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<document id="9E71764D47C4484094D42FF0435A8606" ID-CLB-Dataset="21150" ID-DOI="10.11646/zootaxa.4657.3.7" ID-GBIF-Dataset="5458f1f0-63cd-4a12-9741-a83afbca0482" ID-ISSN="1175-5326" ID-Zenodo-Dep="3998493" ID-ZooBank="7B2B9390-ABAA-4220-9A0B-35ABC2D29AA6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1598305756061" checkinUser="felipe" docAuthor="Borges, Rafael Cabral &amp; Silveira, Orlando Tobias" docDate="2019" docId="03B3D101FFC0D85A7A9AFDDEFD1AFB8E" docLanguage="en" docName="zootaxa.4657.3.7.pdf.imf" docOrigin="Zootaxa 4657 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Mischocyttarus omicron Richards 1978" docType="treatment" docVersion="8" lastPageNumber="558" masterDocId="FF8AA979FFCBD8577A0DFFFBF802FFB9" masterDocTitle="Revision of the species-group of Mischocyttarus (Omega) filiformis (de Saussure 1854), with description of three new species (Hymenoptera, Vespidae)" masterLastPageNumber="564" masterPageNumber="545" pageNumber="556" updateTime="1698845339287" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="F1D87221F6055E9584E0457EC099C7A5">Revision of the species-group of Mischocyttarus (Omega) filiformis (de Saussure 1854), with description of three new species (Hymenoptera, Vespidae)</mods:title>
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<mods:name id="2A1B015084C624FCAE0EA828DABC2802" type="personal">
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<mods:namePart id="2EC004788BD4EF29F4CFCC1FF99FCDD5">Borges, Rafael Cabral</mods:namePart>
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<mods:namePart id="9FEDCDB9AD4FA46C4067CC3FE789A867">Silveira, Orlando Tobias</mods:namePart>
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<mods:title id="9D6E8720A422743D77DF3B9FEEB994AB">Zootaxa</mods:title>
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<mods:date id="78446B5F74EB851CC58C65BAA6666A20">2019</mods:date>
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<treatment id="03B3D101FFC0D85A7A9AFDDEFD1AFB8E" ID-DOI="http://doi.org/10.5281/zenodo.4330039" ID-GBIF-Taxon="167293724" ID-Zenodo-Dep="4330039" LSID="urn:lsid:plazi:treatment:03B3D101FFC0D85A7A9AFDDEFD1AFB8E" httpUri="http://treatment.plazi.org/id/03B3D101FFC0D85A7A9AFDDEFD1AFB8E" lastPageId="13" lastPageNumber="558" pageId="11" pageNumber="556">
<subSubSection id="C300339CFFC0D85C7A9AFDDEFA72FD86" box="[151,624,549,575]" pageId="11" pageNumber="556" type="nomenclature">
<paragraph id="8BA56017FFC0D85C7A9AFDDEFA72FD86" blockId="11.[151,624,549,611]" box="[151,624,549,575]" pageId="11" pageNumber="556">
<heading id="D0EDD77BFFC0D85C7A9AFDDEFA72FD86" box="[151,624,549,575]" pageId="11" pageNumber="556">
<taxonomicName id="4C1A1B94FFC0D85C7A9AFDDEFA72FD86" ID-CoL="43MZZ" authority="Richards 1978" authorityName="Richards" authorityYear="1978" box="[151,624,549,575]" class="Insecta" family="Vespidae" genus="Mischocyttarus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="556" phylum="Arthropoda" rank="species" species="omicron">
<emphasis id="B96EBC05FFC0D85C7A9AFDDEF9B6FD86" box="[151,436,549,575]" italics="true" pageId="11" pageNumber="556">Mischocyttarus omicron</emphasis>
<bibRefCitation id="EF8B1DE6FFC0D85C7BB6FDDEFA72FD86" author="Richards, O. W." box="[443,624,549,575]" pageId="11" pageNumber="556" refId="ref15263" refString="Richards, O. W. (1978) The social wasps of the Americas excluding the Vespinae. British Museum (Natural History), London, 580 pp." type="book" year="1978">
<emphasis id="B96EBC05FFC0D85C7BB6FDDEFA72FD86" bold="true" box="[443,624,549,575]" pageId="11" pageNumber="556">Richards 1978</emphasis>
</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C300339CFFC0D85C7A9AFDB2F96FFDDA" box="[151,365,584,611]" pageId="11" pageNumber="556" type="description">
<paragraph id="8BA56017FFC0D85C7A9AFDB2F96FFDDA" blockId="11.[151,624,549,611]" box="[151,365,584,611]" pageId="11" pageNumber="556">
(
<figureCitation id="13217C92FFC0D85C7A92FDB2F8E5FDDA" box="[159,231,585,611]" captionStart="FIGURES 6–13" captionStartId="4.[151,264,1872,1897]" captionTargetBox="[245,1341,237,1846]" captionTargetId="figure@4.[245,1341,237,1847]" captionTargetPageId="4" captionText="FIGURES 6–13. General dorsal or lateral body views (females; all from Brazil, except 10 and 11). 6 a-b: holotype of M. filiformis (NHM); 7: same species (Pará, Belém, Utinga; MPEG); 8: holotype of M. omicron (NHM); 9: M. buyssoni (R. de Janeiro; MPEG); 10–11: holotype of M. napoensis (Ecuador; NHM); 12: holotype of M. kallindusfloren sp. nov. (Pará, Carajás; MPEG); 13: holotype of M. asahi sp. nov. (Pará, Carajás; MPEG). all scales = 1.00 mm." figureDoi="http://doi.org/10.5281/zenodo.3998497" httpUri="https://zenodo.org/record/3998497/files/figure.png" pageId="11" pageNumber="556">Figs 8</figureCitation>
