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<document id="53F84155A1FCFB3758F8C3C96ACC81DB" ID-CLB-Dataset="29875" ID-DOI="10.11646/zootaxa.4434.3.3" ID-GBIF-Dataset="3ebb3cb7-3b8f-4c4d-8858-d37185292c28" ID-ISSN="1175-5326" ID-Zenodo-Dep="1292189" ID-ZooBank="8B95D692-E8F7-402B-8485-3D3AE20111E8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1529330577762" checkinUser="plazi" docAuthor="Seeman, Owen D., Minor, Maria, Baker, Michelle R. &amp; Walter, David Evans" docDate="2018" docId="0398D71B524073515FFAFE953D14EB90" docLanguage="en" docName="zootaxa.4434.3.3.pdf.imf" docOrigin="Zootaxa 4434 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Heatherella acanthocharis Walter 1997" docType="treatment" docVersion="7" lastPageNumber="450" masterDocId="FFA1AF63524973585F6DFFFA3F4BE851" masterDocTitle="A revision of the Heatherellidae (Parasitiformes: Mesostigmata) with a new genus and two new species from Australasia" masterLastPageNumber="465" masterPageNumber="441" pageNumber="450" updateTime="1698549773811" updateUser="ExternalLinkService">
<mods:mods id="37D12583B722465A0A91E38923EB87B8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B7043BF677065C006E430A4FE0C24BC5">
<mods:title id="1B150DF2987190B7BAEDE42C9C9E721A">A revision of the Heatherellidae (Parasitiformes: Mesostigmata) with a new genus and two new species from Australasia</mods:title>
</mods:titleInfo>
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<mods:namePart id="9F08E45B5085E9573D1CE1001294E88A">Seeman, Owen D.</mods:namePart>
</mods:name>
<mods:name id="A9D64F5DDF37718F961688B94647912F" type="personal">
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<mods:namePart id="80640DCC3D1602E8DE24B51A5524CBB9">Minor, Maria</mods:namePart>
</mods:name>
<mods:name id="9138B8569D44BDA4794BE45D62089456" type="personal">
<mods:role id="A944ACA3D7891989F1D131EDF42446FD">
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<mods:namePart id="D1573EF7ABF15470AD12FB2A0C16FFA5">Baker, Michelle R.</mods:namePart>
</mods:name>
<mods:name id="57F0ADC5FEAFF523AA0ABB486BE656E6" type="personal">
<mods:role id="5F8F7D78A9AE25E48DAEA2E427237AC6">
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<mods:namePart id="CD044CE5D0B270798F0E6B6D529C7D93">Walter, David Evans</mods:namePart>
</mods:name>
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<mods:title id="286FDC9418BB628D74EAA36B10EBD6A4">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="EC5497DD53E5189A182CE5412252DC77">
<mods:date id="89AC07F73194B08A1B295A9F11E558B9">2018</mods:date>
<mods:detail id="96ABDD87DF3FEA26AE0274446C91BCF0" type="pubDate">
<mods:number id="20658B9F0436AD9F1DA7F6BF2AD16764">2018-06-18</mods:number>
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<mods:detail id="3A37E0CB68703F9ED8845D9B681F4248" type="volume">
<mods:number id="BFD4833C05226A9A94AAFD0878302320">4434</mods:number>
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<mods:number id="C46511DA04066B7FE601C8D30356BC5F">3</mods:number>
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<mods:classification id="FE90DBF7DCC50D5124F9EFA7D54EA7B0">journal article</mods:classification>
<mods:identifier id="68638C592D88B0EE07D92B42E4B0573E" type="CLB-Dataset">29875</mods:identifier>
