549 lines
59 KiB
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549 lines
59 KiB
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<document id="C399299B90BF81BA28E58BE12205E07D" ID-DOI="10.5281/zenodo.10831085" ID-ISSN="1942-1354" ID-Zenodo-Dep="10831085" ID-ZooBank="C1D5B819-A964-4679-B090-84CDBBC59D6A" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1688389961601" checkinUser="felipe" docAuthor="Otto, Robert L., Muona, Jyrki & Córdoba-Alfaro, Jim" docDate="2023" docId="954287CDB71EFFA569F0FDD846C85FF4" docLanguage="en" docName="InsectaMundi.2023.0991.1-36.pdf" docOrigin="Insecta Mundi 2023 (991)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.4:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="4" docTitle="Onichodon confluentus Otto, Muona and Cordoba-Alfaro 2023, new species" docType="treatment" docVersion="4" lastPageNumber="17" masterDocId="697BFFB5B70FFFB76930FFD643695A4D" masterDocTitle="A new genus and sixteen new species of false click beetles (Coleoptera: Eucnemidae) described from the Heredia Province of Costa Rica with several additional records from the Osa Peninsula and Panama" masterLastPageNumber="36" masterPageNumber="1" pageNumber="16" updateTime="1710774778166" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
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<mods:mods id="8494D3840CBBFF353878135CA0865E92" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="2BB16BC4C1D5E0ABC56B7AF3F3B9159A">
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<mods:title id="AD77F73416B9780965EBC89CC127E1FE">A new genus and sixteen new species of false click beetles (Coleoptera: Eucnemidae) described from the Heredia Province of Costa Rica with several additional records from the Osa Peninsula and Panama</mods:title>
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</mods:titleInfo>
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<mods:name id="49D5F96B432B23BBAE8FE0AB6069416B" type="personal">
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<mods:role id="5C0A9A4DD6D970654EA19AFD5534214D">
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<mods:roleTerm id="2EF9F2FAB9AF6A479AA95686258B0156">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F676EA3A59CFC3E751E9F9EAC770F2B3">Otto, Robert L.</mods:namePart>
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<mods:affiliation id="E7927846FF70D22105D46FBAA0751BAF">W 4806 Chrissie Circle, Shawano, WI 54166, U. S. A.</mods:affiliation>
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</mods:name>
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<mods:name id="F5FF6161452F86F99AE218563BD7E4B5" type="personal">
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<mods:role id="9110559E4183A5C5324D3E810DF2D8F9">
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<mods:roleTerm id="D701EBCB046964876C7A5732D64C9234">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="19771C64AA778F995C3FB82A737EBC2C">Muona, Jyrki</mods:namePart>
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<mods:affiliation id="3A6DB8F66CEA0E01B3357C5946AC5A16">Finnish Museum of Natural History, Zoology Unit PO Box 17, University of Helsinki, Finland FIN- 00014</mods:affiliation>
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</mods:name>
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<mods:name id="5D01DB62C46455FDA7C78021966CFCF7" type="personal">
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<mods:role id="A1CC6FE2EA5FFAB9D5B9BAF2BB3D877E">
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<mods:roleTerm id="2AC1CBC9C3A1FD2CE245BEE641EB712A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6EBFBA4E841DF390FD1ED23F9A8D3793">Córdoba-Alfaro, Jim</mods:namePart>
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<mods:affiliation id="F94987EE2B1F6BFB6FE7E9FC5CF3D22A">Insectopia Insect Museum, BioSur Foundation Puerto Jiménez, Península de Osa, Costa Rica jim. cordoba @ biosurcorcovado. org</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="0F125C2B4CAC27FD4DFBC7E7A5AE89E3">text</mods:typeOfResource>
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<mods:relatedItem id="5459A0B47B68F061FE82A1C991128B7A" type="host">
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<mods:titleInfo id="7E3F4C410C7AC95DCB4202AF63D3E859">
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<mods:title id="4C8AA6ABDD3E40D9D6A70A6B83207F56">Insecta Mundi</mods:title>
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</mods:titleInfo>
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<mods:part id="8588830B38AE69A2DBDBCAAFF85C73BE">
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<mods:date id="7206FBE7F8E4D79BE4044483DED86E6B">2023</mods:date>
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<mods:detail id="638974C3871D9A54ABFB73ECC437E1DD" type="pubDate">
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<mods:number id="233D03896B09C49CCE01EAF2B64EEF97">2023-05-26</mods:number>
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</mods:detail>
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<mods:detail id="C4506BAAB06B88DEE352B87973B45A2D" type="volume">
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<mods:number id="38A083F6884F5374B9A5BA89F65424C8">2023</mods:number>
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</mods:detail>
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<mods:detail id="32FE37753148953A68E606FB59B31E17" type="issue">
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<mods:number id="566350CC608822098DC0C8AEF0B999E5">991</mods:number>
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</mods:detail>
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<mods:extent id="6DE6E6FF1BD31FE82DC6CAAD393709CE" unit="page">
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<mods:start id="6D3F918832828515D256EDF93D71FAD2">1</mods:start>
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<mods:end id="463481264A260C3F3775F1D1DE653ABA">36</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="333939D8C1875782845149BB73A28D87">journal article</mods:classification>
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<mods:identifier id="F416F1DA82FFB410631D379827DA6876" type="DOI">10.5281/zenodo.10831085</mods:identifier>
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<mods:identifier id="9D6F88AC3D799191767503887B8F9D38" type="ISSN">1942-1354</mods:identifier>
