265 lines
20 KiB
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265 lines
20 KiB
XML
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<document ID-DOI="http://dx.doi.org/10.3897/zse.92.9948" ID-GBIF-Taxon="182236996" ID-Pensoft-Pub="1860-0743-2-163" ID-Pensoft-UUID="FFCC824A3E5F060CFFF2FF83B964FFD2" ID-Zenodo-Dep="155320" ID-ZooBank="9DA068DAB881409199FE252D31DDC7D1" ModsDocID="1860-0743-92-2-163" checkinTime="1474970821929" checkinUser="pensoft" docAuthor="Conradie, Werner, Bittencourt-Silva, Gabriela B., Engelbrecht, Hanlie M., Loader, Simon P., Menegon, Michele, Nanvonamuquitxo, Cristovao, Scott, Michael & Tolley, Krystal A." docDate="2016" docId="E31BE66825E4596E86EDD409FBF1878F" docLanguage="en" docName="ZoosystEvol 92(2): 163-180" docOrigin="Zoosystematics and Evolution 92 (2)" docPubDate="2016-09-26" docSource="http://dx.doi.org/10.3897/zse.92.9948" docTitle="Nothophryne broadleyi Poynton 1963" docType="treatment" docVersion="2" id="FFCC824A3E5F060CFFF2FF83B964FFD2" lastPageId="6" lastPageNumber="169" masterDocId="FFCC824A3E5F060CFFF2FF83B964FFD2" masterDocTitle="Exploration into the hidden world of Mozambique's sky island forests: new discoveries of reptiles and amphibians" masterLastPageNumber="180" masterPageNumber="163" pageId="5" pageNumber="168" updateTime="1643495415339" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Exploration into the hidden world of Mozambique's sky island forests: new discoveries of reptiles and amphibians</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Conradie, Werner</mods:namePart>
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<mods:affiliation>Port Elizabeth Museum (Bayworld), P. O. Box 13147, Humewood 6013, South Africa & South African Institute for Aquatic Biodiversity, P / Bag 1015, Grahamstown, 6140, South Africa</mods:affiliation>
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<mods:nameIdentifier type="email">werner@bayworld.co.za</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bittencourt-Silva, Gabriela B.</mods:namePart>
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<mods:affiliation>University of Basel, Biogeography Research Group, Department of Environmental Sciences, Basel 4056, Switzerland</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Engelbrecht, Hanlie M.</mods:namePart>
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<mods:affiliation>South African National Biodiversity Institute, Private Bag X 7, Claremont, 7735, South Africa & Department of Botany and Zoology, Stellenbosch University, Matieland 7602, Stellenbosch, South Africa</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Loader, Simon P.</mods:namePart>
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<mods:affiliation>University of Roehampton, Department of Life Sciences, London, SW 15 4 JD, United Kingdom</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Menegon, Michele</mods:namePart>
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<mods:affiliation>MUSE, Museo delle Scienze, Viale del Lavoro e delle Scienza, 3 Trento 38122, Italy</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Nanvonamuquitxo, Cristovao</mods:namePart>
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<mods:affiliation>Faculty of Natural Sciences, Lurio University, Pemba, 958, Mozambique</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Scott, Michael</mods:namePart>
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<mods:affiliation>Khangela Safaris, www. khangelasafaris. com, Bulawayo, Zimbabwe</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tolley, Krystal A.</mods:namePart>
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<mods:affiliation>South African National Biodiversity Institute, Private Bag X 7, Claremont, 7735, South Africa & Department of Botany and Zoology, Stellenbosch University, Matieland 7602, Stellenbosch, South Africa</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zoosystematics and Evolution</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2016</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2016-09-26</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>92</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>163</mods:start>
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<mods:end>180</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zse.92.9948</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.92.9948</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1860-0743-2-163</mods:identifier>
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<mods:identifier type="ZooBank">9DA068DAB881409199FE252D31DDC7D1</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FFCC824A3E5F060CFFF2FF83B964FFD2</mods:identifier>
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<mods:identifier type="Zenodo-Dep">155320</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="182236996" LSID="urn:lsid:plazi:treatment:E31BE66825E4596E86EDD409FBF1878F" httpUri="http://treatment.plazi.org/id/E31BE66825E4596E86EDD409FBF1878F" lastPageId="6" lastPageNumber="169" pageId="5" pageNumber="168">
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<subSubSection pageId="5" pageNumber="168" type="nomenclature">
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<paragraph pageId="5" pageNumber="168">
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<taxonomicName LSID="E31BE668-25E4-596E-86ED-D409FBF1878F" authority="Poynton, 1963" authorityName="Poynton" authorityYear="1963" class="Amphibia" family="Pyxicephalidae" genus="Nothophryne" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothophryne broadleyi" order="Anura" pageId="5" pageNumber="168" phylum="Chordata" rank="species" species="broadleyi">Nothophryne cf. broadleyi Poynton, 1963</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="6" lastPageNumber="169" pageId="5" pageNumber="168" type="materials_examined">
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<paragraph pageId="5" pageNumber="168">Material</paragraph>
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<paragraph lastPageId="6" lastPageNumber="169" pageId="5" pageNumber="168">
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<materialsCitation accessionNumber="A11320, A11321, A11322, A11323, A11324, A11325, A11326, A11327, A11328, A11329, A11330, A11331, A11332, A11333, A11334, A11335, A11336, A11337, A11338, A11339, A11340, A11342, A11343, A11344, A11345, A11346, A11347, A11369, A11370, A11371, A11372" collectingDate="2016-01-01" collectingDateMax="2016-12-31" collectingDateMin="2016-01-01" location="Mt. Namuli" specimenCount="1">
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<location LSID="urn:lsid:plazi:treatment:E31BE66825E4596E86EDD409FBF1878F:7294921934D7F725E19C12C08D4C5259" name="Mt. Namuli">Mt. Namuli</location>
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(PEM
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/A11320">A11320</accessionNumber>
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, male,
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<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.7899999999999998" unit="mm" value="17.9">17.9 mm</quantity>
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SUL; PEM
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/A11321">A11321</accessionNumber>
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, female,
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<quantity metricMagnitude="-2" metricUnit="m" metricValue="2.27" unit="mm" value="22.7">22.7 mm</quantity>
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SUL; PEM
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/A11322">A11322</accessionNumber>
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, male,
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<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.81" unit="mm" value="18.1">18.1 mm</quantity>
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SUL; PEM
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/A11322">A11322</accessionNumber>
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, PEM
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/A11323">A11323</accessionNumber>
