<documentid="5DA3CEA2DC781BBB748B4FFD741C6EE9"ID-CLB-Dataset="25754"ID-DOI="http://dx.doi.org/10.3897/zookeys.58.507"ID-GBIF-Dataset="1c4e9564-16cc-41fd-bdd2-d4a77484f100"ID-PMC="PMC3088339"ID-Pensoft-Pub="1313-2970-58-1"ID-PubMed="21594191"ModsDocAuthor=""ModsDocDate="2010"ModsDocID="1313-2970-58-1"ModsDocOrigin="ZooKeys 58"ModsDocTitle="Australian gall-inducing scale insects on Eucalyptus: revision of Opisthoscelis Schrader (Coccoidea, Eriococcidae) and descriptions of a new genus and nine new species"checkinTime="1451250974324"checkinUser="pensoft"docAuthor="Hardy, Nate B. & Gullan, Penny J."docDate="2010"docId="5FD984E7BB8870FD836C6DA0CD122A9F"docLanguage="en"docName="ZooKeys 58: 1-74"docOrigin="ZooKeys 58"docSource="http://dx.doi.org/10.3897/zookeys.58.507"docTitle="Tanyscelis verrucula, comb. n."docType="treatment"docVersion="6"lastPageNumber="41"masterDocId="8968FFC5A95F7F195B2A10735E18FFA1"masterDocTitle="Australian gall-inducing scale insects on Eucalyptus: revision of Opisthoscelis Schrader (Coccoidea, Eriococcidae) and descriptions of a new genus and nine new species"masterLastPageNumber="74"masterPageNumber="1"pageNumber="39"updateTime="1732685506351"updateUser="ExternalLinkService">
<mods:titleid="35F5B9B82AAA2CA7CF12D953F6AA1745">Australian gall-inducing scale insects on Eucalyptus: revision of Opisthoscelis Schrader (Coccoidea, Eriococcidae) and descriptions of a new genus and nine new species</mods:title>
<bibRefCitationid="25E09ED8833A656FF66B2FCC9A091D8E"author="Froggatt, WW"journalOrPublisher="Proceedings of the Linnean Society of New South Wales"pageId="44"pageNumber="45"pagination="335 - 348"title="Notes on the family Brachyscelidae, with descriptions of new species. Part III."volume="8"year="1894 b">Froggatt 1894b: 338-339</bibRefCitation>
<bibRefCitationid="7A31055ABDE6D57EA22B06998928CA4A"author="Froggatt, WW"journalOrPublisher="Proceedings of the Linnean Society of New South Wales"pageId="44"pageNumber="45"pagination="335 - 348"title="Notes on the family Brachyscelidae, with descriptions of new species. Part III."volume="8"year="1894 b">Froggatt 1894b: 344</bibRefCitation>
<bibRefCitationid="9DB6C5FACF44947D19BD4FF139CA79AD"author="Froggatt, WW"journalOrPublisher="Proceedings of the Linnean Society of New South Wales"pageId="44"pageNumber="45"pagination="335 - 348"title="Notes on the family Brachyscelidae, with descriptions of new species. Part III."volume="8"year="1894 b">Froggatt (1894b)</bibRefCitation>
described them as distinct species on different pages of the same article, but the two are identical based on the morphology of the adult females and their galls. Froggatt claimed that the adult female of
<bibRefCitationid="813B905DB86A386A57F3E1604DBEFC51"author="Froggatt, WW"journalOrPublisher="Proceedings of the Linnean Society of New South Wales"pageId="44"pageNumber="45"pagination="335 - 348"title="Notes on the family Brachyscelidae, with descriptions of new species. Part III."volume="8"year="1894 b">Froggatt 1894b: 338</bibRefCitation>
<bibRefCitationid="746416BB21D99062B46BC4EDA47FBC5B"author="Froggatt, WW"journalOrPublisher="Proceedings of the Linnean Society of New South Wales"pageId="44"pageNumber="45"pagination="335 - 348"title="Notes on the family Brachyscelidae, with descriptions of new species. Part III."volume="8"year="1894 b">Froggatt 1894b: 344</bibRefCitation>
). However all specimens that we examined had one pair of large spines, each with a small spine near its base, on abdominal segment VIII, and one pair of medium-sized spines on abdominal segment VII, with the latter sometimes appearing very close to the spines on segment VIII. The type locality of
(Fig. 3g,h). Female (Fig. 3g). On stem, leaf petiole and midrib. Gall nipple-like, broadly rugose, shape somewhat irregular, with orifice on opposite side of leaf; height 3.8-8.7 mm, width 2.7-6.4 mm, length of basal attachment 3.2-6.7 mm. Gall
<paragraphid="0DBAD2984AC594A822B3C5905E5260A4"pageId="39"pageNumber="40">Male (Fig. 3h). On stem and leaf (probably either surface). Height 1.5-5.0 mm, width 0.7-3.0 mm, length of basal attachment 1.1-4.2 mm. Gall cylindrical to conical, surface with shrivelled appearance, opening round to oblong, 0.1-1.3 mm wide, with opposite side of leaf swollen.</paragraph>
