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<document id="B059E21F70D9D0C8042A09CE7173E34A" ID-CLB-Dataset="29875" ID-DOI="10.11646/zootaxa.4434.3.3" ID-GBIF-Dataset="3ebb3cb7-3b8f-4c4d-8858-d37185292c28" ID-ISSN="1175-5326" ID-Zenodo-Dep="1292189" ID-ZooBank="8B95D692-E8F7-402B-8485-3D3AE20111E8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1529330577762" checkinUser="plazi" docAuthor="Seeman, Owen D., Minor, Maria, Baker, Michelle R. &amp; Walter, David Evans" docDate="2018" docId="0398D71B525E734F5FFAFF623AD0EB6F" docLanguage="en" docName="zootaxa.4434.3.3.pdf.imf" docOrigin="Zootaxa 4434 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Heatherellidae Walter 1997" docType="key" docVersion="7" lastPageNumber="464" masterDocId="FFA1AF63524973585F6DFFFA3F4BE851" masterDocTitle="A revision of the Heatherellidae (Parasitiformes: Mesostigmata) with a new genus and two new species from Australasia" masterLastPageNumber="465" masterPageNumber="441" pageNumber="464" updateTime="1698549773811" updateUser="ExternalLinkService">
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<mods:title id="90254886B14892DE52C2149A11A398BE">A revision of the Heatherellidae (Parasitiformes: Mesostigmata) with a new genus and two new species from Australasia</mods:title>
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<mods:namePart id="7473FF7D56982ECECC65AD777B30E7DC">Seeman, Owen D.</mods:namePart>
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<mods:namePart id="267B67F7D99AF92FD446A5AAEF073598">Minor, Maria</mods:namePart>
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<mods:namePart id="DE507EAB2813AF90F8818AC6E7E681B4">Baker, Michelle R.</mods:namePart>
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<mods:namePart id="487BDFC61CB7B398AF68A5C3BF65333F">Walter, David Evans</mods:namePart>
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<mods:date id="CAAD55C9B77013D4D87C2693C5CCACC6">2018</mods:date>
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<mods:number id="674F94940D4203D9A64C585F8E21A265">2018-06-18</mods:number>
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<mods:number id="C8D54D278112110B43A917B7F901E1C0">4434</mods:number>
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<mods:identifier id="D47EBA307494B3F7A9A09D491930B529" type="DOI">10.11646/zootaxa.4434.3.3</mods:identifier>
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<treatment id="0398D71B525E734F5FFAFF623AD0EB6F" ID-DOI="http://doi.org/10.5281/zenodo.5966643" ID-GBIF-Taxon="145968983" ID-Zenodo-Dep="5966643" LSID="urn:lsid:plazi:treatment:0398D71B525E734F5FFAFF623AD0EB6F" httpUri="http://treatment.plazi.org/id/0398D71B525E734F5FFAFF623AD0EB6F" lastPageNumber="464" pageId="23" pageNumber="464">
<subSubSection id="C32B3586525E734F5FFAFF623C54E8E3" box="[151,799,152,178]" pageId="23" pageNumber="464" type="nomenclature">
<paragraph id="8B8E660D525E734F5FFAFF623C54E8E3" blockId="23.[151,799,152,178]" box="[151,799,152,178]" pageId="23" pageNumber="464">
<heading id="D0C6D161525E734F5FFAFF623C54E8E3" bold="true" box="[151,799,152,178]" fontSize="11" level="1" pageId="23" pageNumber="464" reason="1">
<emphasis id="B945BA1F525E734F5FFAFF623C54E8E3" bold="true" box="[151,799,152,178]" pageId="23" pageNumber="464">
Key to genera and species of
<taxonomicName id="4C311D8E525E734F5D6DFF623DFCE8E3" ID-CoL="624QN" authorityName="Walter" authorityYear="1997" box="[512,695,152,178]" class="Arachnida" family="Heatherellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="23" pageNumber="464" phylum="Arthropoda" rank="family">Heatherellidae</taxonomicName>