,
<figureCitation id="13217C92FFC0D85C7AF9FDB3F93EFDDA" box="[244,316,584,611]" captionStart="FIGURES 14–20" captionStartId="5.[151,264,1875,1900]" captionTargetBox="[244,1343,186,1845]" captionTargetId="figure@5.[226,1362,181,1850]" captionTargetPageId="5" captionText="FIGURES 14–20. Head in frontal view (except 17, posterior view) (females; all from Brazil, except 14 and 16) 14: M. filiformis (Bolivia, Beni; NHM); 15 and 17: M. omicron (Pará, Carajás; MPEG); 16: holotype of M. napoensis (Ecuador; NHM); 18: holotype of M. kallindusfloren sp. nov. (Pará, Carajás; MPEG); 19: holotype of M. asahi sp. nov. (Pará, Carajás; MPEG); 20: M. buyssoni (R. de Janeiro; MPEG); all scales = 0.50 mm. Figure 21: colony and nest of M. filiformis (Pará, Belém; campus of MPEG)—nest size 1,5 × 1,2 cm." figureDoi="http://doi.org/10.5281/zenodo.3998499" httpUri="https://zenodo.org/record/3998499/files/figure.png" pageId="11" pageNumber="556">15, 17</figureCitation>
,
<figureCitation id="13217C92FFC0D85C7B45FDB2F967FDDA" box="[328,357,585,611]" captionStart="FIGURES 22–27" captionStartId="8.[151,264,1671,1696]" captionTargetBox="[156,1427,187,1631]" captionTargetId="figure@8.[151,1436,181,1645]" captionTargetPageId="8" captionText="FIGURES 22–27. Anterior dorsal view of mesosoma (females, except 27; all from Brazil). 22: M. filiformis (Pará, Belém, Utinga; MPEG); 23: M. buyssoni (R. de Janeiro; MPEG); 24: M. omicron (Pará, Carajás; MPEG); 25: holotype of M. asahi sp. nov. (Pará, Carajás; MPEG); 26: holotype of M. kallindusfloren sp. nov. (Pará, Carajás; MPEG); 27: holotype of M. nazgul sp. nov. (Pará, Carajás; MPEG); all scales = 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.3998501" httpUri="https://zenodo.org/record/3998501/files/figure.png" pageId="11" pageNumber="556">24</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C300339CFFC0D85C7A9AFD76FBE3FD5A" pageId="11" pageNumber="556" type="reference_group">
<paragraph id="8BA56017FFC0D85C7A9AFD76F9B1FD7D" blockId="11.[151,1436,652,739]" pageId="11" pageNumber="556">
<treatmentCitationGroup id="AB0A4739FFC0D85C7A9AFD76F9B1FD7D" pageId="11" pageNumber="556">
<treatmentCitation id="0ABB4606FFC0D85C7A9AFD76FB47FD1C" author="Richards, O. W." box="[151,837,652,677]" page="372" pageId="11" pageNumber="556" year="1978">
<taxonomicName id="4C1A1B94FFC0D85C7A9AFD76FB47FD1C" ID-CoL="43MZZ" authority="Richards, 1978: 372" authorityName="Richards" authorityPageNumber="372" authorityYear="1978" box="[151,837,652,677]" class="Insecta" family="Vespidae" genus="Mischocyttarus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="556" phylum="Arthropoda" rank="species" species="omicron" subGenus="Monacanthocnemis">
<emphasis id="B96EBC05FFC0D85C7A9AFD76FA6EFD1C" box="[151,620,653,677]" italics="true" pageId="11" pageNumber="556">Mischocyttarus (Monacanthocnemis) omicron</emphasis>
<bibRefCitation id="EF8B1DE6FFC0D85C7879FD77FB47FD1C" author="Richards, O. W." box="[628,837,652,677]" pageId="11" pageNumber="556" refId="ref15263" refString="Richards, O. W. (1978) The social wasps of the Americas excluding the Vespinae. British Museum (Natural History), London, 580 pp." type="book" year="1978">Richards, 1978: 372</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="54A1DEB5FFC0D85C795DFD77FBACFD1C" box="[848,942,652,677]" pageId="11" pageNumber="556" type="holotype">Holotype</typeStatus>
<specimenCount id="9D1CAB9EFFC0D85C79B8FD75FBCAFD1C" box="[949,968,654,677]" pageId="11" pageNumber="556" type="female">♀</specimenCount>
,
<collectingCountry id="F30D2087FFC0D85C79DBFD77FC14FD1C" box="[982,1046,652,677]" name="Brazil" pageId="11" pageNumber="556">Brazil</collectingCountry>
,
<collectingRegion id="49DEAEF5FFC0D85C7E2CFD75FCAAFD1C" box="[1057,1192,653,677]" country="Brazil" name="Mato Grosso" pageId="11" pageNumber="556">Mato Grosso</collectingRegion>
,
<date id="FFA446D7FFC0D85C7EBEFD77FD23FD1C" box="[1203,1313,652,677]" pageId="11" pageNumber="556" value="1968-03-23">23.iii.1968</date>
O.W. Rich- ards (NHM); examined
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BA56017FFC0D85C7A9AFD31FBE3FD5A" blockId="11.[151,1436,652,739]" box="[151,993,714,739]" pageId="11" pageNumber="556">
