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<document id= "463B8211EB6A28CB693DBCE9C24DDC91" ID-DOI= "10.11646/phytotaxa.638.3.9" ID-ISSN= "1179-3163" ID-Zenodo-Dep= "13213312" IM.bibliography_approvedBy= "felipe" IM.illustrations_approvedBy= "felipe" IM.materialsCitations_approvedBy= "felipe" IM.tables_requiresApprovalFor= "GgImagineBatch" IM.taxonomicNames_approvedBy= "felipe" IM.treatments_approvedBy= "felipe" checkinTime= "1722602183299" checkinUser= "felipe" docAuthor= "Cho, Sung-Eun, Shin, Keumchul, Kim, Yong-Ho & Lee, Dong-Hyeon" docDate= "2024" docId= "4F1BB14CFFE9FFE763CBFF09FA2C6CBE" docLanguage= "en" docName= "phytotaxa.638.3.9.pdf" docOrigin= "Phytotaxa 638 (3)" docSource= "http://dx.doi.org/10.11646/phytotaxa.638.3.9" docStyle= "DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId= "96748F8F1B6C902996E134952A3A36B9" docStyleName= "Phytotaxa.2014-.journal_article" docStyleVersion= "13" docTitle= "Tubakia byeongjinii D. Hyeon Lee, S. E. Cho & K. C. Shin, sp. nov." docType= "treatment" docVersion= "2" lastPageNumber= "298" masterDocId= "B322C934FFEDFFE36343FFFDFF8E6B44" masterDocTitle= "Tubakia byeongjinii, a new species of Tubakiaceae, isolated from Quercus variabilis in South Korea" masterLastPageNumber= "300" masterPageNumber= "294" pageNumber= "298" updateTime= "1722803027052" updateUser= "ExternalLinkService" zenodo-license-figures= "UNSPECIFIED" zenodo-license-treatments= "UNSPECIFIED" >
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<mods:title id= "5823EEC4C616A93E9B6935E6E8FE63BA" > Tubakia byeongjinii, a new species of Tubakiaceae, isolated from Quercus variabilis in South Korea</mods:title>
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<mods:namePart id= "0934B221369DE9233AF6629FF312ED86" > Cho, Sung-Eun</mods:namePart>
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<mods:affiliation id= "92ED0E873E82066EB414E83A20E68106" > Institute of Agriculture and Life Science, Gyeongsang National University, Jinju 52828, South Korea.</mods:affiliation>
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<mods:namePart id= "1515290328671749507ECBF0F6BD550A" > Shin, Keumchul</mods:namePart>
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<mods:affiliation id= "ABECBDC1134B43D528CC4FEC06236952" > Institute of Agriculture and Life Science, Gyeongsang National University, Jinju 52828, South Korea. & Department of Forest Environmental Resources, Gyeongsang National University, Jinju 52828, South Korea.</mods:affiliation>
<mods:nameIdentifier id= "85BA651BD8AECBBA3CD4A89C5D8C7C15" type= "email" > kcshin@gnu.ac.kr</mods:nameIdentifier>
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<mods:namePart id= "9E914142D57EA539F1D2436EC1577929" > Kim, Yong-Ho</mods:namePart>
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<mods:namePart id= "A4FCCF5CCFA1BC069198A94CD0EDCE36" > Lee, Dong-Hyeon</mods:namePart>
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<treatment id= "4F1BB14CFFE9FFE763CBFF09FA2C6CBE" LSID= "urn:lsid:plazi:treatment:4F1BB14CFFE9FFE763CBFF09FA2C6CBE" httpUri= "http://treatment.plazi.org/id/4F1BB14CFFE9FFE763CBFF09FA2C6CBE" lastPageNumber= "298" pageId= "4" pageNumber= "298" >
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<taxonomicName id= "00B27BD9FFE9FFE763CBFF09FC926A4A" authority= "D. Hyeon Lee, S.E. Cho & K.C. Shin" authorityName= "D. Hyeon Lee, S. E. Cho & K. C. Shin" box= "[136,796,244,271]" class= "Sordariomycetes" family= "Tubakiaceae" genus= "Tubakia" kingdom= "Fungi" order= "Diaporthales" pageId= "4" pageNumber= "298" phylum= "Ascomycota" rank= "species" species= "byeongjinii" status= "sp. nov." >