(adults)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32B3586525E734F5FFAFF273AD0EB6F" pageId="23" pageNumber="464" type="key">
<keyStep id="30C57EA8525E734F5FFAFF273AD0E9F2" pageId="23" pageNumber="464">
<paragraph id="8B8E660D525E734F5FFAFF273AD7E91A" blockId="23.[151,1436,221,830]" pageId="23" pageNumber="464">
<keyLead id="30C0C538525E734F5FFAFF273AD7E91A" pageId="23" pageNumber="464">
1. Peritremes absent (
<figureCitation id="130A7A88525E734F5EF5FF273E9EE8A2" box="[408,469,221,243]" captionStart="FIGURE 1" captionStartId="4.[151,250,1999,2021]" captionTargetBox="[158,1436,230,1948]" captionTargetId="figure@4.[151,1436,227,1948]" captionTargetPageId="4" captionText="FIGURE 1. Heatherella osleri n. sp. female holotype, venter." httpUri="https://zenodo.org/record/1292191/files/figure.png" pageId="23" pageNumber="464">Figs 1</figureCitation>
,
<figureCitation id="130A7A88525E734F5E8CFF273EB7E8A2" box="[481,508,221,243]" captionStart="FIGURES 34–37" captionStartId="22.[151,264,1832,1854]" captionTargetBox="[326,1233,232,1771]" captionTargetId="figure@22.[309,1273,196,1784]" captionTargetPageId="22" captionText="FIGURES 34–37. (34) Heatherella acanthocharis walter, 1997, dorsomarginal protuberances, pustules and pores of paratype female (P-s5 pustule; P-s6 pore; P-S1 protuberance), dorsal view of flattened specimen, right side; (35) Aheatherella mira n. sp., dorsomarginal protuberances, pustules and pores of paratype female (P-s5 pustule; P-s6 protuberance with closely associated seta), dorsal view of ventrolateral fragment with broken r-setae, left side; (36) Heatherella osleri n. sp., peritreme of paratype female (ventral view, right peritreme of mite); (37) Aheatherella mira n. sp. peritreme of paratype female (ventral view, left peritreme of mite; sclerotised eXopodal portion of eXopodal-peritrematal shield shaded)." httpUri="https://zenodo.org/record/1292223/files/figure.png" pageId="23" pageNumber="464">36</figureCitation>
); female with setae
<emphasis id="B945BA1F525E734F5DD1FF243DBBE8A2" box="[700,752,222,243]" italics="true" pageId="23" pageNumber="464">st3–4</emphasis>
on common shield or
<emphasis id="B945BA1F525E734F5CA4FF243CAEE8A2" box="[969,997,222,243]" italics="true" pageId="23" pageNumber="464">
st
<quantity id="4CC9CBE8525E734F5CB4FF243CAEE8A2" box="[985,997,222,243]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="23" pageNumber="464" unit="in" value="3.0">3</quantity>
</emphasis>
in soft cuticle; dorsal shields sexually dimorphic: female (
<figureCitation id="130A7A88525E734F5E0BFF013EEEE940" box="[358,421,251,273]" captionStart="FIGURE 2" captionStartId="5.[151,250,1997,2019]" captionTargetBox="[201,1387,277,1878]" captionTargetId="figure@5.[151,1436,265,1888]" captionTargetPageId="5" captionText="FIGURE 2. Heatherella osleri n. sp. female holotype, dorsum." httpUri="https://zenodo.org/record/1292193/files/figure.png" pageId="23" pageNumber="464">Figs 2</figureCitation>
,
<figureCitation id="130A7A88525E734F5EDDFF013E83E940" box="[432,456,251,273]" captionStart="FIGURE 15" captionStartId="10.[151,250,1930,1952]" captionTargetBox="[206,1367,343,1897]" captionTargetId="figure@10.[195,1387,327,1916]" captionTargetPageId="10" captionText="FIGURE 15. Heatherella callimaulos walter, 1997, female holotype, dorsum. First pair of tubercles are somewhat concealed. The only marginal seta shown is r1. Two types of pores: 12 pairs large (shaded grey) and ten pairs of smaller pores (unshaded). Platelets behind main dorsal shield not obvious (dotted lines)." httpUri="https://zenodo.org/record/1292201/files/figure.png" pageId="23" pageNumber="464">15</figureCitation>