<mods:identifier id="07CA18D15A3F58EDEEB54C33656713B4" type="DOI">10.11646/zootaxa.4434.3.3</mods:identifier>
<mods:identifier id="154673CB04A09C3B120C441C6EF41F7C" type="GBIF-Dataset">3ebb3cb7-3b8f-4c4d-8858-d37185292c28</mods:identifier>
<mods:identifier id="D8899F290EBF3CF995609F9160FB8796" type="ISSN">1175-5326</mods:identifier>
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<treatment id="0398D71B524073515FFAFE953D14EB90" ID-DOI="http://doi.org/10.5281/zenodo.5966635" ID-GBIF-Taxon="145968994" ID-Zenodo-Dep="5966635" LSID="urn:lsid:plazi:treatment:0398D71B524073515FFAFE953D14EB90" httpUri="http://treatment.plazi.org/id/0398D71B524073515FFAFE953D14EB90" lastPageNumber="450" pageId="9" pageNumber="450">
<subSubSection id="C32B3586524073515FFAFE953E6CE9FA" pageId="9" pageNumber="450" type="nomenclature">
<paragraph id="8B8E660D524073515FFAFE953D34E9DB" blockId="9.[151,639,367,427]" box="[151,639,367,394]" pageId="9" pageNumber="450">
<heading id="D0C6D161524073515FFAFE953D34E9DB" bold="true" box="[151,639,367,394]" fontSize="11" level="1" pageId="9" pageNumber="450" reason="1">
<taxonomicName id="4C311D8E524073515FFAFE953D34E9DB" authority="Walter, 1997" authorityName="Walter" authorityYear="1997" box="[151,639,367,394]" class="Arachnida" family="Heatherellidae" genus="Heatherella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="450" phylum="Arthropoda" rank="species" species="acanthocharis">
<emphasis id="B945BA1F524073515FFAFE953D34E9DB" bold="true" box="[151,639,367,394]" pageId="9" pageNumber="450">
<emphasis id="B945BA1F524073515FFAFE953E92E9D8" bold="true" box="[151,473,367,393]" italics="true" pageId="9" pageNumber="450">Heatherella acanthocharis</emphasis>
Walter, 1997
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B8E660D524073515FFAFE683E6CE9FA" blockId="9.[151,639,367,427]" box="[151,295,402,427]" pageId="9" pageNumber="450">
(
<figureCitation id="130A7A88524073515FF2FE683E54E9FA" box="[159,287,402,427]" captionStart-0="FIGURES 30–31" captionStart-1="FIGURES 32–33" captionStart-2="FIGURES 34–37" captionStartId-0="20.[151,264,1814,1836]" captionStartId-1="21.[151,264,1907,1929]" captionStartId-2="22.[151,264,1832,1854]" captionTargetBox-0="[153,1434,1302,1788]" captionTargetBox-1="[184,1389,196,1839]" captionTargetBox-2="[326,1233,232,1771]" captionTargetId-0="figure@20.[151,1436,1302,1788]" captionTargetId-1="figure@21.[184,1389,196,1839]" captionTargetId-2="figure@22.[309,1273,196,1784]" captionTargetPageId-0="20" captionTargetPageId-1="21" captionTargetPageId-2="22" captionText-0="FIGURES 30–31. Heatherella acanthocharis walter, 1997, scanning electron micrographs. (30) idiosoma, showing sloughed certotegument, scale bar = 100 µm; (31) podonotum, with mostly intact cerotegument (fractured behind setae j3 and laterad z3)." captionText-1="FIGURES 32–33. Heatherella acanthocharis walter, 1997, scanning electron micrographs. (32) lateral habitus, with protuberances coloured orange and the enlarged pore (P-s4, white arrow) and raised pustule (P-s5) coloured yellow. (33) detail of protuberance P-S1 (rotated approXimately 45° clockwise); note cleft margin and pores deep within." captionText-2="FIGURES 34–37. (34) Heatherella acanthocharis walter, 1997, dorsomarginal protuberances, pustules and pores of paratype female (P-s5 