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<mods:identifier id="086BA0676191499B9A86F67C6173BD6B" type="Zenodo-Dep">10831085</mods:identifier>
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<mods:identifier id="A3E8A2F5AE9544E01898AFA17A704392" type="ZooBank">C1D5B819-A964-4679-B090-84CDBBC59D6A</mods:identifier>
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</mods:mods>
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<treatment id="954287CDB71EFFA569F0FDD846C85FF4" ID-DOI="http://doi.org/10.5281/zenodo.8109091" ID-Zenodo-Dep="8109091" LSID="urn:lsid:plazi:treatment:954287CDB71EFFA569F0FDD846C85FF4" httpUri="http://treatment.plazi.org/id/954287CDB71EFFA569F0FDD846C85FF4" lastPageId="18" lastPageNumber="17" pageId="17" pageNumber="16">
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<subSubSection id="55F16550B71EFFA669F0FDD847545866" box="[192,1085,525,555]" pageId="17" pageNumber="16" type="nomenclature">
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<paragraph id="1D5436DBB71EFFA669F0FDD847545866" blockId="17.[192,1085,525,588]" box="[192,1085,525,555]" pageId="17" pageNumber="16">
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<heading id="461C81B7B71EFFA669F0FDD847545866" box="[192,1085,525,555]" fontSize="12" level="2" pageId="17" pageNumber="16" reason="2">
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<emphasis id="2F9FEAC9B71EFFA669F0FDD847545866" bold="true" box="[192,1085,525,555]" pageId="17" pageNumber="16">
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<taxonomicName id="DAEB4D58B71EFFA669F0FDD840F35867" authority="Otto, Muona and Cordoba-Alfaro" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[192,922,525,555]" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="confluentus" status="sp. nov.">
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<emphasis id="2F9FEAC9B71EFFA669F0FDD8428A5866" bold="true" box="[192,483,526,555]" italics="true" pageId="17" pageNumber="16">Onichodon confluentus</emphasis>
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Otto, Muona and Córdoba-Alfaro
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</taxonomicName>
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,
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<taxonomicNameLabel id="34AC57B2B71EFFA66A98FDD847545866" box="[936,1085,526,555]" pageId="17" pageNumber="16" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="55F16550B71EFFA669F0FDE442345801" box="[192,349,562,588]" pageId="17" pageNumber="16" type="description">
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<paragraph id="1D5436DBB71EFFA669F0FDE442345801" blockId="17.[192,1085,525,588]" box="[192,349,562,588]" pageId="17" pageNumber="16">
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<figureCitation id="85D02A5EB71EFFA669F0FDE442345801" box="[192,349,562,588]" captionStart-0="Figures 27–30" captionStart-1="Figures 31–34" captionStartId-0="16.[192,279,631,657]" captionStartId-1="18.[192,279,656,682]" captionTargetBox-0="[378,1254,213,601]" captionTargetBox-1="[353,1279,213,626]" captionTargetId-0="figure-67@16.[378,1254,213,601]" captionTargetId-1="figure-451@18.[353,1279,213,626]" captionTargetPageId-0="16" captionTargetPageId-1="18" captionText-0="Figures 27–30. Costa Rican macraulacine eucnemids. 27) Maelodrus costaricensis sp. nov., dorsal habitus. 28) Maelodrus costaricensis sp. nov., lateral habitus. 29) Maelodrus costaricensis sp. nov., ventral habitus. 30) Onichodon confluentus sp. nov., male holotype, dorsal habitus. (Scale: 27–29 = 1.0 mm; 30 = 5.0 mm)" captionText-1="Figures 31–34. Costa Rican macraulacine eucnemids. 31) Onichodon confluentus sp. nov., male holotype, ventral habitus. 32) Onichodon confluentus sp. nov., female allotype, dorsal habitus. 33) Onichodon rufus sp. nov., dorsal habitus. 34) Onichodon rufus sp. nov., ventral habitus. (Scale: 31–32 = 5.0 mm; 33–34 = 1.0 mm)" figureDoi-0="http://doi.org/10.5281/zenodo.10831097" figureDoi-1="http://doi.org/10.5281/zenodo.10831099" httpUri-0="https://zenodo.org/record/10831097/files/figure.png" httpUri-1="https://zenodo.org/record/10831099/files/figure.png" pageId="17" pageNumber="16">Figures 30–32</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="55F16550B71EFFA669F0FD8D409858DA" pageId="17" pageNumber="16" type="diagnosis">
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<paragraph id="1D5436DBB71EFFA669F0FD8D409858DA" blockId="17.[192,1440,603,664]" pageId="17" pageNumber="16">
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<emphasis id="2F9FEAC9B71EFFA669F0FD8D42555838" bold="true" box="[192,316,603,629]" pageId="17" pageNumber="16">Diagnosis.</emphasis>
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The new eucnemid species can be readily distinguished by the presence of confluent surfaces of the frons from any members of
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<taxonomicName id="DAEB4D58B71EFFA668C7FDA8412F58DA" box="[503,582,638,663]" class="Insecta" family="Eucnemidae" genus="Fornax" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="genus">
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<emphasis id="2F9FEAC9B71EFFA668C7FDA8412F58DA" box="[503,582,638,663]" italics="true" pageId="17" pageNumber="16">Fornax</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="DAEB4D58B71EFFA66B4DFDAB419F58DA" authorityName="Newman" authorityYear="1838" box="[637,758,637,663]" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="genus">
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<emphasis id="2F9FEAC9B71EFFA66B4DFDAB419F58DA" box="[637,758,637,663]" italics="true" pageId="17" pageNumber="16">Onichodon</emphasis>
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</taxonomicName>
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present in
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<collectingCountry id="65FC764BB71EFFA66A42FDAB408458DA" box="[882,1005,637,663]" name="Costa Rica" pageId="17" pageNumber="16">Costa Rica</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="55F16550B71EFFA669F0FD7146EE5C60" pageId="17" pageNumber="16" type="materials_examined">
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<paragraph id="1D5436DBB71EFFA669F0FD71416959C6" blockId="17.[191,1441,679,1582]" pageId="17" pageNumber="16">
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<emphasis id="2F9FEAC9B71EFFA669F0FD7142C5588C" bold="true" box="[192,428,679,705]" pageId="17" pageNumber="16">Type material. Male</emphasis>