) with podonotal-mesonotal shield; males (
<figureCitation id="130A7A88525E734F5C08FF013CE1E940" box="[869,938,251,273]" captionStart="FIGURES 12–14" captionStartId="8.[151,264,1439,1461]" captionTargetBox="[213,1369,199,1413]" captionTargetId="figure@8.[212,1376,185,1424]" captionTargetPageId="8" captionText="FIGURES 12–14. Heatherella osleri n. sp. male (12) venter; (13) dorsum (marginal shields not shown laterally; torn anteriorly from marginal shield); (14) leg II, anterior view." httpUri="https://zenodo.org/record/1292199/files/figure.png" pageId="23" pageNumber="464">Fig. 13</figureCitation>
) with podonotal shield and three mesonotal shields (one medial, two lateral); female latinotal shield undivided; ventral seta
<emphasis id="B945BA1F525E734F5CFCFEE33CFCE97F" box="[913,951,281,302]" italics="true" pageId="23" pageNumber="464">JF3</emphasis>
reduced, on anal rim (
<figureCitation id="130A7A88525E734F5BE0FEE23B8FE97F" box="[1165,1220,280,302]" captionStart="FIGURE 1" captionStartId="4.[151,250,1999,2021]" captionTargetBox="[158,1436,230,1948]" captionTargetId="figure@4.[151,1436,227,1948]" captionTargetPageId="4" captionText="FIGURE 1. Heatherella osleri n. sp. female holotype, venter." httpUri="https://zenodo.org/record/1292191/files/figure.png" pageId="23" pageNumber="464">Fig. 1</figureCitation>
); male
<emphasis id="B945BA1F525E734F5A63FEE33A61E97F" box="[1294,1322,281,302]" italics="true" pageId="23" pageNumber="464">st5</emphasis>
on separate shield from
<emphasis id="B945BA1F525E734F5E39FECC3EC3E91A" box="[340,392,310,331]" italics="true" pageId="23" pageNumber="464">st1–4</emphasis>
(
<figureCitation id="130A7A88525E734F5EFBFECF3E97E91A" box="[406,476,309,331]" captionStart="FIGURES 12–14" captionStartId="8.[151,264,1439,1461]" captionTargetBox="[213,1369,199,1413]" captionTargetId="figure@8.[212,1376,185,1424]" captionTargetPageId="8" captionText="FIGURES 12–14. Heatherella osleri n. sp. male (12) venter; (13) dorsum (marginal shields not shown laterally; torn anteriorly from marginal shield); (14) leg II, anterior view." httpUri="https://zenodo.org/record/1292199/files/figure.png" pageId="23" pageNumber="464">Fig. 12</figureCitation>
) ...
<taxonomicName id="4C311D8E525E734F5D6DFECC3D39E91A" authorityName="walter" authorityYear="1997" box="[512,626,310,331]" class="Arachnida" family="Heatherellidae" genus="Heatherella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="23" pageNumber="464" phylum="Arthropoda" rank="genus">
<emphasis id="B945BA1F525E734F5D6DFECC3D39E91A" box="[512,626,310,331]" italics="true" pageId="23" pageNumber="464">Heatherella</emphasis>
</taxonomicName>
................................................................. 2
</keyLead>
</paragraph>
<paragraph id="8B8E660D525E734F5FFAFEA93AD0E9F2" blockId="23.[151,1436,221,830]" pageId="23" pageNumber="464">
<keyLead id="30C0C538525E734F5FFAFEA93AD0E9F2" pageId="23" pageNumber="464">
- Peritremes present, short (