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, female,
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<quantity metricMagnitude="-2" metricUnit="m" metricValue="2.34" unit="mm" value="23.4">23.4 mm</quantity>
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SUL; PEM
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/A11324">A11324</accessionNumber>
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, male, 18.0 mm SUL; PEM
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/A11325">A11325</accessionNumber>
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, juvenile, 19.0 mm SUL; PEM
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/A11326">A11326</accessionNumber>
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, male,
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<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.75" unit="mm" value="17.5">17.5 mm</quantity>
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SUL; PEM
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/A11327">A11327</accessionNumber>
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, male,
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<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.8699999999999999" unit="mm" value="18.7">18.7 mm</quantity>
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SUL; PEM
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/A11328">A11328</accessionNumber>
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, female,
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<quantity metricMagnitude="-2" metricUnit="m" metricValue="2.53" unit="mm" value="25.3">25.3 mm</quantity>
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SUL; PEM
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/A11329">A11329</accessionNumber>
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, male,
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<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.95" unit="mm" value="19.5">19.5 mm</quantity>
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SUL; PEM
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/A11330">A11330</accessionNumber>
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, female,
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<quantity metricMagnitude="-2" metricUnit="m" metricValue="2.07" unit="mm" value="20.7">20.7 mm</quantity>
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SUL; PEM
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/A11331">A11331</accessionNumber>
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, male,
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<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.8199999999999998" unit="mm" value="18.2">18.2 mm</quantity>
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SUL; PEM
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/A11332">A11332</accessionNumber>
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, juvenile,
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<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.71" unit="mm" value="17.1">17.1 mm</quantity>
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SUL; PEM
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/A11333">A11333</accessionNumber>
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, male,
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<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.7399999999999998" unit="mm" value="17.4">17.4 mm</quantity>
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SUL; PEM
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/A11334">A11334</accessionNumber>
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, male,
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<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.8199999999999998" unit="mm" value="18.2">18.2 mm</quantity>
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SUL; PEM
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/A11335">A11335</accessionNumber>
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, male,
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<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.75" unit="mm" value="17.5">17.5 mm</quantity>
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SUL; PEM
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/A11336">A11336</accessionNumber>
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, juvenile,
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<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.39" unit="mm" value="13.9">13.9 mm</quantity>
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SUL; PEM
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/A11337">A11337</accessionNumber>
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, juvenile,
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<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.33" unit="mm" value="13.3">13.3 mm</quantity>
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SUL; PEM
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/A11338">A11338</accessionNumber>
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, juvenile,
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<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.5699999999999998" unit="mm" value="15.7">15.7 mm</quantity>
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SUL; PEM
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/A11339">A11339</accessionNumber>
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, female,
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<quantity metricMagnitude="-2" metricUnit="m" metricValue="2.59" unit="mm" value="25.9">25.9 mm</quantity>
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SUL; PEM
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/A11340">A11340</accessionNumber>
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, male,
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<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.75" unit="mm" value="17.5">17.5 mm</quantity>
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SUL; PEM
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/A11342">A11342</accessionNumber>
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, male,
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<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.81" unit="mm" value="18.1">18.1 mm</quantity>
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SUL; PEM
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/A11343">A11343</accessionNumber>
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, female, 22.0 mm SUL; PEM
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/A11344">A11344</accessionNumber>
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, male,
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<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.8399999999999999" unit="mm" value="18.4">18.4 mm</quantity>
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SUL; PEM
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/A11345">A11345</accessionNumber>
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, male, 18.0 mm SUL; PEM
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/A11346">A11346</accessionNumber>
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, male,
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<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.77" unit="mm" value="17.7">
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<pageBreakToken pageId="6" pageNumber="169" start="start">17.7</pageBreakToken>
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mm
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</quantity>
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SUL; PEM
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/A11347">A11347</accessionNumber>
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, male,
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<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.7399999999999998" unit="mm" value="17.4">17.4 mm</quantity>
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SUL), Mt.