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<materialsCitation id="AD833C86B71EFFA66883FD71429559C6" collectingDate="1993-06-02" collectingDateMax="2001-04-09" collectingDateMin="1993-06-02" collectionCode="ALAS, R, INBC" collectorName="L. Otto" country="Costa Rica" elevation="150" latitude="10.433333" location="Bosque" longLatPrecision="1298" longitude="-84.01667" municipality="Est. Bio. La Selva" pageId="17" pageNumber="16" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Heredia" typeStatus="holotype">
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<emphasis id="2F9FEAC9B71EFFA66883FD714173588C" bold="true" box="[435,538,679,705]" pageId="17" pageNumber="16">
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<typeStatus id="C2508879B71EFFA66883FD714173588C" box="[435,538,679,705]" pageId="17" pageNumber="16" type="holotype">holotype</typeStatus>
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</emphasis>
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: “
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<collectingCountry id="65FC764BB71EFFA66B02FD7141BF588C" box="[562,726,679,705]" name="Costa Rica" pageId="17" pageNumber="16">COSTA RICA</collectingCountry>
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:
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<collectingRegion id="DF2FF839B71EFFA66BD1FD714054588C" box="[737,829,679,705]" country="Costa Rica" name="Heredia" pageId="17" pageNumber="16">Heredia</collectingRegion>
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;
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<collectingMunicipality id="FD30ACA1B71EFFA66A79FD71476C588C" box="[841,1029,679,705]" pageId="17" pageNumber="16">Est.Bio. La Selva</collectingMunicipality>
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, 50–;
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<quantity id="DA139B3EB71EFFA66D79FD7147F9588C" box="[1097,1168,679,706]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="16" unit="m" value="150.0">
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<elevation id="96C6D1E8B71EFFA66D79FD7147F9588C" box="[1097,1168,679,706]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="16" unit="m" value="150.0">150 m</elevation>
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</quantity>
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<geoCoordinate id="78DF501CB71EFFA66DA8FD71479A588C" box="[1176,1267,679,705]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="17" pageNumber="16" precision="925" value="10.433333">10°26′N</geoCoordinate>
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<geoCoordinate id="78DF501CB71EFFA66DCAFD714634588C" box="[1274,1373,679,705]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="17" pageNumber="16" precision="925" value="-84.01667">84°01′W</geoCoordinate>
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; Proy.
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<collectionCode id="7BFAAE1EB71EFFA669F0FD1F426F58AE" box="[192,262,713,739]" pageId="17" pageNumber="16">ALAS</collectionCode>
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, INBio-OET” / “M/15/121;
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<date id="6955101BB71EFFA66B07FD1F41B858AE" box="[567,721,712,739]" pageId="17" pageNumber="16" value="1993-06-02">
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<collectingDate id="7911E9F3B71EFFA66B07FD1F41B858AE" box="[567,721,712,739]" pageId="17" pageNumber="16" value="1993-06-02">02 Junio 1993</collectingDate>
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</date>
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;
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<location id="18346000B71EFFA66BEAFD1F404458AE" LSID="urn:lsid:plazi:treatment:954287CDB71EFFA569F0FDD846C85FF4:18346000B71EFFA66BEAFD1F404458AE" box="[730,813,713,739]" country="Costa Rica" latitude="10.433333" longLatPrecision="1298" longitude="-84.01667" municipality="Est. Bio. La Selva" name="Bosque" pageId="17" pageNumber="16" stateProvince="Heredia">Bosque</location>
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secundario” / “
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<emphasis id="2F9FEAC9B71EFFA66AE6FD1E471B58AF" bold="true" box="[982,1138,712,738]" pageId="17" pageNumber="16">
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<typeStatus id="C2508879B71EFFA66AE6FD1E470458AF" box="[982,1133,712,738]" pageId="17" pageNumber="16" type="holotype">HOLOTYPE</typeStatus>
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:
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</emphasis>
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;
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<taxonomicName id="DAEB4D58B71EFFA66D4DFD1E479F58AF" authorityName="Newman" authorityYear="1838" box="[1149,1270,712,738]" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="genus">
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<emphasis id="2F9FEAC9B71EFFA66D4DFD1E479F58AF" box="[1149,1270,712,738]" italics="true" pageId="17" pageNumber="16">Onichodon</emphasis>
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</taxonomicName>
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;
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<taxonomicName id="DAEB4D58B71EFFA66C32FD1E46E958AF" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[1282,1408,712,738]" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="confluentus">
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<emphasis id="2F9FEAC9B71EFFA66C32FD1E46E958AF" box="[1282,1408,712,738]" italics="true" pageId="17" pageNumber="16">confluentus</emphasis>
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</taxonomicName>
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<specimenCount id="0BEDFD52B71EFFA66CB5FD1A46F358AF" box="[1413,1434,716,738]" pageId="17" pageNumber="16" type="male">♂</specimenCount>
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; Otto, Muona & Córdoba-Alfaro’ Det.
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<collectionCode id="7BFAAE1EB71EFFA66B51FD3D411C5948" box="[609,629,747,773]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="16">R</collectionCode>
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.
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<collectorName id="B01E530DB71EFFA66B49FD3D41A35949" box="[633,714,746,773]" pageId="17" pageNumber="16">L. Otto</collectorName>
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; 2019” (red printed label).
