNumber="172" phylum="Arthropoda" rank="species" species="incrassatus">Laccophilus incrassatus</taxonomicName>
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<bibRefCitation id="6815F2484B2310BC1B2BEC0E1F9391E6" author="Gschwendtner, L" journalOrPublisher="Revue de Zoologie et Botanique Africaines" pageId="256" pageNumber="257" pagination="84 - 95" title="Neue Schwimmkaefer aus Belgisch Kongo. IV Teil." volume="23" year="1933">Gschwendtner 1933</bibRefCitation>
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: 85 (original description, faunistics);
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<bibRefCitation id="D7C01CAC0EAF4A1A03FB4D0BD62F205D" author="Guignot, F" journalOrPublisher="Memoires du Museum National d'Histoire Naturelle Paris" pageId="256" pageNumber="257" pagination="215 - 322" title="Mission scientifique de l'Omo 6 (58). ColeopteraDytiscidae (2 e partie)." volume="19" year="1946 c">Guignot 1946c</bibRefCitation>
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: 263, 264, 265, 312 (description, faunistics, discussion);
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<bibRefCitation id="C5F329881F6CF81D1B4E8AC9758D1627" author="Legros, C" journalOrPublisher="Memoires de l'Institut Francais d'Afrique Noire" pageId="259" pageNumber="260" pagination="265 - 280" title="XXI. Coleopteres Hydrocanthares (Mission M. Lamotte, 1942)." volume="40" year="1954">Legros 1954</bibRefCitation>
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: 268 (discussion);
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<bibRefCitation id="B973A4DBDB058E24F9FE27F583A4EE9A" author="Guignot, F" journalOrPublisher="Annales du Musee Royal du Congo Belge (Serie 8 vo) Sciences Zoologiques" pageId="258" pageNumber="259" pagination="323 - 648" title="Revision des hydrocanthares d'Afrique (ColeopteraDytiscoidea). 2." volume="78" year="1959 a">Guignot 1959a</bibRefCitation>
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: 544, 549, 550 (description, faunistics);
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<bibRefCitation id="147FBD52C9EE747E4BB45204EF7A4AE6" author="Nilsson, AN" journalOrPublisher="World catalogue of Insects" pageId="260" pageNumber="261" pagination="1 - 395" title="Dytiscidae." volume="3" year="2001">Nilsson 2001</bibRefCitation>
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: 245 (catalogue, faunistics);
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<bibRefCitation id="81BD50BF9985BE5BDC0042E3440AD6E1" author="Nilsson, AN" journalOrPublisher="Apollo books, Stenstrup" pageId="260" pageNumber="261" title="A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga). Version 1. I. 2015" url="http://www2.emg.umu.se/projects/biginst/andersn/" year="2015">Nilsson 2015</bibRefCitation>
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: 212 (catalogue, faunistics).