<treatmentCitationGroup id="AB0A4739FFC0D85C7A9AFD31FBE3FD5A" box="[151,993,714,739]" pageId="11" pageNumber="556">
<taxonomicName id="4C1A1B94FFC0D85C7A9AFD31FACDFD5A" ID-CoL="43MZZ" authority="Richards" authorityName="Richards" authorityYear="1978" box="[151,719,714,739]" class="Insecta" family="Vespidae" genus="Mischocyttarus" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="556" phylum="Arthropoda" rank="species" species="omicron" subGenus="Monacanthocnemis">
<emphasis id="B96EBC05FFC0D85C7A9AFD31FA68FD5B" box="[151,618,714,738]" italics="true" pageId="11" pageNumber="556">Mischocyttarus (Monacanthocnemis) omicron</emphasis>
Richards
</taxonomicName>
:
<treatmentCitation id="0ABB4606FFC0D85C78D4FD31FBE3FD5A" author="Colomo, M. V. &amp; Berta, D. C." box="[729,993,714,739]" page="81" pageId="11" pageNumber="556" year="2005">
<bibRefCitation id="EF8B1DE6FFC0D85C78D4FD31FBE3FD5A" author="Colomo, M. V. &amp; Berta, D. C." box="[729,993,714,739]" pageId="11" pageNumber="556" pagination="71 - 84" refId="ref14553" refString="Colomo, M. V. &amp; Berta, D. C. (2005) Los ejemplares tipo de Masarinae y Polistinae (Hymenoptera: Vespidae) depositados en la coleccion del Instituto Fundacion Miguel Lillo (IFML), Argentina. Rev. Soc. Entomol. Argent, 64 (1 - 2), 71 - 84." type="journal article" year="2005">Colomo &amp; Berta 2005: 81</bibRefCitation>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C300339CFFC0D85C7A9AFCEBF9EEFB9E" pageId="11" pageNumber="556" type="diagnosis">
<paragraph id="8BA56017FFC0D85C7A9AFCEBF9EEFB9E" blockId="11.[151,1437,784,1963]" pageId="11" pageNumber="556">
<emphasis id="B96EBC05FFC0D85C7A9AFCEBF974FC92" bold="true" box="[151,374,784,811]" pageId="11" pageNumber="556">Diagnosis. Female.</emphasis>
Fore wing length
<quantity id="4CE2CDF2FFC0D85C784EFCEAFABCFC92" box="[579,702,785,811]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="11" pageNumber="556" unit="mm" value="11.5" valueMax="12.0" valueMin="11.0">11–12 mm</quantity>
. Apex of clypeus weakly but noticeably bidentate; humeral lobes large and roughly quadrangular, rather detached from pronotal profile as seen from above (
<figureCitation id="13217C92FFC0D85C7E89FCCEFCD4FCF6" box="[1156,1238,821,847]" captionStart="FIGURES 22–27" captionStartId="8.[151,264,1671,1696]" captionTargetBox="[156,1427,187,1631]" captionTargetId="figure@8.[151,1436,181,1645]" captionTargetPageId="8" captionText="FIGURES 22–27. Anterior dorsal view of mesosoma (females, except 27; all from Brazil). 22: M. filiformis (Pará, Belém, Utinga; MPEG); 23: M. buyssoni (R. de Janeiro; MPEG); 24: M. omicron (Pará, Carajás; MPEG); 25: holotype of M. asahi sp. nov. (Pará, Carajás; MPEG); 26: holotype of M. kallindusfloren sp. nov. (Pará, Carajás; MPEG); 27: holotype of M. nazgul sp. nov. (Pará, Carajás; MPEG); all scales = 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.3998501" httpUri="https://zenodo.org/record/3998501/files/figure.png" pageId="11" pageNumber="556">Fig. 24</figureCitation>
), humeral surface just behind with an elongate shallow concavity; propodeal median furrow linear and straight, rather deep, with an anterior median low keel; propodeal valve high, with lamella narrow throughout; first metasomal segment short for this group (
<figureCitation id="13217C92FFC0D85C7B16FC5AF962FC02" box="[283,352,929,955]" captionStart="FIGURES 6–13" captionStartId="4.[151,264,1872,1897]" captionTargetBox="[245,1341,237,1846]" captionTargetId="figure@4.[245,1341,237,1847]" captionTargetPageId="4" captionText="FIGURES 6–13. General dorsal or lateral body views (females; all from Brazil, except 10 and 11). 6 a-b: holotype of M. filiformis (NHM); 7: same species (Pará, Belém, Utinga; MPEG); 8: holotype of M. omicron (NHM); 9: M. buyssoni (R. de Janeiro; MPEG); 10–11: holotype of M. napoensis (Ecuador; NHM); 12: holotype of M. kallindusfloren sp. nov. (Pará, Carajás; MPEG); 13: holotype of M. asahi sp. nov. (Pará, Carajás; MPEG). all scales = 1.00 mm." figureDoi="http://doi.org/10.5281/zenodo.3998497" httpUri="https://zenodo.org/record/3998497/files/figure.png" pageId="11" pageNumber="556">Fig. 8</figureCitation>
), length less than [mesoscutum + scutellum + metanotum], but narrow and linear basally, with sides diverging only after spiracles; mesoscutum sculpture dense, with homogeneous aspect, puncture diameter varying rather continuously between relatively narrow limits, interstices rather dull; overall color yellow; dark mark on vertex wide, reaching eyes.