(Fig. 22) (n = 51). Body turbinate, margin incised at intersegmental boundaries, length 1.5-4.0 mm, greatest width 0.8-2.1 mm; abdomen tapered, about as long as head + thorax, extending far beyond femur. Eyespots each 30-65 mm wide, highly convex, on dorsal margin. Antennal segmentation poor, ca 3-segmented; each antenna 50-229 mm long. Frontal lobes difficult to discern, each 120-360
wide. Tentorial box 230-550 mm long. Labium 50-120 mm long, 60-140 mm wide. Spiracles 75-150 mm long, 35-85 mm wide across atrium. Fore and mid legs small sclerotic protuberances, 13-80
long; translucent pores scattered on both surfaces of tibia, plus a few on tarsus; trochanter with 2 campaniform sensilla on each side; femur-tibia articulation non-functional, tibia fixed in orientation almost parallel to long axis of femur; claw and digitules present but reduced. Anal opening 8-35
wide, sclerotisation of ring frequently uneven, weaker at posteroventral end, usually appearing horseshoe-shaped; anal ring with 4-6 minute setae. Anal area with 2 fleshy protuberances on each side of body, 1 laterad of anal ring, 1 posterolateral of anal ring, each protuberance terminating in stout spine, 20-35
Dorsum. Derm membranous. Dorsal setae minute, 5-8 mm long; arranged in an irregular transverse row across each abdominal segment, scattered over surface of head and thorax. Stout spinose setae, 5-10
long, found on margin of each posterior abdominal segment plus along margin of head and prothorax. Macrotubular ducts 13-15 mm long, dermal orifice with rim 5 mm wide; in transverse row across each abdominal segment, scattered over head and thorax. Microtubular ducts absent. Quinquelocular pores 5-7
Venter. Membranous lobe-like protuberance immediately posterior of each spiracle, each protuberance having a few slender setae. Setae flagellate, 30-118 mm long, in a transverse row or narrow band across each abdominal segment, and scattered along margin of head and thorax and mesad to each metacoxa. Macrotubular ducts absent. Quinquelocular pores similar to those on dorsum, in transverse band across each of abdominal segments
(here designated): AUSTRALIA: New South Wales: 1 adult female (3.27 mm long, 2.14 mm wide): ex dry gall on leaf, with printed label: "No. 1788 E / GALL MAKING COCCIDS.
Paralectotypes: AUSTRALIA: New South Wales: 2 adult females, 1 second-instar female: same data as lectotype (ASCU); 2 adult females, numerous first-instar nymphs 5 adult males: ex leaf galls,
sp., Bathurst, ex WWF coll. (BMNH). The latter BMNH specimens are deemed to be paralectotypes because the collection is from the type locality and Froggatt (1894b, 1921) only ever recorded the species from this one locality.
(here designated): AUSTRALIA: Victoria: 1 adult female (2.83 mm long, 2.04 mm wide): ex dry gall on leaf, with printed label: "No. 1789 E / GALL MAKING COCCIDS. /
<paragraphid="B8EF66F2C0D1BBA1BAD7808B50FF1E0B"pageId="40"pageNumber="41">Paralectotypes: AUSTRALIA: Victoria: 2 adult females, 6 slides with hundreds of first-instar nymphs: same data as lectotype (ASCU).</paragraph>
sp., Tallong cementery, -34.72°; 150.08°, 16 Oct., 1993, PJG (ANIC); Northern Territory: 1 adult female: Eucalyptus sp. N of Alice Springs, near Todd River, 19 Nov, 1978, M. Kotzman (ANIC). South Australia: 18 adult females, 8 adult males: ex galls,
sp., Para Wirra National Park, 31 Dec., 1965 [label of several slides erroneously records date as 1 Dec., 1965], HMB, Boratynski #1743 (BMNH). Tasmania: 1 adult female:
, ca 20 km E of Bendigo, near Axedale Flora and Fauna Reserve, roadside on Calder Highway, -36.78°; 144.49°, 3 Feb., 2005, PJG and NBH, NH73 (ANIC); 15 adult females: ex galls,
, Cranbourne, Royal Botanic Gardens, ca 150 m SE of Trig Point Lookout, -38.13°; 145.28°, 9 Feb., 2005, PJG, NH52, NH79 (ANIC, NMV); 300 first-instar nymphs, ex gall cavity,
, the body of the females shrunk to about one-fifth of their size prior to oviposition. It was estimated that a single female could lay more than 1000 eggs.