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|
|
<emphasis id="2F9FEAC9B71EFFA66AC0FD3C47CE5949" bold="true" box="[1008,1191,746,772]" pageId="17" pageNumber="16">
|
|||
|
|
Female
|
|||
|
|
<typeStatus id="C2508879B71EFFA66D79FD3C47CE5949" box="[1097,1191,746,772]" pageId="17" pageNumber="16" type="allotype">allotype</typeStatus>
|
|||
|
|
</emphasis>
|
|||
|
|
: “
|
|||
|
|
<collectingCountry id="65FC764BB71EFFA66D8DFD3C46365949" box="[1213,1375,746,772]" name="Costa Rica" pageId="17" pageNumber="16">COSTA RICA</collectingCountry>
|
|||
|
|
: Prov.
|
|||
|
|
<collectingRegion id="DF2FF839B71EFFA6698FFCDA4277596B" box="[191,286,780,806]" country="Costa Rica" name="Heredia" pageId="17" pageNumber="16">Heredia</collectingRegion>
|
|||
|
|
:;
|
|||
|
|
<locationDeviation id="0CE62FF9B71EFFA6681CFCDA4172596B" box="[300,539,780,806]" location="La Virgen" pageId="17" pageNumber="16">
|
|||
|
|
<quantity id="DA139B3EB71EFFA6681CFCDA421A596B" box="[300,371,780,806]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="17" pageNumber="16" unit="km" value="16.0">16 km</quantity>
|
|||
|
|
SSE La Virgen
|
|||
|
|
</locationDeviation>
|
|||
|
|
, 1050–;
|
|||
|
|
<quantity id="DA139B3EB71EFFA66B47FCDA41A4596B" box="[631,717,780,807]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="17" pageNumber="16" unit="m" value="1150.0">
|
|||
|
|
<elevation id="96C6D1E8B71EFFA66B47FCDA41A4596B" box="[631,717,780,807]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="17" pageNumber="16" unit="m" value="1150.0">1150 m</elevation>
|
|||
|
|
</quantity>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="78DF501CB71EFFA66BE8FCDA405A596B" box="[728,819,780,806]" degrees="10" direction="north" minutes="16" orientation="latitude" pageId="17" pageNumber="16" precision="925" value="10.266666">10°16′N</geoCoordinate>
|
|||
|
|
<geoCoordinate id="78DF501CB71EFFA66A0AFCDA40F3596B" box="[826,922,780,807]" degrees="84" direction="west" minutes="05" orientation="longitude" pageId="17" pageNumber="16" precision="925" value="-84.083336">84°05′W</geoCoordinate>
|
|||
|
|
;
|
|||
|
|
<date id="6955101BB71EFFA66A95FCDA475B596B" box="[933,1074,780,806]" pageId="17" pageNumber="16" value="2001-04-09">
|
|||
|
|
<collectingDate id="7911E9F3B71EFFA66A95FCDA475B596B" box="[933,1074,780,806]" pageId="17" pageNumber="16" value="2001-04-09">9 Abril 2001</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
; INBio-OET-ALAS transect” (“9 Abril”, “1” were handwritten on label) / “
|
|||
|
|
<date id="6955101BB71EFFA66B4FFCF840715905" box="[639,792,814,840]" pageId="17" pageNumber="16" value="2001-04-09">
|
|||
|
|
<collectingDate id="7911E9F3B71EFFA66B4FFCF840715905" box="[639,792,814,840]" pageId="17" pageNumber="16" value="2001-04-09">09 Abril 2001</collectingDate>
|
|||
|
|
</date>
|
|||
|
|
; 11/M/01/061; Transect” / “
|
|||
|
|
<emphasis id="2F9FEAC9B71EFFA66D7BFCFB47B7590A" bold="true" box="[1099,1246,813,839]" pageId="17" pageNumber="16">
|
|||
|
|
<typeStatus id="C2508879B71EFFA66D7BFCFB47B2590A" box="[1099,1243,813,839]" pageId="17" pageNumber="16" type="allotype">ALLOTYPE</typeStatus>
|
|||
|
|
:
|
|||
|
|
</emphasis>
|
|||
|
|
;
|
|||
|
|
<taxonomicName id="DAEB4D58B71EFFA66DDBFCF8460D5905" authorityName="Newman" authorityYear="1838" box="[1259,1380,814,840]" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="2F9FEAC9B71EFFA66DDBFCF8460D5905" box="[1259,1380,814,840]" italics="true" pageId="17" pageNumber="16">Onichodon</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
;
|
|||
|
|
<taxonomicName id="DAEB4D58B71EFFA66C40FCF8427E5924" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="confluentus">
|
|||
|
|
<emphasis id="2F9FEAC9B71EFFA66C40FCF8427E5924" italics="true" pageId="17" pageNumber="16">confluentus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<specimenCount id="0BEDFD52B71EFFA6682FFC85425D5924" box="[287,308,851,873]" pageId="17" pageNumber="16" type="female">♀</specimenCount>
|
|||
|
|
; Otto, Muona & Córdoba-Alfaro; Det.
|
|||
|
|
<collectionCode id="7BFAAE1EB71EFFA66BDBFC8641965927" box="[747,767,848,874]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="16">R</collectionCode>
|
|||
|
|
.
|
|||
|
|
<collectorName id="B01E530DB71EFFA66A33FC86403F5924" box="[771,854,847,874]" pageId="17" pageNumber="16">L. Otto</collectorName>
|
|||
|
|
; 2019” (yellow printed label).