pustule; P-s6 pore; P-S1 protuberance), dorsal view of flattened specimen, right side; (35) Aheatherella mira n. sp., dorsomarginal protuberances, pustules and pores of paratype female (P-s5 pustule; P-s6 protuberance with closely associated seta), dorsal view of ventrolateral fragment with broken r-setae, left side; (36) Heatherella osleri n. sp., peritreme of paratype female (ventral view, right peritreme of mite); (37) Aheatherella mira n. sp. peritreme of paratype female (ventral view, left peritreme of mite; sclerotised eXopodal portion of eXopodal-peritrematal shield shaded)." httpUri-0="https://zenodo.org/record/1292219/files/figure.png" httpUri-1="https://zenodo.org/record/1292221/files/figure.png" httpUri-2="https://zenodo.org/record/1292223/files/figure.png" pageId="9" pageNumber="450">Figs 30–34</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32B3586524073515FFAFE203D23E9A1" box="[151,616,474,496]" pageId="9" pageNumber="450" type="description">
<paragraph id="8B8E660D524073515FFAFE203D23E9A1" blockId="9.[151,616,474,496]" box="[151,616,474,496]" pageId="9" pageNumber="450">
<taxonomicName id="4C311D8E524073515FFAFE203D7AE9A1" authority="walter, 1997" authorityName="Walter" authorityYear="1997" box="[151,561,474,496]" class="Arachnida" family="Heatherellidae" genus="Heatherella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="464" phylum="Arthropoda" rank="species" species="acanthocharis">
<emphasis id="B945BA1F524073515FFAFE203EEDE9A1" box="[151,422,474,496]" italics="true" pageId="9" pageNumber="450">Heatherella acanthocharis</emphasis>
walter, 1997
</taxonomicName>
: 168.
</paragraph>
</subSubSection>
<subSubSection id="C32B3586524073515FFAFDE73C12EB5F" pageId="9" pageNumber="450" type="diagnosis">
<paragraph id="8B8E660D524073515FFAFDE73C12EB5F" blockId="9.[151,1436,541,962]" pageId="9" pageNumber="450">
<emphasis id="B945BA1F524073515FFAFDE73E41EA67" bold="true" box="[151,266,541,566]" pageId="9" pageNumber="450">Diagnosis</emphasis>
: With the characteristics of the genus. Adult: sternal shield (bearing
<emphasis id="B945BA1F524073515B5DFDE43B1BEA67" box="[1072,1104,542,566]" italics="true" pageId="9" pageNumber="450">st2</emphasis>
) rounded anteriorly, without endopodal elements; most ventral setae short, smooth, including setae
<emphasis id="B945BA1F524073515CD1FDBB3B4CEA08" box="[956,1031,577,601]" italics="true" pageId="9" pageNumber="450">ZV2–3</emphasis>
; setae
<emphasis id="B945BA1F524073515B35FDBB3BCFEA08" box="[1112,1156,577,601]" italics="true" pageId="9" pageNumber="450">JV4</emphasis>
,
<emphasis id="B945BA1F524073515BF9FDBB3B88EA08" box="[1172,1219,577,601]" italics="true" pageId="9" pageNumber="450">ZV4</emphasis>
thickened, barbed; dorsal setae thickened, barbed, without ciliate crests; marginal setae bifurcate but crest small, not highly ciliate; dorsal shield heavily ornamented; idiosomal margin with seven pairs of idiosomatic protuberances (protuberances
<emphasis id="B945BA1F524073515FFAFD543F9EEA97" box="[151,213,686,710]" italics="true" pageId="9" pageNumber="450">P - s4</emphasis>
and
<emphasis id="B945BA1F524073515E79FD543E03EA97" box="[276,328,685,710]" italics="true" pageId="9" pageNumber="450">P -s6</emphasis>
absent), distinct pustule