<figureCitation id="130A7A88525E734F5EB0FEA93D63E938" box="[477,552,339,361]" captionStart="FIGURE 16" captionStartId="12.[151,250,2006,2028]" captionTargetBox="[168,1434,498,1982]" captionTargetId="figure@12.[151,1436,481,1985]" captionTargetPageId="12" captionText="FIGURE 16. Aheatherella mira n. sp. female holotype, venter." httpUri="https://zenodo.org/record/1292203/files/figure.png" pageId="23" pageNumber="464">Figs 16</figureCitation>
,
<figureCitation id="130A7A88525E734F5D5EFEA93D05E938" box="[563,590,339,361]" captionStart="FIGURES 34–37" captionStartId="22.[151,264,1832,1854]" captionTargetBox="[326,1233,232,1771]" captionTargetId="figure@22.[309,1273,196,1784]" captionTargetPageId="22" captionText="FIGURES 34–37. (34) Heatherella acanthocharis walter, 1997, dorsomarginal protuberances, pustules and pores of paratype female (P-s5 pustule; P-s6 pore; P-S1 protuberance), dorsal view of flattened specimen, right side; (35) Aheatherella mira n. sp., dorsomarginal protuberances, pustules and pores of paratype female (P-s5 pustule; P-s6 protuberance with closely associated seta), dorsal view of ventrolateral fragment with broken r-setae, left side; (36) Heatherella osleri n. sp., peritreme of paratype female (ventral view, right peritreme of mite); (37) Aheatherella mira n. sp. peritreme of paratype female (ventral view, left peritreme of mite; sclerotised eXopodal portion of eXopodal-peritrematal shield shaded)." httpUri="https://zenodo.org/record/1292223/files/figure.png" pageId="23" pageNumber="464">37</figureCitation>
); female with
<emphasis id="B945BA1F525E734F5DB4FEA93DBEE939" box="[729,757,339,360]" italics="true" pageId="23" pageNumber="464">st3</emphasis>
and
<emphasis id="B945BA1F525E734F5C48FEA93C0AE939" box="[805,833,339,360]" italics="true" pageId="23" pageNumber="464">st4</emphasis>
on separate platelets; dorsal shields monomorphic: both sexes with podonotal-mesonotal shield (
<figureCitation id="130A7A88525E734F5D4AFE8A3D25E9D7" box="[551,622,368,390]" captionStart="FIGURE 17" captionStartId="13.[151,250,1788,1810]" captionTargetBox="[161,1414,210,1732]" captionTargetId="figure@13.[151,1436,193,1766]" captionTargetPageId="13" captionText="FIGURE 17. Aheatherella mira n. sp. female holotype, dorsum." httpUri="https://zenodo.org/record/1292205/files/figure.png" pageId="23" pageNumber="464">Fig. 17</figureCitation>
); latinotal shield divided into podonotal and mesonotal regions; ventral seta
<emphasis id="B945BA1F525E734F5A24FE8B3A24E9D7" box="[1353,1391,369,390]" italics="true" pageId="23" pageNumber="464">JF3</emphasis>
normal, removed from anus (
<figureCitation id="130A7A88525E734F5EB5FE773D54E9F2" box="[472,543,397,419]" captionStart="FIGURE 16" captionStartId="12.[151,250,2006,2028]" captionTargetBox="[168,1434,498,1982]" captionTargetId="figure@12.[151,1436,481,1985]" captionTargetPageId="12" captionText="FIGURE 16. Aheatherella mira n. sp. female holotype, venter." httpUri="https://zenodo.org/record/1292203/files/figure.png" pageId="23" pageNumber="464">Fig. 16</figureCitation>
); male
<emphasis id="B945BA1F525E734F5D0BFE743DC9E9F2" box="[614,642,398,419]" italics="true" pageId="23" pageNumber="464">st5</emphasis>
on intercoxal shield (
<figureCitation id="130A7A88525E734F5C3FFE773CD3E9F2" box="[850,920,397,419]" captionStart="FIGURE 24" captionStartId="16.[151,250,1729,1751]" captionTargetBox="[173,1407,196,1716]" captionTargetId="figure@16.[151,1436,183,1722]" captionTargetPageId="16" captionText="FIGURE 24. Aheatherella mira n. sp. male, venter." httpUri="https://zenodo.org/record/1292211/files/figure.png" pageId="23" pageNumber="464">Fig. 24</figureCitation>
)...........................