|
|||
|
|
<typeStatus id="C2508879B71EFFA66D97FC9946665924" box="[1191,1295,847,873]" pageId="17" pageNumber="16" type="holotype">Holotype</typeStatus>
|
|||
|
|
and
|
|||
|
|
<typeStatus id="C2508879B71EFFA66C77FC9946C95924" box="[1351,1440,847,873]" pageId="17" pageNumber="16" type="allotype">allotype</typeStatus>
|
|||
|
|
are to be deposited in
|
|||
|
|
<collectionCode id="7BFAAE1EB71EFFA66887FCA7429559C6" box="[439,508,881,907]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="17" pageNumber="16">INBC</collectionCode>
|
|||
|
|
</materialsCitation>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="1D5436DBB71EFFA66830FC4546EE5C60" blockId="17.[191,1441,679,1582]" pageId="17" pageNumber="16">
|
|||
|
|
<emphasis id="2F9FEAC9B71EFFA66830FC45421359E0" bold="true" box="[256,378,915,941]" pageId="17" pageNumber="16">
|
|||
|
|
<typeStatus id="C2508879B71EFFA66830FC45421C59E0" box="[256,373,915,941]" pageId="17" pageNumber="16" type="paratype">Paratypes</typeStatus>
|
|||
|
|
.
|
|||
|
|
</emphasis>
|
|||
|
|
<specimenCount id="0BEDFD52B71EFFA668B3FC4542AD59E1" box="[387,452,915,941]" pageId="17" pageNumber="16" type="male">7 ♂♂</specimenCount>
|
|||
|
|
:
|
|||
|
|
<emphasis id="2F9FEAC9B71EFFA668E4FC4441EB59E1" bold="true" box="[468,642,914,940]" pageId="17" pageNumber="16">
|
|||
|
|
<collectingCountry id="65FC764BB71EFFA668E4FC44411459E1" box="[468,637,914,940]" name="Costa Rica" pageId="17" pageNumber="16">COSTA RICA</collectingCountry>
|
|||
|
|
:
|
|||
|
|
</emphasis>
|
|||
|
|
<specimenCount id="0BEDFD52B71EFFA66BBBFC4541DE59E1" box="[651,695,915,941]" pageId="17" pageNumber="16" type="male">1 ♂</specimenCount>
|
|||
|
|
, “COSTA RICA:
|
|||
|
|
<collectingRegion id="DF2FF839B71EFFA66AB3FC4540B659E0" box="[899,991,915,941]" country="Costa Rica" name="Heredia" pageId="17" pageNumber="16">Heredia</collectingRegion>
|
|||
|
|
; Est.Bio. La Selva, 50–;
|
|||
|
|
<quantity id="DA139B3EB71EFFA66DC3FC45465559E0" box="[1267,1340,915,942]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="16" unit="m" value="150.0">150 m</quantity>
|
|||
|
|
<geoCoordinate id="78DF501CB71EFFA66C76FC4546C959E0" box="[1350,1440,915,941]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="17" pageNumber="16" precision="925" value="10.433333">10°26′N</geoCoordinate>
|
|||
|
|
<geoCoordinate id="78DF501CB71EFFA669F0FC6342495982" box="[192,288,949,975]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="17" pageNumber="16" precision="925" value="-84.01667">84°01′W</geoCoordinate>
|
|||
|
|
; Proy.
|
|||
|
|
<collectionCode id="7BFAAE1EB71EFFA66853FC6342C35982" box="[355,426,949,975]" pageId="17" pageNumber="16">ALAS</collectionCode>
|
|||
|
|
, INBio-OET” / “M/14/061;
|
|||
|
|
<date id="6955101BB71EFFA66BEFFC6340115982" box="[735,888,948,975]" pageId="17" pageNumber="16" value="1993-04-02">02 Abril 1993</date>
|
|||
|
|
; Bosque secundario” / “
|
|||
|
|
<emphasis id="2F9FEAC9B71EFFA66DB1FC62467A5983" bold="true" box="[1153,1299,948,974]" pageId="17" pageNumber="16">
|
|||
|
|
<typeStatus id="C2508879B71EFFA66DB1FC6246665983" box="[1153,1295,948,974]" pageId="17" pageNumber="16" type="paratype">PARATYPE</typeStatus>
|
|||
|
|
:
|
|||
|
|
</emphasis>
|
|||
|
|
;
|
|||
|
|
<taxonomicName id="DAEB4D58B71EFFA66C10FC6246F05983" authorityName="Newman" authorityYear="1838" box="[1312,1433,948,974]" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="2F9FEAC9B71EFFA66C10FC6246F05983" box="[1312,1433,948,974]" italics="true" pageId="17" pageNumber="16">Onichodon</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
;
|
|||
|
|
<taxonomicName id="DAEB4D58B71EFFA669F0FC00425759BD" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[192,318,982,1008]" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="confluentus">
|
|||
|
|
<emphasis id="2F9FEAC9B71EFFA669F0FC00425759BD" box="[192,318,982,1008]" italics="true" pageId="17" pageNumber="16">confluentus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<specimenCount id="0BEDFD52B71EFFA66872FC0C423E59BD" box="[322,343,986,1008]" pageId="17" pageNumber="16" type="male">♂</specimenCount>
|
|||
|
|
; Otto, Muona & Córdoba-Alfaro; Det.
|
|||
|
|
<collectionCode id="7BFAAE1EB71EFFA66A33FC01407E59BC" box="[771,791,983,1009]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="16">R</collectionCode>
|
|||
|
|
.L. Otto; 2019” (yellow printed label) (
|
|||
|
|
<collectionCode id="7BFAAE1EB71EFFA66D83FC00479259BD" box="[1203,1275,982,1008]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="17" pageNumber="16">INBC</collectionCode>
|
|||
|
|
);
|
|||
|
|
<specimenCount id="0BEDFD52B71EFFA66C3CFC00465B59BD" box="[1292,1330,982,1008]" pageId="17" pageNumber="16" type="male">1 ♂</specimenCount>
|
|||
|
|
, “COSTA RICA:
|
|||
|
|
<collectingRegion id="DF2FF839B71EFFA6683CFC2E42015E5F" box="[268,360,1016,1042]" country="Costa Rica" name="Heredia" pageId="17" pageNumber="16">Heredia</collectingRegion>
|
|||
|
|
; Est.Bio. La Selva, 50–;
|
|||
|
|
<quantity id="DA139B3EB71EFFA66B5EFC2E41DD5E5F" box="[622,692,1016,1043]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="16" unit="m" value="150.0">150 m</quantity>
|
|||
|
|
<geoCoordinate id="78DF501CB71EFFA66B8BFC2E407E5E5F" box="[699,791,1016,1042]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="17" pageNumber="16" precision="925" value="10.433333">10°26′N</geoCoordinate>
|
|||
|
|
<geoCoordinate id="78DF501CB71EFFA66A2DFC2E40175E5F" box="[797,894,1016,1042]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="17" pageNumber="16" precision="925" value="-84.01667">84°01′W</geoCoordinate>
|
|||
|
|
; Proy.