<emphasis id="B945BA1F524073515D1DFD543DE8EA94" box="[624,675,686,709]" italics="true" pageId="9" pageNumber="450">P -s5</emphasis>
present; dorsum lacking seta
<emphasis id="B945BA1F524073515B6BFD543B55EA97" box="[1030,1054,686,710]" italics="true" pageId="9" pageNumber="450">z2</emphasis>
; seta
<emphasis id="B945BA1F524073515B05FD573BCAEA97" box="[1128,1153,685,710]" italics="true" pageId="9" pageNumber="450">z6</emphasis>
posterior to
<emphasis id="B945BA1F524073515A71FD573A7EEA97" box="[1308,1333,685,710]" italics="true" pageId="9" pageNumber="450">s6</emphasis>
, seta
<emphasis id="B945BA1F524073515A13FD543AD7EA97" box="[1406,1436,686,710]" italics="true" pageId="9" pageNumber="450">Z1</emphasis>
posterior to
<emphasis id="B945BA1F524073515E72FD283E70EABB" box="[287,315,722,746]" italics="true" pageId="9" pageNumber="450">S1</emphasis>
; seta
<emphasis id="B945BA1F524073515E11FD283EDDEAB8" box="[380,406,722,745]" italics="true" pageId="9" pageNumber="450">z5</emphasis>
placed obliquely to seta
<emphasis id="B945BA1F524073515DC3FD283D8FEAB8" box="[686,708,722,745]" italics="true" pageId="9" pageNumber="450">j5</emphasis>
; marginal
<emphasis id="B945BA1F524073515C50FD283C29EAB8" box="[829,866,722,745]" italics="true" pageId="9" pageNumber="450">r -R</emphasis>
series setae duplicated, 22 pairs. Female: setae
<emphasis id="B945BA1F524073515A16FD283AD7EABB" box="[1403,1436,722,746]" italics="true" pageId="9" pageNumber="450">st3</emphasis>
and
<emphasis id="B945BA1F524073515FABFD0C3FACEB5F" box="[198,231,758,782]" italics="true" pageId="9" pageNumber="450">st4</emphasis>
on single shield. Male: seta
<emphasis id="B945BA1F524073515D4BFD0C3D04EB5F" box="[550,591,758,782]" italics="true" pageId="9" pageNumber="450">av1</emphasis>
on genu II unmodified.
</paragraph>
</subSubSection>
<subSubSection id="C32B3586524073515FAAFCE33B01EB07" pageId="9" pageNumber="450" type="materials_examined">
<paragraph id="8B8E660D524073515FAAFCE33B01EB07" blockId="9.[151,1436,541,962]" pageId="9" pageNumber="450">
<materialsCitation id="3B596C50524073515FAAFCE33B01EB07" ID-GBIF-Occurrence="1849335294" collectingDate="1998-07-14" collectorName="D. E. Walter &amp; H. Klompen" county="New" latitude="-26.666666" location="Lobster Creek" longLatPrecision="1241" longitude="152.65" municipality="Conondale Ranges" pageId="9" pageNumber="450" specimenCount="1" specimenCount-female="1" stateProvince="Queensland">
<emphasis id="B945BA1F524073515FAAFCE33EA5EB63" bold="true" box="[199,494,793,818]" pageId="9" pageNumber="450">
<collectingCounty id="62EF1E81524073515FAAFCE33FB7EB63" box="[199,252,793,818]" pageId="9" pageNumber="450">New</collectingCounty>
material examined:
</emphasis>
<specimenCount id="9D37AD84524073515E9AFCE03D17EB63" box="[503,604,793,818]" pageId="9" pageNumber="450" type="female">1 female</specimenCount>
, 1 deutonymph,
<collectingRegion id="49F5A8EF524073515C76FCE33CE9EB63" box="[795,930,793,818]" country="Australia" name="Queensland" pageId="9" pageNumber="450">Queensland</collectingRegion>
,
<collectingMunicipality id="6BEAFC77524073515CC2FCE33BCDEB63" box="[943,1158,793,818]" pageId="9" pageNumber="450">Conondale Ranges</collectingMunicipality>
,