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</paragraph>
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<paragraph id="BCBC505116FD7EB35D83BCE2A27F3747" pageId="171" pageNumber="172">
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<taxonomicName id="C30FD171082A611F983795D800C7466C" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus virgatus" order="Coleoptera" pageId="171" pageNumber="172" phylum="Arthropoda" rank="species" species="virgatus">Laccophilus virgatus</taxonomicName>
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<bibRefCitation id="EAFDCE4D3CBEE6AD54CCA0A19BE0BAF3" author="Guignot, F" journalOrPublisher="Bulletin de l'Institut Royal des Sciences Naturelles de Belgique" pageId="257" pageNumber="258" pagination="1 - 4" title="Nouveaux dytiscides de Parc National de l'Upemba. Diagnoses provisoires (suite) (1)." volume="29" year="1953 e">Guignot 1953e</bibRefCitation>
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: 4 (original description, faunistics);
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<bibRefCitation id="0929FB29D095D8F392109279B783562A" author="Guignot, F" journalOrPublisher="Exploration du Parc National de l'Upemba, Mission G. F. de Witte (1946 - 1949)" pageId="257" pageNumber="258" pagination="3 - 59" title="Dytiscidae. (Coleoptera, Adephaga)." volume="33" year="1954">Guignot 1954</bibRefCitation>
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: 27 (description, faunistics);
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<bibRefCitation id="F96412ABD75C6918D8E178755A493DF0" author="Guignot, F" journalOrPublisher="Annales du Musee Royal du Congo Belge (Serie 8 vo) Sciences Zoologiques" pageId="258" pageNumber="259" pagination="323 - 648" title="Revision des hydrocanthares d'Afrique (ColeopteraDytiscoidea). 2." volume="78" year="1959 a">Guignot 1959a</bibRefCitation>
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: 544, 546, 550 (description, faunistics);
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<bibRefCitation id="92A303C6FF1D73A9EF01F05710B3A841" author="Nilsson, AN" journalOrPublisher="World catalogue of Insects" pageId="260" pageNumber="261" pagination="1 - 395" title="Dytiscidae." volume="3" year="2001">Nilsson 2001</bibRefCitation>
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: 253 (catalogue, faunistics);
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<bibRefCitation id="79B59F992310AE767D8606E8EF186249" author="Nilsson, AN" journalOrPublisher="Apollo books, Stenstrup" pageId="260" pageNumber="261" title="A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga). Version 1. I. 2015" url="http://www2.emg.umu.se/projects/biginst/andersn/" year="2015">Nilsson 2015</bibRefCitation>
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: 219 (catalogue, faunistics). New synonym.
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</paragraph>
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</subSubSection>
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<subSubSection id="6AF903F227039509CBB8B0B686D534B8" pageId="171" pageNumber="172" type="type localities">
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<paragraph id="A1ADEF88B58FA614360C2F07C4D04388" pageId="171" pageNumber="172">Type localities.</paragraph>
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<paragraph id="E9C9D4E73E2716D48E29D5FFB64E2E38" pageId="171" pageNumber="172">
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<taxonomicName id="2977C10CCFFB6162D109E8FA711FF57B" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus incrassatus" order="Coleoptera" pageId="171" pageNumber="172" phylum="Arthropoda" rank="species" species="incrassatus">Laccophilus incrassatus</taxonomicName>
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: Zaire: Moero, Kasiki.
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</paragraph>
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<paragraph id="AB1783CF20353DAE746884FD707693F5" pageId="171" pageNumber="172">
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<taxonomicName id="09ADD973A87294F424ECF9BC51C0A90F" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus virgatus" order="Coleoptera" pageId="171" pageNumber="172" phylum="Arthropoda" rank="species" species="virgatus">Laccophilus virgatus</taxonomicName>
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: Zaire: Upemba National Park, Riv. Dipwa.
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</paragraph>
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</subSubSection>
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<subSubSection id="89D8C258044AB80C9B84673CE035F62F" pageId="171" pageNumber="172" type="type material">
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<paragraph id="6FD622D39B902618CA4376ED6ED4C31B" pageId="171" pageNumber="172">Type material studied</paragraph>
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<paragraph id="293376EEF44B71D5F2D205836970418C" pageId="171" pageNumber="172">
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(14 exs.).
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<taxonomicName id="FC0F765926721B15CBF521A457A4BDA0" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus incrassatus" order="Coleoptera" pageId="171" pageNumber="172" phylum="Arthropoda" rank="species" species="incrassatus">Laccophilus incrassatus</taxonomicName>
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: Holotype: male: "Holotypus /
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<normalizedToken id="C9ADC119AEC9A50FB069E1E2A7340C27" originalValue="Musée">Musee</normalizedToken>
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du Congo Tang. - Moero: Kasiki 20/27-VI-1931 G.F. de Witte / Type Gschw. / R.
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<normalizedToken id="CCA37AA64566C4E64714AFEA77C0EFFD" originalValue="DÈT">DET</normalizedToken>
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. A 2223 /
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<taxonomicName id="B29CAE9BC0535C5B22407F4BC15FCED2" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus incrassatus" order="Coleoptera" pageId="171" pageNumber="172" phylum="Arthropoda" rank="species" species="incrassatus">Laccophilus incrassatus</taxonomicName>
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Gschw. det. Gschwendt." (MRAC; habitus in Fig. 472). - Paratype: female (considered male in original description): Same data as holotype, but: "Paratype Gschw. / Coll. Gschwendtner / Paratype" (1 ex. OLML).