<taxonomicName id="4C311D8E525E734F5B80FE743AD0E9F2" authorityName="Seeman &amp; Minor &amp; Baker &amp; Walter" authorityYear="2018" box="[1261,1435,398,419]" class="Arachnida" family="Heatherellidae" genus="Aheatherella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="23" pageNumber="464" phylum="Arthropoda" rank="species" species="mira">
<emphasis id="B945BA1F525E734F5B80FE743AD0E9F2" box="[1261,1435,398,419]" italics="true" pageId="23" pageNumber="464">Aheatherella mira</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C57EA8525E734F5FFAFE513AD7EADF" pageId="23" pageNumber="464">
<paragraph id="8B8E660D525E734F5FFAFE513AD0EA48" blockId="23.[151,1436,221,830]" pageId="23" pageNumber="464">
<keyLead id="30C0C538525E734F5FFAFE513AD0EA48" pageId="23" pageNumber="464">
2. Dorsal setae somewhat thickened, barbed, without ciliate crests, dissimilar to marginal (
<emphasis id="B945BA1F525E734F5B2FFE513B29E991" box="[1090,1122,427,448]" italics="true" pageId="23" pageNumber="464">r -R</emphasis>
) setae (
<figureCitation id="130A7A88525E734F5BDDFE513BB2E990" box="[1200,1273,427,449]" captionStart="FIGURE 15" captionStartId="10.[151,250,1930,1952]" captionTargetBox="[206,1367,343,1897]" captionTargetId="figure@10.[195,1387,327,1916]" captionTargetPageId="10" captionText="FIGURE 15. Heatherella callimaulos walter, 1997, female holotype, dorsum. First pair of tubercles are somewhat concealed. The only marginal seta shown is r1. Two types of pores: 12 pairs large (shaded grey) and ten pairs of smaller pores (unshaded). Platelets behind main dorsal shield not obvious (dotted lines)." httpUri="https://zenodo.org/record/1292201/files/figure.png" pageId="23" pageNumber="464">Fig. 15</figureCitation>
); seta
<emphasis id="B945BA1F525E734F5A56FE513A04E991" box="[1339,1359,427,448]" italics="true" pageId="23" pageNumber="464">z2</emphasis>
absent; marginal setae duplicated (22 pairs); anterior opisthogastric setae simple; 8–9 pairs of submarginal protuberances (always lacking protuberance
<emphasis id="B945BA1F525E734F5EDFFE1C3E95E9AA" box="[434,478,486,507]" italics="true" pageId="23" pageNumber="464">P -s6</emphasis>
, between
<emphasis id="B945BA1F525E734F5D2FFE1C3D1DE9AA" box="[578,598,486,507]" italics="true" pageId="23" pageNumber="464">s6</emphasis>
and
<emphasis id="B945BA1F525E734F5DEAFE1C3DD4E9AA" box="[647,671,486,507]" italics="true" pageId="23" pageNumber="464">S1</emphasis>
); protuberance posterior to seta
<emphasis id="B945BA1F525E734F5CB6FE1C3CBBE9AA" box="[987,1008,486,507]" italics="true" pageId="23" pageNumber="464">s5</emphasis>
a raised pustule, different from other protuberances............................................................................................. 3
</keyLead>
</paragraph>
<paragraph id="8B8E660D525E734F5FFAFDDB3AD7EADF" blockId="23.[151,1436,221,830]" pageId="23" pageNumber="464">
<keyLead id="30C0C538525E734F5FFAFDDB3AD7EADF" pageId="23" pageNumber="464">
- Dorsal setae bifurcate, with ciliate crests, similar to marginal (
<emphasis id="B945BA1F525E734F5C5AFDDB3C0BEA67" box="[823,832,545,566]" italics="true" pageId="23" pageNumber="464">r</emphasis>
-R) setae (
<figureCitation id="130A7A88525E734F5CCEFDDA3C95EA67" box="[931,990,544,566]" captionStart="FIGURE 2" captionStartId="5.[151,250,1997,2019]" captionTargetBox="[201,1387,277,1878]" captionTargetId="figure@5.[151,1436,265,1888]" captionTargetPageId="5" captionText="FIGURE 2. Heatherella osleri n. sp. female holotype, dorsum." httpUri="https://zenodo.org/record/1292193/files/figure.png" pageId="23" pageNumber="464">Fig. 2</figureCitation>
); seta
<emphasis id="B945BA1F525E734F5B71FDDB3B7AEA67" box="[1052,1073,545,566]" italics="true" pageId="23" pageNumber="464">z2</emphasis>
present; marginal setae highly hypertrichous (&gt; 24 pairs); opisthogastric setae all short, bulbous and barbed; ten pairs of submarginal protuberances (with protuberance
<emphasis id="B945BA1F525E734F5E7EFDA63E74EA20" box="[275,319,604,625]" italics="true" pageId="23" pageNumber="464">P -s6</emphasis>
between
<emphasis id="B945BA1F525E734F5EF6FDA63EE4EA20" box="[411,431,604,625]" italics="true" pageId="23" pageNumber="464">s6</emphasis>
and
<emphasis id="B945BA1F525E734F5EB3FDA63EBDEA20" box="[478,502,604,625]" italics="true" pageId="23" pageNumber="464">S1</emphasis>
); protuberance posterior to seta
<emphasis id="B945BA1F525E734F5C40FDA63C09EA20" box="[813,834,604,625]" italics="true" pageId="23" pageNumber="464">s5</emphasis>
not obviously different from other protuberances................................................................................................