<emphasis id= "F5C6DC48FFE9FFE763CBFF09FEE16A4B" bold= "true" box= "[136,367,244,271]" italics= "true" pageId= "4" pageNumber= "298" > Tubakia byeongjinii</emphasis>
D. Hyeon Lee, S.E. Cho & K.C. Shin
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,
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<taxonomicNameLabel id= "EEF56133FFE9FFE7606BFF08FCF36A4A" box= "[808,893,245,270]" pageId= "4" pageNumber= "298" rank= "species" > sp. nov.</taxonomicNameLabel>
</emphasis>
(
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<figureCitation id= "5F891CDFFFE9FFE760C8FF09FC416A4A" box= "[907,975,244,270]" captionStart= "FIGURE 2" captionStartId= "4.[136,229,1335,1357]" captionTargetBox= "[340,1247,619,1310]" captionTargetId= "figure-124@4.[340,1247,619,1310]" captionTargetPageId= "4" captionText= "FIGURE 2. Leaf spot on seedlings of Quercus variabilis caused by Tubakia byeongjinii. A. Symptoms on leaves. B. Scutellum and conidia in germination. C. Close-up view of symptoms and conidiomata. D. Top view of radiate scutellum. E. Conidia. Scale bar: 10 µm." figureDoi= "http://doi.org/10.5281/zenodo.13213316" httpUri= "https://zenodo.org/record/13213316/files/figure.png" pageId= "4" pageNumber= "298" > Fig. 2</figureCitation>
2024-08-04 22:55:04 +02:00
)
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<emphasis id= "F5C6DC48FFE9FFE763CBFEE5FEB66A76" box= "[136,312,280,306]" italics= "true" pageId= "4" pageNumber= "298" > MycoBank No.:</emphasis>
MB 851634
</paragraph>
</subSubSection>
<subSubSection id= "8FA853D1FFE9FFE763CBFE9DFE9A6ADB" pageId= "4" pageNumber= "298" type= "etymology" >
<paragraph id= "C70D005AFFE9FFE763CBFE9DFE9A6ADB" blockId= "4.[136,1452,352,594]" pageId= "4" pageNumber= "298" >
<emphasis id= "F5C6DC48FFE9FFE763CBFE9DFEA06A3E" bold= "true" box= "[136,302,352,378]" pageId= "4" pageNumber= "298" > Etymology:—</emphasis>
<taxonomicName id= "00B27BD9FFE9FFE7626DFE9DFE226A3E" box= "[302,428,352,378]" class= "Sordariomycetes" family= "Tubakiaceae" genus= "Tubakia" kingdom= "Fungi" order= "Diaporthales" pageId= "4" pageNumber= "298" phylum= "Ascomycota" rank= "species" species= "byeongjinii" >
<emphasis id= "F5C6DC48FFE9FFE7626DFE9DFE226A3E" box= "[302,428,352,378]" italics= "true" pageId= "4" pageNumber= "298" > byeongjinii</emphasis>
</taxonomicName>
is given in honor of Professor Byeongjin Cha, for his contributions as a tree pathologist in
<collectingCountry id= "BFA540CAFFE9FFE763CBFE79FE9A6ADB" box= "[136,276,388,415]" name= "South Korea" pageId= "4" pageNumber= "298" > South Korea</collectingCountry>
</paragraph>
</subSubSection>
<subSubSection id= "8FA853D1FFE9FFE763FEFE55FF7B6C65" pageId= "4" pageNumber= "298" type= "materials_examined" >
<paragraph id= "C70D005AFFE9FFE763FEFE55FED86916" blockId= "4.[136,1452,352,594]" pageId= "4" pageNumber= "298" >
<emphasis id= "F5C6DC48FFE9FFE763FEFE55FE906A86" bold= "true" box= "[189,286,424,450]" pageId= "4" pageNumber= "298" > Type:—</emphasis>
<collectingCountry id= "BFA540CAFFE9FFE7625EFE55FE696A86" box= "[285,487,424,450]" name= "South Korea" pageId= "4" pageNumber= "298" > SOUTH KOREA</collectingCountry>
. Sejong-si,
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<geoCoordinate id= "A286669DFFE9FFE7604DFE55FC3A6A86" box= "[782,948,424,450]" degrees= "127" direction= "east" minutes= "17" orientation= "longitude" pageId= "4" pageNumber= "298" precision= "1" seconds= "41.8" value= "127.294945" > 127°17'41.8" E</geoCoordinate>
, isolated from leaves on
<taxonomicName id= "00B27BD9FFE9FFE76790FE55FA2A6A86" box= "[1235,1444,424,450]" class= "Magnoliopsida" family= "Fagaceae" genus= "Quercus" kingdom= "Plantae" order= "Fagales" pageId= "4" pageNumber= "298" phylum= "Tracheophyta" rank= "species" species= "variabilis" >