|
|||
|
|
<collectionCode id="7BFAAE1EB71EFFA66AF1FC2E47615E5F" box="[961,1032,1016,1042]" pageId="17" pageNumber="16">ALAS</collectionCode>
|
|||
|
|
, INBio-OET” / “M/15/094;
|
|||
|
|
<date id="6955101BB71EFFA66C0FFC2E43915E79" pageId="17" pageNumber="16" value="1993-05-03">03 Mayo 1993</date>
|
|||
|
|
; Bosque secundario” / “
|
|||
|
|
<emphasis id="2F9FEAC9B71EFFA66B31FBCF41FA5E7E" bold="true" box="[513,659,1049,1075]" pageId="17" pageNumber="16">
|
|||
|
|
<typeStatus id="C2508879B71EFFA66B31FBCF41E65E7E" box="[513,655,1049,1075]" pageId="17" pageNumber="16" type="paratype">PARATYPE</typeStatus>
|
|||
|
|
:
|
|||
|
|
</emphasis>
|
|||
|
|
;
|
|||
|
|
<taxonomicName id="DAEB4D58B71EFFA66B91FBCF40735E7E" authorityName="Newman" authorityYear="1838" box="[673,794,1049,1075]" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="2F9FEAC9B71EFFA66B91FBCF40735E7E" box="[673,794,1049,1075]" italics="true" pageId="17" pageNumber="16">Onichodon</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
;
|
|||
|
|
<taxonomicName id="DAEB4D58B71EFFA66A16FBCF40CD5E7E" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[806,932,1049,1075]" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="confluentus">
|
|||
|
|
<emphasis id="2F9FEAC9B71EFFA66A16FBCF40CD5E7E" box="[806,932,1049,1075]" italics="true" pageId="17" pageNumber="16">confluentus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<specimenCount id="0BEDFD52B71EFFA66A9AFBCB40D65E7E" box="[938,959,1053,1075]" pageId="17" pageNumber="16" type="male">♂</specimenCount>
|
|||
|
|
; Otto, Muona & Córdoba-Alfaro; Det.
|
|||
|
|
<collectionCode id="7BFAAE1EB71EFFA66C42FBCC46EF5E79" box="[1394,1414,1050,1076]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="16">R</collectionCode>
|
|||
|
|
.L. Otto; 2019” (yellow printed label) (
|
|||
|
|
<collectionCode id="7BFAAE1EB71EFFA66B7AFBED41FB5E18" box="[586,658,1083,1109]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="17" pageNumber="16">INBC</collectionCode>
|
|||
|
|
);
|
|||
|
|
<specimenCount id="0BEDFD52B71EFFA66B95FBED41A65E18" box="[677,719,1083,1109]" pageId="17" pageNumber="16" type="male">1 ♂</specimenCount>
|
|||
|
|
, “COSTA RICA:
|
|||
|
|
<collectingRegion id="DF2FF839B71EFFA66AA3FBED40865E18" box="[915,1007,1083,1109]" country="Costa Rica" name="Heredia" pageId="17" pageNumber="16">Heredia</collectingRegion>
|
|||
|
|
; Est.Bio. La Selva, 50–;
|
|||
|
|
<quantity id="DA139B3EB71EFFA66DC8FBED46565E1B" box="[1272,1343,1083,1110]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="16" unit="m" value="150.0">150 m</quantity>
|
|||
|
|
<geoCoordinate id="78DF501CB71EFFA66C76FBED46C95E1B" box="[1350,1440,1083,1110]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="17" pageNumber="16" precision="925" value="10.433333">10°26′N</geoCoordinate>
|
|||
|
|
<geoCoordinate id="78DF501CB71EFFA669F0FB8B42495E3A" box="[192,288,1117,1143]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="17" pageNumber="16" precision="925" value="-84.01667">84°01′W</geoCoordinate>
|
|||
|
|
; Proy.
|
|||
|
|
<collectionCode id="7BFAAE1EB71EFFA66853FB8B42C35E3A" box="[355,426,1117,1143]" pageId="17" pageNumber="16">ALAS</collectionCode>
|
|||
|
|
, INBio-OET” / “M/16/122;
|
|||
|
|
<date id="6955101BB71EFFA66BD3FB8B40E95E3A" box="[739,896,1117,1143]" pageId="17" pageNumber="16" value="1993-06-02">02 Junio 1993</date>
|
|||
|
|
; Bosque secundario” / “Collection of the Global; Eucnemid Research Project; (Robert L. Otto)” (green framed white label) / “
|
|||
|
|
<emphasis id="2F9FEAC9B71EFFA66D23FBA847CF5ED5" bold="true" box="[1043,1190,1150,1176]" pageId="17" pageNumber="16">
|
|||
|
|
<typeStatus id="C2508879B71EFFA66D23FBA847C85ED5" box="[1043,1185,1150,1176]" pageId="17" pageNumber="16" type="paratype">PARATYPE</typeStatus>
|
|||
|
|
:
|
|||
|
|
</emphasis>
|
|||
|
|
;
|
|||
|
|
<taxonomicName id="DAEB4D58B71EFFA66D83FBA946455ED4" authorityName="Newman" authorityYear="1838" box="[1203,1324,1151,1177]" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="2F9FEAC9B71EFFA66D83FBA946455ED4" box="[1203,1324,1151,1177]" italics="true" pageId="17" pageNumber="16">Onichodon</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
;
|
|||
|
|
<taxonomicName id="DAEB4D58B71EFFA66C09FBA943885EF7" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="confluentus">
|
|||
|
|
<emphasis id="2F9FEAC9B71EFFA66C09FBA943885EF7" italics="true" pageId="17" pageNumber="16">confluentus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<specimenCount id="0BEDFD52B71EFFA669C5FB7242635EF7" box="[245,266,1188,1210]" pageId="17" pageNumber="16" type="male">♂</specimenCount>
|
|||
|
|
; Otto, Muona & Córdoba-Alfaro; Det.