<taxonomicName id="4C311D8E525E734F5B80FD833AD7EADF" authorityName="Seeman &amp; Minor &amp; Baker &amp; Walter" authorityYear="2018" box="[1261,1436,633,654]" class="Arachnida" family="Heatherellidae" genus="Heatherella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="23" pageNumber="464" phylum="Arthropoda" rank="species" species="osleri">
<emphasis id="B945BA1F525E734F5B80FD833AD7EADF" box="[1261,1436,633,654]" italics="true" pageId="23" pageNumber="464">Heatherella osleri</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C57EA8525E734F5FFAFD6F3AD0EB6F" pageId="23" pageNumber="464">
<paragraph id="8B8E660D525E734F5FFAFD6F3AD0EAB7" blockId="23.[151,1436,221,830]" pageId="23" pageNumber="464">
<keyLead id="30C0C538525E734F5FFAFD6F3AD0EAB7" pageId="23" pageNumber="464">
3. Adult sternal shield (bearing
<emphasis id="B945BA1F525E734F5E97FD6C3D5CEAFA" box="[506,535,662,683]" italics="true" pageId="23" pageNumber="464">st2</emphasis>
) rounded anteriorly in both sexes, without endopodal elements; dorsum lacking protuberance
<emphasis id="B945BA1F525E734F5F8CFD4E3E47EA98" box="[225,268,692,713]" italics="true" pageId="23" pageNumber="464">P -s4</emphasis>
between setae
<emphasis id="B945BA1F525E734F5EF3FD4E3EFFEA98" box="[414,436,692,713]" italics="true" pageId="23" pageNumber="464">z4</emphasis>
and
<emphasis id="B945BA1F525E734F5E8FFD4E3EBCEA98" box="[482,503,692,713]" italics="true" pageId="23" pageNumber="464">s4</emphasis>
; seta
<emphasis id="B945BA1F525E734F5D42FD4E3D0EEA98" box="[559,581,692,713]" italics="true" pageId="23" pageNumber="464">z6</emphasis>
posterior to
<emphasis id="B945BA1F525E734F5DD3FD4E3D9FEA98" box="[702,724,692,713]" italics="true" pageId="23" pageNumber="464">s6</emphasis>
, seta
<emphasis id="B945BA1F525E734F5C66FD4E3C6FEA98" box="[779,804,692,713]" italics="true" pageId="23" pageNumber="464">Z1</emphasis>
posterior to
<emphasis id="B945BA1F525E734F5CF3FD4E3CFDEA98" box="[926,950,692,713]" italics="true" pageId="23" pageNumber="464">S1</emphasis>
; seta
<emphasis id="B945BA1F525E734F5C83FD4E3B48EA98" box="[1006,1027,692,713]" italics="true" pageId="23" pageNumber="464">z5</emphasis>
placed obliquely to seta
<emphasis id="B945BA1F525E734F5B9FFD4E3A4EEA98" box="[1266,1285,692,713]" italics="true" pageId="23" pageNumber="464">j5</emphasis>
; male with seta
<emphasis id="B945BA1F525E734F5F8CFD2B3E4FEAB7" box="[225,260,721,742]" italics="true" pageId="23" pageNumber="464">av1</emphasis>
on genu II unmodified...............................................