</paragraph>
</subSubSection>
<subSubSection id="C300339CFFC0D85A7ACAFBCBFA1FFC82" lastPageId="13" lastPageNumber="558" pageId="11" pageNumber="556" type="description">
<paragraph id="8BA56017FFC0D85C7ACAFBCBFD4AF9FA" blockId="11.[151,1437,784,1963]" pageId="11" pageNumber="556">
<emphasis id="B96EBC05FFC0D85C7ACAFBCBF9D8FBF2" bold="true" box="[199,474,1072,1099]" pageId="11" pageNumber="556">Redescription. Female.</emphasis>
Length of fore wing
<quantity id="4CE2CDF2FFC0D85C78C5FBCAFB40FBF2" box="[712,834,1073,1099]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="11" pageNumber="556" unit="mm" value="11.5" valueMax="12.0" valueMin="11.0">11–12 mm</quantity>
; head distinctly higher than wide in frontal view (
<figureCitation id="13217C92FFC0D85C7F62FBCAF8B4FBD6" captionStart="FIGURES 14–20" captionStartId="5.[151,264,1875,1900]" captionTargetBox="[244,1343,186,1845]" captionTargetId="figure@5.[226,1362,181,1850]" captionTargetPageId="5" captionText="FIGURES 14–20. Head in frontal view (except 17, posterior view) (females; all from Brazil, except 14 and 16) 14: M. filiformis (Bolivia, Beni; NHM); 15 and 17: M. omicron (Pará, Carajás; MPEG); 16: holotype of M. napoensis (Ecuador; NHM); 18: holotype of M. kallindusfloren sp. nov. (Pará, Carajás; MPEG); 19: holotype of M. asahi sp. nov. (Pará, Carajás; MPEG); 20: M. buyssoni (R. de Janeiro; MPEG); all scales = 0.50 mm. Figure 21: colony and nest of M. filiformis (Pará, Belém; campus of MPEG)—nest size 1,5 × 1,2 cm." figureDoi="http://doi.org/10.5281/zenodo.3998499" httpUri="https://zenodo.org/record/3998499/files/figure.png" pageId="11" pageNumber="556">Fig. 15</figureCitation>
); FHH/IntOW 1.16; clypeus clearly higher than wide H/WCLP 1.19; ventral margin distinctly angular, apex rather wide and slightly bidentate; malar space very narrow; tentorial pit much closer to eye margin than to antennal socket; antennal scape moderately elongated, L/Wesc 2.71; posterior occeli less widely separated than in most species of the group, POL/OOL 1.25; occipital carina well developed, distinct in lateral view and ending fairly below level of ocular sinus; gena considerably narrower than upper lobe of the eye; pronotum without lateral fovea; anterior margin with lamella raised and reflexed especially on center, with lateral profile as a loop reaching more than 180 degrees; region immediately behind the lamella without secondary margin; humeral angle well developed WCAR/WMS 1.05; pronotal carina high, not distinctly lowered at center, but forming large rather quadrate humeral lobes (
<figureCitation id="13217C92FFC0D85C7AEFFA8EF931FA36" box="[226,307,1397,1423]" captionStart="FIGURES 22–27" captionStartId="8.[151,264,1671,1696]" captionTargetBox="[156,1427,187,1631]" captionTargetId="figure@8.[151,1436,181,1645]" captionTargetPageId="8" captionText="FIGURES 22–27. Anterior dorsal view of mesosoma (females, except 27; all from Brazil). 22: M. filiformis (Pará, Belém, Utinga; MPEG); 23: M. buyssoni (R. de Janeiro; MPEG); 24: M. omicron (Pará, Carajás; MPEG); 25: holotype of M. asahi sp. nov. (Pará, Carajás; MPEG); 26: holotype of M. kallindusfloren sp. nov. (Pará, Carajás; MPEG); 27: holotype of M. nazgul sp. nov. (Pará, Carajás; MPEG); all scales = 0.50 mm." figureDoi="http://doi.org/10.5281/zenodo.3998501" httpUri="https://zenodo.org/record/3998501/files/figure.png" pageId="11" pageNumber="556">Fig. 24</figureCitation>
) with slightly reflexed extremities, area just behind lobes with an elongate shallow concavity; mesoscutum as long as wide, L/WMS 1.0; fore wing very elongated, LDIS/HMP 2.48; hind tarsi with asymmetric claws, inner claw longer with apex uniformly rounded, but not enlarged or spoon-shaped; propodeal median furrow linear, narrow and deep, with a short keel on its anterior 2/3; propodeal valve very high and poorly expanded backward, lamella narrow with rounded margin; first metasomal segment relatively short (