<emphasis id= "F5C6DC48FFE9FFE76790FE55FA2A6A86" box= "[1235,1444,424,450]" italics= "true" pageId= "4" pageNumber= "298" > Quercus variabilis</emphasis>
</taxonomicName>
,
<date id= "B30C269AFFE9FFE763CBFE31FEE86AA2" box= "[136,358,460,486]" pageId= "4" pageNumber= "298" value= "2020-09-28" > 28 September 2020</date>
. The
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,
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23-0901, a dried culture of
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041 was deposited in the herbarium collection (
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) of Korea National Arboretum. The ex-holotype culture (
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041=
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49779) was deposited in the culture collection of the National Institute of Forest Science (
<collectionCode id= "A1A3989FFFE9FFE761B8FDE9FCB7696A" box= "[763,825,532,558]" pageId= "4" pageNumber= "298" > CDH</collectionCode>
) and Korean Agricultural Culture Collection for Type Cultures (
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).
</paragraph>
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2024-08-04 22:55:04 +02:00
<paragraph id= "C70D005AFFE9FFE763CBFACAFAEE6E34" blockId= "4.[136,1452,1335,1393]" pageId= "4" pageNumber= "298" >
<emphasis id= "F5C6DC48FFE9FFE763CBFACAFF726E08" bold= "true" box= "[136,252,1335,1357]" pageId= "4" pageNumber= "298" > FIGURE 2.</emphasis>
Leaf spot on seedlings of
<taxonomicName id= "00B27BD9FFE9FFE762B1FACAFD2D6E08" box= "[498,675,1335,1356]" class= "Magnoliopsida" family= "Fagaceae" genus= "Quercus" kingdom= "Plantae" order= "Fagales" pageId= "4" pageNumber= "298" phylum= "Tracheophyta" rank= "species" species= "variabilis" >
<emphasis id= "F5C6DC48FFE9FFE762B1FACAFD2D6E08" box= "[498,675,1335,1356]" italics= "true" pageId= "4" pageNumber= "298" > Quercus variabilis</emphasis>
</taxonomicName>
caused by
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<emphasis id= "F5C6DC48FFE9FFE76049FACAFC466E08" box= "[778,968,1335,1356]" italics= "true" pageId= "4" pageNumber= "298" > Tubakia byeongjinii</emphasis>
</taxonomicName>
. A. Symptoms on leaves. B. Scutellum and conidia in germination. C. Close-up view of symptoms and conidiomata. D. Top view of radiate scutellum. E. Conidia. Scale bar: 10 µm.
</paragraph>
</caption>
<paragraph id= "C70D005AFFE9FFE763FEFA62FC7B6E99" blockId= "4.[136,1452,1439,2042]" pageId= "4" pageNumber= "298" >
<emphasis id= "F5C6DC48FFE9FFE763FEFA62FDD26EFD" bold= "true" box= "[189,604,1439,1465]" pageId= "4" pageNumber= "298" > Additional specimen examined:—</emphasis>
<collectingCountry id= "BFA540CAFFE9FFE7611FFA62FCA76EFD" box= "[604,809,1439,1465]" name= "South Korea" pageId= "4" pageNumber= "298" > SOUTH KOREA</collectingCountry>
. Sejong-si,
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<geoCoordinate id= "A286669DFFE9FFE76718FA62FA8F6EFD" box= "[1115,1281,1439,1465]" degrees= "127" direction= "east" minutes= "17" orientation= "longitude" pageId= "4" pageNumber= "298" precision= "1" seconds= "41.8" value= "127.294945" > 127°17'41.8" E</geoCoordinate>
, isolated from leaves on
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<emphasis id= "F5C6DC48FFE9FFE763BBFA39FE466E99" box= "[248,456,1475,1502]" italics= "true" pageId= "4" pageNumber= "298" > Quercus variabilis</emphasis>
</taxonomicName>
,
<date id= "B30C269AFFE9FFE76295FA3EFD3D6E99" box= "[470,691,1475,1501]" pageId= "4" pageNumber= "298" value= "2020-09-28" > 28 September 2020</date>
, D.