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<normalizedToken originalValue="M’pàluwé">M'paluwe</normalizedToken>
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(PEM
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/A11369">A11369</accessionNumber>
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, female,
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<quantity metricMagnitude="-2" metricUnit="m" metricValue="2.17" unit="mm" value="21.7">21.7 mm</quantity>
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SUL; PEM
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/A11370">A11370</accessionNumber>
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, female,
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<quantity metricMagnitude="-2" metricUnit="m" metricValue="2.38" unit="mm" value="23.8">23.8 mm</quantity>
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SUL; PEM
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/A11371">A11371</accessionNumber>
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, male,
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<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.92" unit="mm" value="19.2">19.2 mm</quantity>
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SUL; PEM
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<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/A11372">A11372</accessionNumber>
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, male,
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<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.81" unit="mm" value="18.1">18.1 mm</quantity>
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SUL)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="6" pageNumber="169" type="comments">
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<paragraph pageId="6" pageNumber="169">Comments</paragraph>
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<paragraph pageId="6" pageNumber="169">
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Specimens were collected both during the day and at night from rocky outcrops in water seepages (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A selection of amphibians from northern Mozambique inselbergs. A - Arthroleptis francei (Mt. Namuli, PEM A 11303), B - Arthroleptis xenodactyloides (Mt. Mabu), C - Arthroleptis stenodactylus (Mt. Mabu), D - Amietia delalandii (Mt. Namuli, PEM A 11319), E - Nothophryne cf. broadleyi (Mt. M'paluwe, PEM A 11370), F - Nothophryne cf. broadleyi (Mt. Namuli), G - Strongylopus cf. fuelleborni (Mt. Mabu, PEM A 11184), H - Scolecomorphus cf. kirkii (Mt. Mabu, PEM A 11248)." figureDoi="10.3897/zse.92.9948.figure3" httpUri="https://binary.pensoft.net/fig/104227" pageId="6" pageNumber="169">3E</figureCitation>
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,
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A selection of amphibians from northern Mozambique inselbergs. A - Arthroleptis francei (Mt. Namuli, PEM A 11303), B - Arthroleptis xenodactyloides (Mt. Mabu), C - Arthroleptis stenodactylus (Mt. Mabu), D - Amietia delalandii (Mt. Namuli, PEM A 11319), E - Nothophryne cf. broadleyi (Mt. M'paluwe, PEM A 11370), F - Nothophryne cf. broadleyi (Mt. Namuli), G - Strongylopus cf. fuelleborni (Mt. Mabu, PEM A 11184), H - Scolecomorphus cf. kirkii (Mt. Mabu, PEM A 11248)." figureDoi="10.3897/zse.92.9948.figure3" httpUri="https://binary.pensoft.net/fig/104227" pageId="6" pageNumber="169">3F</figureCitation>
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). Historically,
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<taxonomicName authorityName="Poynton" authorityYear="1963" class="Amphibia" family="Pyxicephalidae" genus="Nothophryne" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothophryne" order="Gymnophiona" pageId="6" pageNumber="169" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="6" pageNumber="169">Nothophryne</emphasis>
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</taxonomicName>
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has only been reported from two localities: Mt. Mulanje, Malawi (type locality) and Mt.