<figureCitation id="13217C92FFC0D85C7E05F9FEFC48F9A6" box="[1032,1098,1541,1567]" captionStart="FIGURES 6–13" captionStartId="4.[151,264,1872,1897]" captionTargetBox="[245,1341,237,1846]" captionTargetId="figure@4.[245,1341,237,1847]" captionTargetPageId="4" captionText="FIGURES 6–13. General dorsal or lateral body views (females; all from Brazil, except 10 and 11). 6 a-b: holotype of M. filiformis (NHM); 7: same species (Pará, Belém, Utinga; MPEG); 8: holotype of M. omicron (NHM); 9: M. buyssoni (R. de Janeiro; MPEG); 10–11: holotype of M. napoensis (Ecuador; NHM); 12: holotype of M. kallindusfloren sp. nov. (Pará, Carajás; MPEG); 13: holotype of M. asahi sp. nov. (Pará, Carajás; MPEG). all scales = 1.00 mm." figureDoi="http://doi.org/10.5281/zenodo.3998497" httpUri="https://zenodo.org/record/3998497/files/figure.png" pageId="11" pageNumber="556">Fig. 8</figureCitation>
), LSI/HMP 1.15; L/WSI 3.54, apex only 1.66 × wider than basal petiole, sides diverging after spiracles, these only moderately prominent.
</paragraph>
<paragraph id="8BA56017FFC0D85C7ACAF9B6F930F812" blockId="11.[151,1437,784,1963]" pageId="11" pageNumber="556">
<emphasis id="B96EBC05FFC0D85C7ACAF9B6F933F9DE" box="[199,305,1613,1639]" italics="true" pageId="11" pageNumber="556">Sculpture</emphasis>
. Moderately strong and conspicuous. Disk of clypeus with weak medium-sized punctures and nearly indistinguishable micropunctures, area close to ventral margin and mandibles smooth, with medium-sized shallow punctures; upper interantennal area, frons and vertex with close dense punctures of generally small diameter, with still smaller punctures surrounding ocelli; mesoscutum sculpture dense, with homogeneous aspect, puncture diameter varying continuously between rather narrow limits (
<quantity id="4CE2CDF2FFC0D85C790AF926FBABF94E" box="[775,937,1757,1783]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="11" pageNumber="556" unit="mm" value="0.06" valueMax="0.08" valueMin="0.04">0.04–0.08mm</quantity>
), interstices rather dull, area close to margins with smaller punctures; humeral region similar to mesoscutum, diameter slightly larger; mesopleuron with rather shallow punctures of two distinct size-classes, neither large, nor dense, interstices reticulate and shining; metapleuron similar to mesopleuron, but with sparser punctures; propodeum with medium-sized not very dense punctures near anterior margin, interstices perceptible and shining, central area with large and sparser punctures, integument more shining.
</paragraph>
<caption id="DF65309FFFC7D85B7A9AF986F90FF89C" ID-DOI="http://doi.org/10.5281/zenodo.3998503" ID-Zenodo-Dep="3998503" httpUri="https://zenodo.org/record/3998503/files/figure.png" pageId="12" pageNumber="557" startId="12.[151,264,1661,1686]" targetBox="[156,1432,187,1626]" targetPageId="12">
<paragraph id="8BA56017FFC7D85B7A9AF986F9B8F964" blockId="12.[151,1437,1661,1830]" pageId="12" pageNumber="557">
<emphasis id="B96EBC05FFC7D85B7A9AF986F9C8F92F" bold="true" box="[151,458,1661,1686]" pageId="12" pageNumber="557">FIGURES 28–32 (MALES).</emphasis>
28: head of
<taxonomicName id="4C1A1B94FFC7D85B785CF985FACCF92F" authorityName="Ducke. He" authorityYear="1918" box="[593,718,1662,1686]" class="Insecta" family="Vespidae" genus="Mischocyttarus" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="557" phylum="Arthropoda" rank="species" species="buyssoni">
<emphasis id="B96EBC05FFC7D85B785CF985FACCF92F" box="[593,718,1662,1686]" italics="true" pageId="12" pageNumber="557">M. buyssoni</emphasis>
</taxonomicName>
(R. de Janeiro, Itatiaia; IOC); 29: same of
<taxonomicName id="4C1A1B94FFC7D85B7E98F985FCFFF92F" authorityName="Borges &amp; Silveira" authorityYear="2019" box="[1173,1277,1662,1686]" class="Insecta" family="Vespidae" genus="Mischocyttarus" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="557" phylum="Arthropoda" rank="species" species="nazgul">
<emphasis id="B96EBC05FFC7D85B7E98F985FCFFF92F" box="[1173,1277,1662,1686]" italics="true" pageId="12" pageNumber="557">M. nazgul</emphasis>
</taxonomicName>
(Pará, Carajás; MPEG); 30: same of
<taxonomicName id="4C1A1B94FFC7D85B7B63F959F9EEF903" authorityName=": Richards" authorityYear="1978" box="[366,492,1698,1722]" class="Insecta" family="Vespidae" genus="Mischocyttarus" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="557" phylum="Arthropoda" rank="species" species="filiformis">
<emphasis id="B96EBC05FFC7D85B7B63F959F9EEF903" box="[366,492,1698,1722]" italics="true" pageId="12" pageNumber="557">M. filiformis</emphasis>
</taxonomicName>
(Pará, Bujaru; MPEG); scales = 0.50 mm. 31: lateral body view of