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</paragraph>
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<paragraph id="0DE24820047C6960328C0B83EF9D3A62" pageId="171" pageNumber="172">
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<taxonomicName id="D3B89B6E902436A12D663E49A58CBA35" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus virgatus" order="Coleoptera" pageId="171" pageNumber="172" phylum="Arthropoda" rank="species" species="virgatus">Laccophilus virgatus</taxonomicName>
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: Holotype: male: "Holotypus / Congo belge: P.N.U., R. Dipwa (1.900 m) 17-I-1948 Mis. G.F. de Witte, 1239a / Coll. Mus. Congo (ex. coll. I.P.N.C.B.) /
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<taxonomicName id="806C2B6B1090E1B41A583008E29ECAEE" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus virgatus" order="Coleoptera" pageId="171" pageNumber="172" phylum="Arthropoda" rank="species" species="virgatus">Laccophilus virgatus</taxonomicName>
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sp. n. Type, male symbol / F. Guignot det., 1952
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<taxonomicName id="8AB34149BA54E8E670CA68C87BAC774D" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus virgatus" order="Coleoptera" pageId="171" pageNumber="172" phylum="Arthropoda" rank="species" species="virgatus">Laccophilus virgatus</taxonomicName>
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Guign. Type, male symbol" (MRAC). - Paratype: "Congo belge: P.N.U., R. Dipwa (1.900 m) 17-I-1948 Mis. G.F. de Witte, 1242 / Paratype" (5 exs. IRSNB); same but
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<normalizedToken id="5DB9E965FBC83E669008CC570413C6C2" originalValue="“1293”">"1293"</normalizedToken>
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(1 ex. IRSNB); "Congo Belge P.N.U. Katobwe (Mukana 1810 m) 22-III-1947 Mis. G.F. de Witte: 92a / Paratype" (2 exs. IRSNB); "PNU Mukana 1810 m/12.3. 1947 F. de Witte / Paratype" (1 ex. MNHN); "Congo Belge: PNU Lusinga (Mukana) 28-V-1945 / Paratype / Guignot det., 1953:
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<taxonomicName id="C221E4FDFAD97D8D06D04DD9A46D5C30" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus virgatus" order="Coleoptera" pageId="171" pageNumber="172" phylum="Arthropoda" rank="species" species="virgatus">Laccophilus virgatus</taxonomicName>
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sp. n." (1 ex. IRSNB, habitus in Fig. 473); "PNU Lusinga (Galerie) / 22-25.5. 1945 G.F. de Witte / Paratype" (1 ex. MNHN).
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</paragraph>
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</subSubSection>
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<subSubSection id="5E4B4D2E84AEA946F14683EFA460076D" pageId="171" pageNumber="172" type="materials_examined">
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<paragraph id="10614558C1FB6A795A96C0079A108CBB" pageId="171" pageNumber="172">Additional material studied</paragraph>
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<paragraph id="5659A1E64CF4EAB76A01862D3B0ED982" pageId="171" pageNumber="172">
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(1 ex.). Zaire: "PNU Mukana 1810 m, 24.III. 1947 / F. Guignot det., 1953:
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<taxonomicName id="E6CA2091DF457F681FB8A28BBA46EE28" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus virgatus" order="Coleoptera" pageId="171" pageNumber="172" phylum="Arthropoda" rank="species" species="virgatus">Laccophilus virgatus</taxonomicName>
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sp. n." (1 ex. IRSNB). [Comment: labelled as paratype, but not listed as such in the original description.]