<location id="8EEE30D6524073515BFEFCE33A7CEB63" LSID="urn:lsid:plazi:treatment:0398D71B524073515FFAFE953D14EB90:8EEE30D6524073515BFEFCE33A7CEB63" box="[1171,1335,793,818]" county="New" latitude="-26.666666" longLatPrecision="1241" longitude="152.65" municipality="Conondale Ranges" name="Lobster Creek" pageId="9" pageNumber="450" stateProvince="Queensland">Lobster Creek</location>
,
<geoCoordinate id="EE0500CA524073515A2EFCE33AD7EB63" box="[1347,1436,793,818]" direction="south" orientation="latitude" pageId="9" pageNumber="450" precision="925" value="-26.666666">26°40′S</geoCoordinate>
<geoCoordinate id="EE0500CA524073515FFAFCC73E49EB07" box="[151,258,829,854]" direction="east" orientation="longitude" pageId="9" pageNumber="450" precision="925" value="152.65">152°39′E</geoCoordinate>
,
<date id="FF8F40CD524073515E60FCC43EEAEB07" box="[269,417,829,854]" pageId="9" pageNumber="450" value="1998-07-14">
<collectingDate id="EFCBB925524073515E60FCC43EEAEB07" box="[269,417,829,854]" pageId="9" pageNumber="450" value="1998-07-14">14 July 1998</collectingDate>
</date>
,
<collectorName id="26C403DB524073515EC1FCC43D7BEB07" box="[428,560,829,854]" pageId="9" pageNumber="450">D.E. Walter</collectorName>
&amp;
<collectorName id="26C403DB524073515D39FCC43DAAEB07" box="[596,737,829,854]" pageId="9" pageNumber="450">H. Klompen</collectorName>
, ex riparian rainforest leaf litter.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32B3586524073515FAAFC9B3D14EB90" pageId="9" pageNumber="450" type="discussion">
<paragraph id="8B8E660D524073515FAAFC9B3D14EB90" blockId="9.[151,1436,541,962]" pageId="9" pageNumber="450">
<materialsCitation id="3B596C50524073515FAAFC9B3D14EB90" ID-GBIF-Occurrence="1849336033" location="Walter" pageId="9" pageNumber="450" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
<emphasis id="B945BA1F524073515FAAFC9B3E04EB2B" bold="true" box="[199,335,865,890]" pageId="9" pageNumber="450">Comments.</emphasis>
Our diagnosis above is based on
<location id="8EEE30D6524073515DBFFC9B3C57EB2B" LSID="urn:lsid:plazi:treatment:0398D71B524073515FFAFE953D14EB90:8EEE30D6524073515DBFFC9B3C57EB2B" box="[722,796,865,890]" name="Walter" pageId="9" pageNumber="450" stateProvince="Queensland">Walter</location>
(1997) and the re-examination of specimens held in the
<collectingRegion id="49F5A8EF524073515FFAFC7F3E50EBCF" box="[151,283,901,926]" country="Australia" name="Queensland" pageId="9" pageNumber="450">Queensland</collectingRegion>
Museum, including the
<typeStatus id="548AD8AF524073515D5DFC7F3DDDEBCF" box="[560,662,901,926]" pageId="9" pageNumber="450" type="holotype">holotype</typeStatus>
. The new specimens examined are in poor condition but include the only known nymph of
<taxonomicName id="4C311D8E524073515EFBFC503D13EB93" authorityName="Walter" authorityYear="1997" box="[406,600,937,962]" class="Arachnida" family="Heatherellidae" genus="Heatherella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="450" phylum="Arthropoda" rank="species" species="acanthocharis">
<emphasis id="B945BA1F524073515EFBFC503EE1EB90" box="[406,426,938,961]" italics="true" pageId="9" pageNumber="450">H</emphasis>
.
<emphasis id="B945BA1F524073515ED5FC533D13EB93" box="[440,600,937,962]" italics="true" pageId="9" pageNumber="450">acanthocharis</emphasis>
</taxonomicName>
.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>
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