<taxonomicName id="4C1A1B94FFC7D85B7E99F959FCFBF903" authorityName="Borges &amp; Silveira" authorityYear="2019" box="[1172,1273,1698,1722]" class="Insecta" family="Vespidae" genus="Mischocyttarus" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="557" phylum="Arthropoda" rank="species" species="nazgul">
<emphasis id="B96EBC05FFC7D85B7E99F959FCFBF903" box="[1172,1273,1698,1722]" italics="true" pageId="12" pageNumber="557">M. nazgul</emphasis>
</taxonomicName>
; 32: same of
<taxonomicName id="4C1A1B94FFC7D85B7F8CF959F8F6F967" authorityName=": Richards" authorityYear="1978" class="Insecta" family="Vespidae" genus="Mischocyttarus" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="557" phylum="Arthropoda" rank="species" species="filiformis">
<emphasis id="B96EBC05FFC7D85B7F8CF959F8F6F967" italics="true" pageId="12" pageNumber="557">M. filiformis</emphasis>
</taxonomicName>
; scales = 1.00 mm.
</paragraph>
<paragraph id="8BA56017FFC7D85B7A9AF912F90FF89C" blockId="12.[151,1437,1661,1830]" pageId="12" pageNumber="557">
Figure 33 (female hind tarsal claws). a:
<taxonomicName id="4C1A1B94FFC7D85B783CF911FA89F8BB" authorityName="Borges &amp; Silveira" authorityYear="2019" box="[561,651,1770,1794]" class="Insecta" family="Vespidae" genus="Mischocyttarus" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="557" phylum="Arthropoda" rank="species" species="asahi">
<emphasis id="B96EBC05FFC7D85B783CF911FA89F8BB" box="[561,651,1770,1794]" italics="true" pageId="12" pageNumber="557">M. asahi</emphasis>
</taxonomicName>
(Pará, Carajás; MPEG); b:
<taxonomicName id="4C1A1B94FFC7D85B79AAF911FC25F8BB" authorityName=": Richards" authorityYear="1978" box="[935,1063,1770,1794]" class="Insecta" family="Vespidae" genus="Mischocyttarus" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="557" phylum="Arthropoda" rank="species" species="filiformis">
<emphasis id="B96EBC05FFC7D85B79AAF911FC25F8BB" box="[935,1063,1770,1794]" italics="true" pageId="12" pageNumber="557">M. filiformis</emphasis>
</taxonomicName>
(Pará, Belém, Utinga; MPEG) scale = 0.20 mm.
</paragraph>
</caption>
<paragraph id="8BA56017FFC7D85A7ACAF8AEF99DFF6E" blockId="12.[151,1436,1876,2010]" lastBlockId="13.[151,1437,153,1079]" lastPageId="13" lastPageNumber="558" pageId="12" pageNumber="557">
<emphasis id="B96EBC05FFC7D85B7ACAF8AEF92AF8D7" box="[199,296,1877,1902]" italics="true" pageId="12" pageNumber="557">Vestiture</emphasis>
. Eyes with very short inconspicuous hairs; head, thorax and mesosoma with conspicuous appressed or decumbent hairs; longer and stouter whitish hairs on lower portion of clypeus, frons, vertex, pronotum and mesopleuron; those on mesoscutum a little shorter; hairs on metapleuron, and propodeum more erect, finer and longer; metasomal sternum 1 distinctly polished with long fine erect hairs; most of tergum 1 and remaining metasomal segments covered by decumbent short hairs; legs generally with conspicuous appressed hairs; femora and tarsi with sparse stout short hairs.
</paragraph>
<paragraph id="8BA56017FFC6D85A7ACAFF1AFA6FFD86" blockId="13.[151,1437,153,1079]" pageId="13" pageNumber="558">
<emphasis id="B96EBC05FFC6D85A7ACAFF1AF90AFF42" box="[199,264,225,251]" italics="true" pageId="13" pageNumber="558">Color</emphasis>
. Yellow. Antennal scape and pedicel, disk or metasomal terga 2–6, light orange testaceous; antennal flagellum; area surrounding foraminal concavity; part of anterior face of pronotum; posterior lateral areas of mesoscutum; mark on tegula; subalar area; margins and sulci of mesopleuron; ventral angle of lower metapleural plate; area along lateral margin of propodeum; elongated marks or areas on posterior face of hind coxa, mid and hind trochanters, femora, tibiae, and tarsi; metasomal tergum 1, testaceous to light brown; large triangular area on vertex connecting at sides to eyes and by a posterior median short arm to a wide occipital mark which also connects to eyes (letting two triangular yellow spots on vertex); anterior face of pronotum, large humeral mark, central stripe and anterior region of mesoscutum, most of scutellum and metanotum posteriorly, wide anterior area (extending backwards at sides) and median stripe on propodeum (letting two elongated submedian yellow spots), dark brown to black; wings hyaline with brown veins.