|
|||
|
|
<collectionCode id="7BFAAE1EB71EFFA66BE3FB77418E5EF6" box="[723,743,1185,1211]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="16">R</collectionCode>
|
|||
|
|
.L. Otto; 2019” (yellow printed label) (
|
|||
|
|
<collectionCode id="7BFAAE1EB71EFFA66D91FB7647855EF7" box="[1185,1260,1184,1210]" pageId="17" pageNumber="16">GERP</collectionCode>
|
|||
|
|
);
|
|||
|
|
<specimenCount id="0BEDFD52B71EFFA66C32FB7646475EF7" box="[1282,1326,1184,1210]" pageId="17" pageNumber="16" type="male">1 ♂</specimenCount>
|
|||
|
|
, “COSTA RICA:
|
|||
|
|
<collectingRegion id="DF2FF839B71EFFA6683EFB1442035E91" box="[270,362,1218,1244]" country="Costa Rica" name="Heredia" pageId="17" pageNumber="16">Heredia</collectingRegion>
|
|||
|
|
; Est.Bio. La Selva, 50–;
|
|||
|
|
<quantity id="DA139B3EB71EFFA66B44FB1441D25E90" box="[628,699,1218,1245]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="16" unit="m" value="150.0">150 m</quantity>
|
|||
|
|
<geoCoordinate id="78DF501CB71EFFA66BF2FB1440745E90" box="[706,797,1218,1245]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="17" pageNumber="16" precision="925" value="10.433333">10°26′N</geoCoordinate>
|
|||
|
|
<geoCoordinate id="78DF501CB71EFFA66A14FB1440EC5E90" box="[804,901,1218,1245]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="17" pageNumber="16" precision="925" value="-84.01667">84°01′W</geoCoordinate>
|
|||
|
|
; Proy.
|
|||
|
|
<collectionCode id="7BFAAE1EB71EFFA66AF8FB1447665E91" box="[968,1039,1218,1244]" pageId="17" pageNumber="16">ALAS</collectionCode>
|
|||
|
|
, INBio-OET” / “M/14/148;
|
|||
|
|
<date id="6955101BB71EFFA66C79FB1443905EB3" pageId="17" pageNumber="16" value="1993-07-01">01 Julio 1993</date>
|
|||
|
|
; Bosque secundario” / “
|
|||
|
|
<emphasis id="2F9FEAC9B71EFFA66B31FB3541FA5EB0" bold="true" box="[513,659,1251,1277]" pageId="17" pageNumber="16">
|
|||
|
|
<typeStatus id="C2508879B71EFFA66B31FB3541E65EB0" box="[513,655,1251,1277]" pageId="17" pageNumber="16" type="paratype">PARATYPE</typeStatus>
|
|||
|
|
:
|
|||
|
|
</emphasis>
|
|||
|
|
;
|
|||
|
|
<taxonomicName id="DAEB4D58B71EFFA66B91FB3240735EB3" authorityName="Newman" authorityYear="1838" box="[673,794,1252,1278]" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="2F9FEAC9B71EFFA66B91FB3240735EB3" box="[673,794,1252,1278]" italics="true" pageId="17" pageNumber="16">Onichodon</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
;
|
|||
|
|
<taxonomicName id="DAEB4D58B71EFFA66A16FB3240CD5EB3" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[806,932,1252,1278]" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="confluentus">
|
|||
|
|
<emphasis id="2F9FEAC9B71EFFA66A16FB3240CD5EB3" box="[806,932,1252,1278]" italics="true" pageId="17" pageNumber="16">confluentus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<specimenCount id="0BEDFD52B71EFFA66A9AFB3140D65EB0" box="[938,959,1255,1277]" pageId="17" pageNumber="16" type="male">♂</specimenCount>
|
|||
|
|
; Otto, Muona & Córdoba-Alfaro; Det.
|
|||
|
|
<collectionCode id="7BFAAE1EB71EFFA66C42FB3246EF5EB3" box="[1394,1414,1252,1278]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="16">R</collectionCode>
|
|||
|
|
.L. Otto; 2019” (yellow printed label) (
|
|||
|
|
<collectionCode id="7BFAAE1EB71EFFA66B7AFAD041FB5F6D" box="[586,658,1286,1312]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="17" pageNumber="16">INBC</collectionCode>
|
|||
|
|
);
|
|||
|
|
<specimenCount id="0BEDFD52B71EFFA66B95FAD041A65F52" box="[677,719,1286,1312]" pageId="17" pageNumber="16" type="male">1 ♂</specimenCount>
|
|||
|
|
, “COSTA RICA:
|
|||
|
|
<collectingRegion id="DF2FF839B71EFFA66AA3FAD340865F52" box="[915,1007,1285,1311]" country="Costa Rica" name="Heredia" pageId="17" pageNumber="16">Heredia</collectingRegion>
|
|||
|
|
; Est.Bio. La Selva, 50–;
|
|||
|
|
<quantity id="DA139B3EB71EFFA66DC8FAD346565F6D" box="[1272,1343,1285,1312]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="16" unit="m" value="150.0">150 m</quantity>
|
|||
|
|
<geoCoordinate id="78DF501CB71EFFA66C76FAD346C95F6D" box="[1350,1440,1285,1312]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="17" pageNumber="16" precision="925" value="10.433333">10°26′N</geoCoordinate>
|
|||
|
|
<geoCoordinate id="78DF501CB71EFFA669F0FAF142495F0F" box="[192,288,1319,1346]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="17" pageNumber="16" precision="925" value="-84.01667">84°01′W</geoCoordinate>
|
|||
|
|
; Proy.