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</paragraph>
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</subSubSection>
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<subSubSection id="30B30BE26F59D841C836BA845AC9D38B" pageId="171" pageNumber="172" type="comments on synonymy">
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<paragraph id="A1E23DD503FDF71684F88FBEAAA95301" pageId="171" pageNumber="172">Comments on synonymy.</paragraph>
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<paragraph id="FDFF954564C0E111727F8A706ACEB21E" pageId="171" pageNumber="172">
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Holotypes of
|
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<taxonomicName id="007E536A7719AD3112173698296296C4" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus incrassatus" order="Coleoptera" pageId="171" pageNumber="172" phylum="Arthropoda" rank="species" species="incrassatus">Laccophilus incrassatus</taxonomicName>
|
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and
|
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|
<taxonomicName id="1F654716CCF56C94AEBBDD1A4A679F65" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus virgatus" order="Coleoptera" pageId="171" pageNumber="172" phylum="Arthropoda" rank="species" species="virgatus">Laccophilus virgatus</taxonomicName>
|
||
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|
have been examined and compared but no characteristics, justifying separation in two species were found. Accordingly, the two species are considered conspecific and
|
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<taxonomicName id="F33AC031482DDC814F3BF9AAED545F0A" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus incrassatus" order="Coleoptera" pageId="171" pageNumber="172" phylum="Arthropoda" rank="species" species="incrassatus">Laccophilus incrassatus</taxonomicName>
|
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, being older, is the valid name of the species.
|
||
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</paragraph>
|
||
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|
</subSubSection>
|
||
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|
<subSubSection id="E7AB0F9F220B635C2608F60E2E479F62" lastPageId="172" lastPageNumber="173" pageId="171" pageNumber="172" type="diagnosis">
|
||
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|
<paragraph id="3D0BE36CABEA7E0EF5B23D78AB9AAD5A" pageId="171" pageNumber="172">Diagnosis.</paragraph>
|
||
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<paragraph id="BCD1835E0391B77DC3322E67CFDC854D" lastPageId="172" lastPageNumber="173" pageId="171" pageNumber="172">
|
||
|
|
<taxonomicName id="37FA03B8CAD6CD3FE38BE910ADB1937B" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus incrassatus" order="Coleoptera" pageId="171" pageNumber="172" phylum="Arthropoda" rank="species" species="incrassatus">Laccophilus incrassatus</taxonomicName>
|
||
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|
resembles most of
|
||
|
|
<taxonomicName id="E831ECB6F178E532DCACE2817D35E371" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="171" pageNumber="172" phylum="Arthropoda" rank="species" species="cyclopis">Laccophilus cyclopis</taxonomicName>
|
||
|
|
,
|
||
|
|
<taxonomicName id="FCBFC6B2BCD7793D4A56FB4040C4296A" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus quindecimvittatus" order="Coleoptera" pageId="171" pageNumber="172" phylum="Arthropoda" rank="species" species="quindecimvittatus">Laccophilus quindecimvittatus</taxonomicName>
|
||
|
|
and
|
||
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|
<taxonomicName id="3F92F03975E0A047D0E6705EA2E596E5" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus empheres" order="Coleoptera" pageId="171" pageNumber="172" phylum="Arthropoda" rank="species" species="empheres">Laccophilus empheres</taxonomicName>