<taxonomicName id="4C311D8E525E734F5B77FD2B3AD0EAB7" authority="Walter, 1997" authorityName="Walter" authorityYear="1997" box="[1050,1435,720,742]" class="Arachnida" family="Heatherellidae" genus="Heatherella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="23" pageNumber="464" phylum="Arthropoda" rank="species" species="acanthocharis">
<emphasis id="B945BA1F525E734F5B77FD2B3A51EAB7" box="[1050,1306,721,742]" italics="true" pageId="23" pageNumber="464">Heatherella acanthocharis</emphasis>
Walter, 1997
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8B8E660D525E734F5FFAFD143AD0EB6F" blockId="23.[151,1436,221,830]" pageId="23" pageNumber="464">
<keyLead id="30C0C538525E734F5FFAFD143AD0EB6F" pageId="23" pageNumber="464">
- Adult sternal shield (bearing
<emphasis id="B945BA1F525E734F5E94FD143D5FEB52" box="[505,532,750,771]" italics="true" pageId="23" pageNumber="464">st2</emphasis>
) angular anterior corners in both sexes, endopodal elements fused to shield; dorsum with protuberance
<emphasis id="B945BA1F525E734F5E28FCF63E3AEB70" box="[325,369,780,801]" italics="true" pageId="23" pageNumber="464">P -s4</emphasis>
(
<figureCitation id="130A7A88525E734F5EEDFCF13E83EB70" box="[384,456,779,801]" captionStart="FIGURE 15" captionStartId="10.[151,250,1930,1952]" captionTargetBox="[206,1367,343,1897]" captionTargetId="figure@10.[195,1387,327,1916]" captionTargetPageId="10" captionText="FIGURE 15. Heatherella callimaulos walter, 1997, female holotype, dorsum. First pair of tubercles are somewhat concealed. The only marginal seta shown is r1. Two types of pores: 12 pairs large (shaded grey) and ten pairs of smaller pores (unshaded). Platelets behind main dorsal shield not obvious (dotted lines)." httpUri="https://zenodo.org/record/1292201/files/figure.png" pageId="23" pageNumber="464">Fig. 15</figureCitation>
); seta
<emphasis id="B945BA1F525E734F5D64FCF63D55EB70" box="[521,542,780,801]" italics="true" pageId="23" pageNumber="464">z6</emphasis>
anterior to
<emphasis id="B945BA1F525E734F5DFFFCF63DEDEB70" box="[658,678,780,801]" italics="true" pageId="23" pageNumber="464">s6</emphasis>
, seta
<emphasis id="B945BA1F525E734F5D8CFCF63DB1EB70" box="[737,762,780,801]" italics="true" pageId="23" pageNumber="464">Z1</emphasis>
anterior to
<emphasis id="B945BA1F525E734F5C03FCF63CCDEB70" box="[878,902,780,801]" italics="true" pageId="23" pageNumber="464">S1</emphasis>
; seta
<emphasis id="B945BA1F525E734F5CADFCF63C9DEB70" box="[960,982,780,801]" italics="true" pageId="23" pageNumber="464">z5</emphasis>
almost transversely aligned with seta
<emphasis id="B945BA1F525E734F5A21FCF63A14EB70" box="[1356,1375,780,801]" italics="true" pageId="23" pageNumber="464">j5</emphasis>
; male with seta
<emphasis id="B945BA1F525E734F5E51FCD33E14EB6F" box="[316,351,809,830]" italics="true" pageId="23" pageNumber="464">av1</emphasis>
on genu II modified into small blunt spur...........................
<taxonomicName id="4C311D8E525E734F5B5EFCD33AD0EB6F" authority="Walter, 1997" authorityName="Walter" authorityYear="1997" box="[1075,1435,808,830]" class="Arachnida" family="Heatherellidae" genus="Heatherella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="23" pageNumber="464" phylum="Arthropoda" rank="species" species="callimaulos">
<emphasis id="B945BA1F525E734F5B5EFCD33A50EB6F" box="[1075,1307,809,830]" italics="true" pageId="23" pageNumber="464">Heatherella callimaulos</emphasis>
Walter, 1997
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>
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