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<normalizedToken originalValue="M’pàluwé">M'paluwe</normalizedToken>
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in Mozambique (Poynton 1962 &
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<bibRefCitation author="Poynton, JC" journalOrPublisher="Ciencas B" pageId="10" pageNumber="173" pagination="13 - 34" refId="B56" refString="Poynton, JC, 1966. Amphibia of northern Mozambique. Memoirs de Instituto de Investigacao Cientifica de Mocambique Ser. A. Ciencas B 8: 13 - 34" title="Amphibia of northern Mozambique. Memoirs de Instituto de Investigacao Cientifica de Mocambique Ser. A." volume="8" year="1966">1966</bibRefCitation>
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,
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<bibRefCitation author="Blake, DK" journalOrPublisher="The Journal of the Herpetological Association of Rhodesia" pageId="10" pageNumber="173" pagination="31 - 46" publicationUrl="10.1080/0440730X.1965.9650549" refId="B5" refString="Blake, DK, 1965. The fourth Umtali Museum expedition to Mozambique: November -December 1964. The Journal of the Herpetological Association of Rhodesia 23: 31 - 46, DOI: 10.1080/0440730X.1965.9650549" title="The fourth Umtali Museum expedition to Mozambique: November - December 1964." url="10.1080/0440730X.1965.9650549" volume="23" year="1965">Blake 1965</bibRefCitation>
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), both higher than 1200 m asl.
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<bibRefCitation author="Timberlake, J" journalOrPublisher="PLoS ONE" pageId="10" pageNumber="173" publicationUrl="http://www.kew.org/science/directory/projects/annex/Namuli_report_FINAL.pdf" refId="B68" refString="Timberlake, J, Dowsett-Lemaire, F, Bayliss, J, Alves, T, Baena, S, Bento, C, Cook, K, Francisco, J, Harris, T, Smith, P, de Sousa, C, 2009. Mt. Namuli. Mozambique: Biodiversity and Conservation. In: Technical Report: Darwin Initiative Award 15/036: Monitoring and Managing Biodiversity Loss in South-East Africa's Montane Ecosystems., http://www.kew.org/science/directory/projects/annex/Namuli_report_FINAL.pdf" title="Mt. Namuli. Mozambique: Biodiversity and Conservation." url="http://www.kew.org/science/directory/projects/annex/Namuli_report_FINAL.pdf" volumeTitle="Technical Report: Darwin Initiative Award 15 / 036: Monitoring and Managing Biodiversity Loss in South-East Africa's Montane Ecosystems." year="2009">Timberlake et al. (2009)</bibRefCitation>
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were the first to report this species from Mt. Namuli. Some calls similar to
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<taxonomicName authorityName="Poynton" authorityYear="1963" class="Amphibia" family="Pyxicephalidae" genus="Nothophryne" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nothophryne" order="Gymnophiona" pageId="6" pageNumber="169" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="6" pageNumber="169">Nothophryne</emphasis>
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</taxonomicName>
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were heard from the summit of Mt. Mabu, but no frog was visually located to confirm the species occurrence in this mountain. Genetic analyses suggest a cryptic diversity in this genus (
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<bibRefCitation author="Bittencourt-Silva, G" journalOrPublisher="Molecular Phylogenies and Evolution" pageId="10" pageNumber="173" pagination="89 - 102" publicationUrl="10.1016/j.ympev.2016.03.021" refId="B4" refString="Bittencourt-Silva, G, Conradie, W, Siu-Ting, K, Tolley, KA, Channing, A, Cunningham, M, Farooq, HM, Menegon, M, Loader, SP, 2016. The phylogenetic position and diversity of the enigmatic mongrel frog Nothophryne Poynton, 1963 (Amphibia, Anura). Molecular Phylogenies and Evolution 99: 89 - 102, DOI: 10.1016/j.ympev.2016.03.021" title="The phylogenetic position and diversity of the enigmatic mongrel frog Nothophryne Poynton, 1963 (Amphibia, Anura)." url="10.1016/j.ympev.2016.03.021" volume="99" year="2016">Bittencourt-Silva et al. 2016</bibRefCitation>
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) and additional work is underway to update its taxonomy. Based on its small distribution these nominotypical species is listed as Endangered (
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<bibRefCitation author="Mazibuko, L" journalOrPublisher="The IUCN Red List of Threatened Species" pageId="10" pageNumber="173" publicationUrl="10.2305/IUCN.UK.2004.RLTS.T58077A11714410.en" refId="B44" refString="Mazibuko, L, Poynton, J, 2004. Nothophryne broadleyi. The IUCN Red List of Threatened Species 2004, DOI: 10.2305/IUCN.UK.2004.RLTS.T58077A11714410.en" title="Nothophryne broadleyi." url="10.2305/IUCN.UK.2004.RLTS.T58077A11714410.en" volume="2004" year="2004">Mazibuko and Poynton 2004</bibRefCitation>
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). Referring to that the undescribed species will receive similar red list status.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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