|
||
|
|
. From
|
||
|
|
<taxonomicName id="219F1474214C58D4BE9C50547B37B95F" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="171" pageNumber="172" phylum="Arthropoda" rank="species" species="cyclopis">Laccophilus cyclopis</taxonomicName>
|
||
|
|
,
|
||
|
|
<taxonomicName id="87E42FE5C815D787AFEE814E46249551" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus incrassatus" order="Coleoptera" pageId="171" pageNumber="172" phylum="Arthropoda" rank="species" species="incrassatus">Laccophilus incrassatus</taxonomicName>
|
||
|
|
is separated by the elytral reticulation, which is clearly double in
|
||
|
|
<taxonomicName id="92D6D6B56D9C1C8688E12788CCC68E75" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus cyclopis" order="Coleoptera" pageId="171" pageNumber="172" phylum="Arthropoda" rank="species" species="cyclopis">Laccophilus cyclopis</taxonomicName>
|
||
|
|
while simple or almost simple in
|
||
|
|
<taxonomicName id="AF4A2398F28BA6B0F247B2BF1C07D46C" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus incrassatus" order="Coleoptera" pageId="171" pageNumber="172" phylum="Arthropoda" rank="species" species="incrassatus">Laccophilus incrassatus</taxonomicName>
|
||
|
|
(if larger meshes are discerned they are always rather indistinct and reduced). From
|
||
|
|
<taxonomicName id="7FCFF64763645B9BC9F919B829E94AB3" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus quindecimvittatus" order="Coleoptera" pageId="171" pageNumber="172" phylum="Arthropoda" rank="species" species="quindecimvittatus">Laccophilus quindecimvittatus</taxonomicName>
|
||
|
|
,
|
||
|
|
<taxonomicName id="6D7C688971C5FB7735080ADCC81422E9" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus incrassatus" order="Coleoptera" pageId="171" pageNumber="172" phylum="Arthropoda" rank="species" species="incrassatus">Laccophilus incrassatus</taxonomicName>
|
||
|
|
is separated by having distinctly larger body (max. 4.2 mm in
|
||
|
|
<taxonomicName id="EF35A0D81765B7BA42D618E1C283A06F" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus quindecimvittatus" order="Coleoptera" pageId="171" pageNumber="172" phylum="Arthropoda" rank="species" species="quindecimvittatus">Laccophilus quindecimvittatus</taxonomicName>
|
||
|
|
). Shape of penis is also characteristic; apex broader in
|
||
|
|
<taxonomicName id="D65D698EA1BFC3A250312A8F4976BC58" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus incrassatus" order="Coleoptera" pageId="172" pageNumber="173" phylum="Arthropoda" rank="species" species="incrassatus">
|
||
|
|
<pageBreakToken id="51F5025E032EBDD20771D715FC908F3F" pageId="172" pageNumber="173" start="start">Laccophilus</pageBreakToken>
|
||
|
|
incrassatus
|
||
|
|
</taxonomicName>
|
||
|
|
. From
|
||
|
|
<taxonomicName id="8522C01B8E97FC829245ABEC61742A74" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus empheres" order="Coleoptera" pageId="172" pageNumber="173" phylum="Arthropoda" rank="species" species="empheres">Laccophilus empheres</taxonomicName>
|
||
|
|
,
|
||
|
|
<taxonomicName id="34C8C89B86BDCC049A1E83606D383E92" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus incrassatus" order="Coleoptera" pageId="172" pageNumber="173" phylum="Arthropoda" rank="species" species="incrassatus">Laccophilus incrassatus</taxonomicName>
|
||
|
|
is distinguished by differences detected in male genitalia; penis distinctly broader in
|
||
|
|
<taxonomicName id="9779C32E1E4F10E22C06C8A7FB17C06C" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus empheres" order="Coleoptera" pageId="172" pageNumber="173" phylum="Arthropoda" rank="species" species="empheres">Laccophilus empheres</taxonomicName>
|
||
|
|
.