<collectionCode id= "A1A3989FFFE9FFE7619AFA39FD7F6E9A" box= "[729,753,1476,1502]" country= "Finland" lsid= "urn:lsid:biocol.org:col:15618" name= "University of Helsinki" pageId= "4" pageNumber= "298" type= "Herbarium" > H</collectionCode>
. Lee, culture
<collectionCode id= "A1A3989FFFE9FFE760CBFA3EFC4B6E99" box= "[904,965,1475,1501]" pageId= "4" pageNumber= "298" > CDH</collectionCode>
040.
</paragraph>
<paragraph id= "C70D005AFFE9FFE763FEFA1AFCF26D46" blockId= "4.[136,1452,1439,2042]" box= "[189,892,1511,1538]" pageId= "4" pageNumber= "298" >
<emphasis id= "F5C6DC48FFE9FFE763FEFA1AFEB06D45" bold= "true" box= "[189,318,1511,1537]" pageId= "4" pageNumber= "298" > Habitat:—</emphasis>
On leaves of
<taxonomicName id= "00B27BD9FFE9FFE76290FA15FD2D6D45" box= "[467,675,1511,1538]" class= "Magnoliopsida" family= "Fagaceae" genus= "Quercus" kingdom= "Plantae" order= "Fagales" pageId= "4" pageNumber= "298" phylum= "Tracheophyta" rank= "species" species= "variabilis" >
<emphasis id= "F5C6DC48FFE9FFE76290FA15FD2D6D45" box= "[467,675,1511,1538]" italics= "true" pageId= "4" pageNumber= "298" > Quercus variabilis</emphasis>
</taxonomicName>
from
<collectingCountry id= "BFA540CAFFE9FFE761ABFA1AFCF66D46" box= "[744,888,1511,1538]" name= "South Korea" pageId= "4" pageNumber= "298" > South Korea</collectingCountry>
.
</paragraph>
<paragraph id= "C70D005AFFE9FFE763FEF9F6FF7B6C65" blockId= "4.[136,1452,1439,2042]" pageId= "4" pageNumber= "298" >
<emphasis id= "F5C6DC48FFE9FFE763FEF9F6FEAF6D61" box= "[189,289,1547,1573]" italics= "true" pageId= "4" pageNumber= "298" > Colonies</emphasis>
on MEA attaining diameter of
<quantity id= "004AADBFFFE9FFE7613EF9F6FD426D62" box= "[637,716,1547,1574]" metricMagnitude= "-2" metricUnit= "m" metricValue= "6.5" pageId= "4" pageNumber= "298" unit= "mm" value= "65.0" > 65 mm</quantity>
in 10 days, initially white with dense aerial mycelium, spreading, smooth, flat, rapidly turning to olivaceous black, with dark green at 10 days, developing concentric rings of dense mycelium leading to irregular margins.
<emphasis id= "F5C6DC48FFE9FFE76113F9AEFD6C6D29" box= "[592,738,1619,1645]" italics= "true" pageId= "4" pageNumber= "298" > Conidiomata</emphasis>
(pycnothyria) superficial, epiphyllous on leaf, radiate scutella 75–
<emphasis id= "F5C6DC48FFE9FFE762EDF966FDB26DF1" box= "[430,572,1691,1717]" italics= "true" pageId= "4" pageNumber= "298" > Sporodochia</emphasis>
superficial with only a few radiating dark hyphae, mainly hypophyllous, on necrotic interveinal tissues and along leaf veins.
<emphasis id= "F5C6DC48FFE9FFE761E2F942FCC86D9D" box= "[673,838,1727,1753]" italics= "true" pageId= "4" pageNumber= "298" > Conidiophores</emphasis>
on underside of scutella.