</paragraph>
<paragraph id="8BA56017FFC6D85A7ACAFDB2F98AFDDA" blockId="13.[151,1437,153,1079]" box="[199,392,585,611]" pageId="13" pageNumber="558">
<emphasis id="B96EBC05FFC6D85A7ACAFDB2F909FDDA" bold="true" box="[199,267,585,611]" pageId="13" pageNumber="558">Male.</emphasis>
Unknown.
</paragraph>
<paragraph id="8BA56017FFC6D85A7ACAFD96FB84FD4D" blockId="13.[151,1437,153,1079]" pageId="13" pageNumber="558">
<emphasis id="B96EBC05FFC6D85A7ACAFD96F901FD3E" bold="true" box="[199,259,621,647]" pageId="13" pageNumber="558">Nest.</emphasis>
<bibRefCitation id="EF8B1DE6FFC6D85A7B01FD96F9C1FD3E" author="Richards, O. W." box="[268,451,621,647]" pageId="13" pageNumber="558" refId="ref15263" refString="Richards, O. W. (1978) The social wasps of the Americas excluding the Vespinae. British Museum (Natural History), London, 580 pp." type="book" year="1978">Richards (1978)</bibRefCitation>
described the nest of this species as having a whitish carton, with a flattened 5.0 mm pedicel attached at a corner made by three adjacent cells. It seems to resemble the nest of
<taxonomicName id="4C1A1B94FFC6D85A7E86FD69FD1AFD12" authorityName=": Richards" authorityYear="1978" box="[1163,1304,657,683]" class="Insecta" family="Vespidae" genus="Mischocyttarus" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="558" phylum="Arthropoda" rank="species" species="filiformis">
<emphasis id="B96EBC05FFC6D85A7E86FD69FD1AFD12" box="[1163,1304,657,683]" italics="true" pageId="13" pageNumber="558">M. filiformis</emphasis>
</taxonomicName>
. Both nests mentioned by
<bibRefCitation id="EF8B1DE6FFC6D85A7B38FD4EF9EBFD76" author="Richards, O. W." box="[309,489,693,719]" pageId="13" pageNumber="558" refId="ref15263" refString="Richards, O. W. (1978) The social wasps of the Americas excluding the Vespinae. British Museum (Natural History), London, 580 pp." type="book" year="1978">Richards (1978)</bibRefCitation>
were attached to adaxial surfaces of leaves of shrubs about
<quantity id="4CE2CDF2FFC6D85A7E71FD4EFCCDFD76" box="[1148,1231,693,719]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="558" unit="m" value="1.0">1 meter</quantity>
above the ground. The nests were in areas of open vegetation (field and road margin).
</paragraph>
<paragraph id="8BA56017FFC6D85A7ACAFD07FA1FFC82" blockId="13.[151,1437,153,1079]" pageId="13" pageNumber="558">
<emphasis id="B96EBC05FFC6D85A7ACAFD07F9EDFCAE" bold="true" box="[199,495,764,791]" pageId="13" pageNumber="558">Geographic distribution.</emphasis>
<collectingCountry id="F30D2087FFC6D85A7BF8FD06FA5DFCAE" box="[501,607,765,791]" name="Brazil" pageId="13" pageNumber="558">BRAZIL</collectingCountry>
:
<collectingRegion id="49DEAEF5FFC6D85A7864FD06FAA3FCAE" box="[617,673,765,791]" country="Brazil" name="Para" pageId="13" pageNumber="558">Pará</collectingRegion>
*: Serra dos Carajás;
<collectingRegion id="49DEAEF5FFC6D85A7987FD06FC22FCAE" box="[906,1056,765,791]" country="Brazil" name="Mato Grosso" pageId="13" pageNumber="558">Mato Grosso</collectingRegion>
: Ribeirão Cascalheira.