|
||
|
|
</paragraph>
|
||
|
|
</subSubSection>
|
||
|
|
<subSubSection id="C711E1C31BA6C025E687D46BF357EDAC" pageId="172" pageNumber="173" type="description">
|
||
|
|
<paragraph id="32FA5598328A9A30AAE50C9563D9640A" pageId="172" pageNumber="173">Description.</paragraph>
|
||
|
|
<paragraph id="7C8F78D3D5011EBAF725920BB38AA73E" pageId="172" pageNumber="173">Body: Length 4.6-5.0 mm, width 2.7-2.8 mm. Pale ferrugineous with distinct blackish ferrugineous to dark ferrugineous colour pattern (Figs 472-473).</paragraph>
|
||
|
|
<paragraph id="39976C1FE85C4CE85FE42EF9849F9993" pageId="172" pageNumber="173">Head: Pale ferrugineous. Narrowly at pronotum with a dark ferrugineous, fairly well delimited marking. Submat, rather distinctly microsculptured. Reticulation double; meshes of two kinds and large meshes only slightly coarser than fine meshes. Large meshes discernible in medial part of head; posteriorly and anteriorly with simple reticulation. Large meshes, when discernible, contain two to almost ten small meshes. At eyes with some scattered, fine punctures extending towards centre of head.</paragraph>
|
||
|
|
<paragraph id="8CCFB1FFC37CF759333FB1D2BD978D4C" pageId="172" pageNumber="173">Pronotum: Pale ferrugineous, frontally with a quite broad, slightly vague dark ferrugineous marking; mediobasally with a rather narrow, blackish ferrugineous marking. Microsculpture dense and rather distinct; reticulation double. Large meshes only slightly more distinctly developed in comparison with fine meshes. Large meshes contain two to six fine meshes. Almost impunctate. Sparse and irregular, scattered punctures may be discerned.</paragraph>
|
||
|
|
<paragraph id="6FC0E679EC5EAFBE17D6372E848C6C19" pageId="172" pageNumber="173">Elytra: Pale ferrugineous, with distinct dark ferrugineous colour pattern; with longitudinal, blackish to dark ferrugineous markings, which exhibit some variation (Figs 472-473). Submat, rather densely microsculptured; reticulation almost of one kind (large meshes hardly discernible). Discal, dorsolateral and lateral rows of punctures discernible, but fine, irregular and sparse. Lateral, pre-apical furrow finely pubescent.</paragraph>
|
||
|
|
<paragraph id="622EE869806E0B69AFDCE7DFEBD4E1FB" pageId="172" pageNumber="173">Ventral aspect: Pale ferrugineous to ferrugineous, metacoxal plates laterally with a narrow blackish area. Metacoxal plates with transveresely located, very shallow furrows. Rather shiny, although, with very fine microsculpture. Abdomen with sparsely located, fine and somewhat curved striae. Almost impunctate. Prosternal process apex extended, slender and pointed. Apical ventrite with a lateral knob (Fig. 141).</paragraph>
|
||
|
|
<paragraph id="0C174992A4B765CD6D46A0334CFC50A2" pageId="172" pageNumber="173">Legs: Pro- and mesotarsus quite long and slender, provided with distinct suckers.</paragraph>
|
||
|
|
<paragraph id="9F421A5B9AE485F8040A35728F0D903C" pageId="172" pageNumber="173">Male genitalia: Penis comparatively long, slightly curved and extreme apex forms an angulate, medium broad process. External outline of penis provided with a distinct membranous area (Fig. 327).</paragraph>
|
||
|
|
<paragraph id="057066445812CBF4276A4E8DD52FC2B6" pageId="172" pageNumber="173">Female: Apical ventrite lacks knob (Fig. 142). Pro- and mesotarsus slender.</paragraph>
|
||
|
|
</subSubSection>
|
||
|
|
<subSubSection id="280EF610BA7CCB2C7F6104FA529CBE77" pageId="172" pageNumber="173" type="distribution">
|
||
|
|
<paragraph id="DE8C2C9FBAF776B80318C2D440E2D588" pageId="172" pageNumber="173">Distribution.</paragraph>
|
||
|
|
<paragraph id="503D137F374044A72576131A3584FE36" pageId="172" pageNumber="173">Zaire (Fig. 564). Not recorded from other African countries.</paragraph>
|
||
|
|
</subSubSection>
|
||
|
|
<subSubSection id="D6D2EC333225ED3480C3593953E4A222" pageId="172" pageNumber="173" type="collecting circumstances">
|
||
|
|
<paragraph id="AE61DB25A67BAAF6EE5AA4B5188B98CD" pageId="172" pageNumber="173">Collecting circumstances.</paragraph>
|
||
|
|
<paragraph id="000A815A49EE210037761614BABA128E" pageId="172" pageNumber="173">Unknown, not documented.</paragraph>
|
||
|
|
</subSubSection>
|
||
|
|
</treatment>
|
||
|
|
</document>
|