<emphasis id= "F5C6DC48FFE9FFE76722F942FB356D9D" box= "[1121,1211,1727,1753]" italics= "true" pageId= "4" pageNumber= "298" > Conidia</emphasis>
hyaline, turning light brown with age, smooth to slightly varicose, aseptate, obovoid to ovoid, 7.7– × –
<emphasis id= "F5C6DC48FFE9FFE767ABF91EFAF16DB9" box= "[1256,1407,1763,1789]" italics= "true" pageId= "4" pageNumber= "298" > Microconidia</emphasis>
not observed.
</paragraph>
</subSubSection>
<subSubSection id= "8FA853D1FFE9FFE763FEF8D6FA2C6CBE" pageId= "4" pageNumber= "298" type= "discussion" >
<paragraph id= "C70D005AFFE9FFE763FEF8D6FD006CC9" blockId= "4.[136,1452,1439,2042]" pageId= "4" pageNumber= "298" >
<emphasis id= "F5C6DC48FFE9FFE763FEF8D6FEEA6C01" bold= "true" box= "[189,356,1835,1861]" pageId= "4" pageNumber= "298" > Comments:—</emphasis>
<taxonomicName id= "00B27BD9FFE9FFE76227F8D6FDCB6C01" box= "[356,581,1835,1861]" class= "Sordariomycetes" family= "Tubakiaceae" genus= "Tubakia" kingdom= "Fungi" order= "Diaporthales" pageId= "4" pageNumber= "298" phylum= "Ascomycota" rank= "species" species= "byeongjinii" >
<emphasis id= "F5C6DC48FFE9FFE76227F8D6FDCB6C01" box= "[356,581,1835,1861]" italics= "true" pageId= "4" pageNumber= "298" > Tubakia byeongjinii</emphasis>
</taxonomicName>
is phylogenetically closely related
<taxonomicName id= "00B27BD9FFE9FFE76098F8D1FBD46C01" baseAuthorityName= "Saccardo" baseAuthorityYear= "1913" box= "[987,1114,1836,1861]" class= "Sordariomycetes" family= "Tubakiaceae" genus= "Actinopelte" kingdom= "Fungi" order= "Diaporthales" pageId= "4" pageNumber= "298" phylum= "Ascomycota" rank= "species" species= "japonica" >
<emphasis id= "F5C6DC48FFE9FFE76098F8D1FBD46C01" box= "[987,1114,1836,1861]" italics= "true" pageId= "4" pageNumber= "298" > T. japonica</emphasis>
</taxonomicName>
. However,
<taxonomicName id= "00B27BD9FFE9FFE767A3F8D1FAF56C01" box= "[1248,1403,1835,1861]" class= "Sordariomycetes" family= "Tubakiaceae" genus= "Tubakia" kingdom= "Fungi" order= "Diaporthales" pageId= "4" pageNumber= "298" phylum= "Ascomycota" rank= "species" species= "byeongjinii" >
<emphasis id= "F5C6DC48FFE9FFE767A3F8D1FAF56C01" box= "[1248,1403,1835,1861]" italics= "true" pageId= "4" pageNumber= "298" > T. byeongjinii</emphasis>
</taxonomicName>
can be distinguished by relatively smaller pycnothyria and conidia compared to those that can be observed in
<taxonomicName id= "00B27BD9FFE9FFE7666EF8ADFA256C2D" baseAuthorityName= "Saccardo" baseAuthorityYear= "1913" box= "[1325,1451,1872,1897]" class= "Sordariomycetes" family= "Tubakiaceae" genus= "Actinopelte" kingdom= "Fungi" order= "Diaporthales" pageId= "4" pageNumber= "298" phylum= "Ascomycota" rank= "species" species= "japonica" >
<emphasis id= "F5C6DC48FFE9FFE7666EF8ADFA256C2D" box= "[1325,1451,1872,1897]" italics= "true" pageId= "4" pageNumber= "298" > T. japonica</emphasis>
</taxonomicName>
(
<bibRefCitation id= "A3237DABFFE9FFE763D3F88EFEB46CC9" author= "Saccardo, P. A." box= "[144,314,1907,1933]" pageId= "4" pageNumber= "298" pagination= "312 - 325" refId= "ref3882" refString= "Saccardo, P. A. (1913) Notae mycologicae. Annales Mycologici 11: 312 - 325." type= "journal article" year= "1913" > Saccardo 1913</bibRefCitation>
,
<bibRefCitation id= "A3237DABFFE9FFE76206F88EFD0C6CC9" author= "Harrington, T. C. & McNew, D. L." box= "[325,642,1907,1934]" pageId= "4" pageNumber= "298" pagination= "1003 - 1022" refId= "ref3459" refString= "Harrington, T. C. & McNew, D. L. (2018) A re-evaluation of Tubakia, including three new species on Quercus and six new combinations. Antonie van Leeuwenhoek 111: 1003 - 1022. https: // doi. org / 10.1007 / s 10482 - 017 - 1001 - 9" type= "journal article" year= "2018" > Harrington & McNew 2018</bibRefCitation>
).