<collectingCountry id="F30D2087FFC6D85A7F2CFD06FD9BFCAE" box="[1313,1433,765,791]" name="Bolivia" pageId="13" pageNumber="558">BOLIVIA</collectingCountry>
:
<collectingRegion id="49DEAEF5FFC6D85A7A9AFCDAF8D2FC82" box="[151,208,801,827]" country="Bolivia" name="Beni" pageId="13" pageNumber="558">Beni</collectingRegion>
: Guayaramerim. (see
<figureCitation id="13217C92FFC6D85A7BC8FCDAFA17FC82" box="[453,533,801,827]" captionStart="FIGURE 34" captionStartId="14.[151,250,1206,1231]" captionTargetBox="[159,1432,349,1175]" captionTargetId="figure@14.[151,1436,345,1181]" captionTargetPageId="14" captionText="FIGURE 34. Partial truncated map of South America with distributions of species of the M. filiformis group. Embedded frame box shows enlarged area representation with the records of M. filiformis, M. omicron, M. asahi sp. nov., M. kallindusfloren sp. nov. and M. nazgul sp. nov. within the Carajás National Forest (FLONA) in southeastern Pará state." figureDoi="http://doi.org/10.5281/zenodo.3998505" httpUri="https://zenodo.org/record/3998505/files/figure.png" pageId="13" pageNumber="558">Fig. 34</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C300339CFFC6D85A7ACAFCBEFD1AFB8E" pageId="13" pageNumber="558" type="discussion">
<paragraph id="8BA56017FFC6D85A7ACAFCBEF8D6FC56" blockId="13.[151,1437,153,1079]" pageId="13" pageNumber="558">
<emphasis id="B96EBC05FFC6D85A7ACAFCBEF939FCE6" bold="true" box="[199,315,837,863]" pageId="13" pageNumber="558">Remarks.</emphasis>
The known geographic distribution of this species is considerably broadened in this work, with the addition of a record from southeastern
<collectingRegion id="49DEAEF5FFC6D85A7858FC92FAC7FC3A" box="[597,709,873,899]" country="Brazil" name="Para" pageId="13" pageNumber="558">Pará state</collectingRegion>
, in
<collectingCountry id="F30D2087FFC6D85A78FCFC92FB37FC3A" box="[753,821,873,899]" name="Brazil" pageId="13" pageNumber="558">Brazil</collectingCountry>
(
<figureCitation id="13217C92FFC6D85A794BFC92FB9EFC3A" box="[838,924,873,899]" captionStart="FIGURE 34" captionStartId="14.[151,250,1206,1231]" captionTargetBox="[159,1432,349,1175]" captionTargetId="figure@14.[151,1436,345,1181]" captionTargetPageId="14" captionText="FIGURE 34. Partial truncated map of South America with distributions of species of the M. filiformis group. Embedded frame box shows enlarged area representation with the records of M. filiformis, M. omicron, M. asahi sp. nov., M. kallindusfloren sp. nov. and M. nazgul sp. nov. within the Carajás National Forest (FLONA) in southeastern Pará state." figureDoi="http://doi.org/10.5281/zenodo.3998505" httpUri="https://zenodo.org/record/3998505/files/figure.png" pageId="13" pageNumber="558">Fig. 34</figureCitation>
). The collecting points suggest a distribution along the southern border of the Amazon biome, in transitional areas to Cerrado. Specimens from all areas are similar and may be promptly distinguished from other species in this group by their larger size (fore wing length
<quantity id="4CE2CDF2FFC6D85A7F8DFC4AF8C5FC56" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="13" pageNumber="558" unit="mm" value="12.0">12 mm</quantity>
).
</paragraph>
<paragraph id="8BA56017FFC6D85A7ACAFC03FD1AFB8E" blockId="13.[151,1437,153,1079]" pageId="13" pageNumber="558">
<emphasis id="B96EBC05FFC6D85A7ACAFC03F9B6FBAA" bold="true" box="[199,436,1016,1043]" pageId="13" pageNumber="558">Examined material.</emphasis>
<collectingCountry id="F30D2087FFC6D85A7BB1FC02FA36FBAA" box="[444,564,1017,1043]" name="Bolivia" pageId="13" pageNumber="558">BOLIVIA</collectingCountry>
:
<collectingRegion id="49DEAEF5FFC6D85A7832FC02FA7AFBAA" box="[575,632,1017,1043]" country="Bolivia" name="Beni" pageId="13" pageNumber="558">Beni</collectingRegion>
: Guayaramerim,
<specimenCount id="9D1CAB9EFFC6D85A7934FC02FB4CFBAA" box="[825,846,1017,1043]" pageId="13" pageNumber="558" type="female">♀</specimenCount>
(
<typeStatus id="54A1DEB5FFC6D85A7950FC02FBEEFBAA" box="[861,1004,1017,1043]" pageId="13" pageNumber="558" type="paratype">PARATYPE</typeStatus>
)
<date id="FFA446D7FFC6D85A79F6FC03FC6BFBAA" box="[1019,1129,1016,1043]" pageId="13" pageNumber="558" value="1954-05-10">10.v.1954</date>
(W. Foster) (IFML). Other material.
<collectingCountry id="F30D2087FFC6D85A7B0CFBE6F969FB8E" box="[257,363,1053,1079]" name="Brazil" pageId="13" pageNumber="558">BRAZIL</collectingCountry>
:
<collectingRegion id="49DEAEF5FFC6D85A7B78FBE6F9A9FB8E" box="[373,427,1053,1079]" country="Brazil" name="Para" pageId="13" pageNumber="558">Pará</collectingRegion>
: Serra dos Carajás, Serra Norte, Fofoca,
<specimenCount id="9D1CAB9EFFC6D85A797EFBE6FB8AFB8E" box="[883,904,1053,1079]" pageId="13" pageNumber="558" type="female">♀</specimenCount>
,
<date id="FFA446D7FFC6D85A799BFBE7FC0CFB8E" box="[918,1038,1052,1079]" pageId="13" pageNumber="558" value="1985-09-16">16.ix.1985</date>
(M.F. Torres) (MPEG).
</paragraph>
</subSubSection>
</treatment>
</document>
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