</paragraph>
<paragraph id= "C70D005AFFE9FFE763FEF86AFA2C6CBE" blockId= "4.[136,1452,1439,2042]" pageId= "4" pageNumber= "298" >
Based on morphology and phylogenetic analysis, the species of
<taxonomicName id= "00B27BD9FFE9FFE76032F86AFC456CF5" box= "[881,971,1943,1969]" class= "Sordariomycetes" family= "Tubakiaceae" genus= "Tubakia" kingdom= "Fungi" order= "Diaporthales" pageId= "4" pageNumber= "298" phylum= "Ascomycota" rank= "genus" >
<emphasis id= "F5C6DC48FFE9FFE76032F86AFC456CF5" box= "[881,971,1943,1969]" italics= "true" pageId= "4" pageNumber= "298" > Tubakia</emphasis>
</taxonomicName>
found on
<taxonomicName id= "00B27BD9FFE9FFE76775F865FA8D6CF5" box= "[1078,1283,1943,1970]" class= "Magnoliopsida" family= "Fagaceae" genus= "Quercus" kingdom= "Plantae" order= "Fagales" pageId= "4" pageNumber= "298" phylum= "Tracheophyta" rank= "species" species= "variabilis" >
<emphasis id= "F5C6DC48FFE9FFE76775F865FA8D6CF5" box= "[1078,1283,1943,1970]" italics= "true" pageId= "4" pageNumber= "298" > Quercus variabilis</emphasis>
</taxonomicName>
in
<collectingCountry id= "BFA540CAFFE9FFE76662F86AFA256CF6" box= "[1313,1451,1943,1970]" name= "South Korea" pageId= "4" pageNumber= "298" > South Korea</collectingCountry>
is distinct from all previously described species of
<taxonomicName id= "00B27BD9FFE9FFE761FCF846FC976C91" box= "[703,793,1979,2005]" class= "Sordariomycetes" family= "Tubakiaceae" genus= "Tubakia" kingdom= "Fungi" order= "Diaporthales" pageId= "4" pageNumber= "298" phylum= "Ascomycota" rank= "genus" >
<emphasis id= "F5C6DC48FFE9FFE761FCF846FC976C91" box= "[703,793,1979,2005]" italics= "true" pageId= "4" pageNumber= "298" > Tubakia</emphasis>
</taxonomicName>
. In addition to its unique morphological characteristics, its phylogenetic position supports its classification as a novel species of
<taxonomicName id= "00B27BD9FFE9FFE760D3F822FC646CBD" box= "[912,1002,2015,2041]" class= "Sordariomycetes" family= "Tubakiaceae" genus= "Tubakia" kingdom= "Fungi" order= "Diaporthales" pageId= "4" pageNumber= "298" phylum= "Ascomycota" rank= "genus" >
<emphasis id= "F5C6DC48FFE9FFE760D3F822FC646CBD" box= "[912,1002,2015,2041]" italics= "true" pageId= "4" pageNumber= "298" > Tubakia</emphasis>
</taxonomicName>
, proposed here as
<taxonomicName id= "00B27BD9FFE9FFE767FFF822FA156CBD" box= "[1212,1435,2015,2041]" class= "Sordariomycetes" family= "Tubakiaceae" genus= "Tubakia" kingdom= "Fungi" order= "Diaporthales" pageId= "4" pageNumber= "298" phylum= "Ascomycota" rank= "species" species= "byeongjinii" >
<emphasis id= "F5C6DC48FFE9FFE767FFF822FA156CBD" box= "[1212,1435,2015,2041]" italics= "true" pageId= "4" pageNumber= "298" > Tubakia byeongjinii</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