|
|||
|
|
<collectionCode id="7BFAAE1EB71EFFA66855FAF142C55F0C" box="[357,428,1319,1345]" pageId="17" pageNumber="16">ALAS</collectionCode>
|
|||
|
|
, INBio-OET” / “M/12/397;
|
|||
|
|
<date id="6955101BB71EFFA66BD8FAF140EE5F0F" box="[744,903,1319,1346]" pageId="17" pageNumber="16" value="1995-06-30">30 Junio 1995</date>
|
|||
|
|
; Bosque primario” / “
|
|||
|
|
<emphasis id="2F9FEAC9B71EFFA66D4FFAF146785F0C" bold="true" box="[1151,1297,1319,1345]" pageId="17" pageNumber="16">
|
|||
|
|
<typeStatus id="C2508879B71EFFA66D4FFAF146645F0C" box="[1151,1293,1319,1345]" pageId="17" pageNumber="16" type="paratype">PARATYPE</typeStatus>
|
|||
|
|
:
|
|||
|
|
</emphasis>
|
|||
|
|
;
|
|||
|
|
<taxonomicName id="DAEB4D58B71EFFA66C10FAF146F05F0C" authorityName="Newman" authorityYear="1838" box="[1312,1433,1319,1345]" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="2F9FEAC9B71EFFA66C10FAF146F05F0C" box="[1312,1433,1319,1345]" italics="true" pageId="17" pageNumber="16">Onichodon</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
;
|
|||
|
|
<taxonomicName id="DAEB4D58B71EFFA669F0FA9F42575F2E" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[192,318,1353,1379]" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="confluentus">
|
|||
|
|
<emphasis id="2F9FEAC9B71EFFA669F0FA9F42575F2E" box="[192,318,1353,1379]" italics="true" pageId="17" pageNumber="16">confluentus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<specimenCount id="0BEDFD52B71EFFA66872FA9A423E5F2F" box="[322,343,1356,1378]" pageId="17" pageNumber="16" type="male">♂</specimenCount>
|
|||
|
|
; Otto, Muona & Córdoba-Alfaro; Det.
|
|||
|
|
<collectionCode id="7BFAAE1EB71EFFA66A33FA9F407E5F2E" box="[771,791,1353,1379]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="16">R</collectionCode>
|
|||
|
|
.L. Otto; 2019” (yellow printed label) (
|
|||
|
|
<collectionCode id="7BFAAE1EB71EFFA66D83FA9F47925F2E" box="[1203,1275,1353,1379]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="17" pageNumber="16">INBC</collectionCode>
|
|||
|
|
);
|
|||
|
|
<specimenCount id="0BEDFD52B71EFFA66C3CFA9F465B5F2F" box="[1292,1330,1353,1379]" pageId="17" pageNumber="16" type="male">1 ♂</specimenCount>
|
|||
|
|
, “COSTA RICA:
|
|||
|
|
<collectingRegion id="DF2FF839B71EFFA6683CFABC42015FC9" box="[268,360,1386,1412]" country="Costa Rica" name="Heredia" pageId="17" pageNumber="16">Heredia</collectingRegion>
|
|||
|
|
; Est.Bio. La Selva, 50–;
|
|||
|
|
<quantity id="DA139B3EB71EFFA66B5EFABC41DD5FC8" box="[622,692,1386,1413]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="16" unit="m" value="150.0">150 m</quantity>
|
|||
|
|
<geoCoordinate id="78DF501CB71EFFA66B8BFABC407E5FC8" box="[699,791,1386,1413]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="17" pageNumber="16" precision="925" value="10.433333">10°26′N</geoCoordinate>
|
|||
|
|
<geoCoordinate id="78DF501CB71EFFA66A2DFABD40175FC8" box="[797,894,1387,1413]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="17" pageNumber="16" precision="925" value="-84.01667">84°01′W</geoCoordinate>
|
|||
|
|
; Proy.
|
|||
|
|
<collectionCode id="7BFAAE1EB71EFFA66AF1FABD47615FC8" box="[961,1032,1387,1413]" pageId="17" pageNumber="16">ALAS</collectionCode>
|
|||
|
|
, INBio-OET” / “M/05/655;
|
|||
|
|
<date id="6955101BB71EFFA66C0FFABD43915FEB" pageId="17" pageNumber="16" value="1996-05-31">31 Mayo 1996</date>
|
|||
|
|
; Bosque primario” / “
|
|||
|
|
<emphasis id="2F9FEAC9B71EFFA668C0FA5A41EA5FEB" bold="true" box="[496,643,1420,1446]" pageId="17" pageNumber="16">
|
|||
|
|
<typeStatus id="C2508879B71EFFA668C0FA5A41175FEB" box="[496,638,1420,1446]" pageId="17" pageNumber="16" type="paratype">PARATYPE</typeStatus>
|
|||
|
|
:
|
|||
|
|
</emphasis>
|
|||
|
|
;
|
|||
|
|
<taxonomicName id="DAEB4D58B71EFFA66BA2FA5A40625FEB" authorityName="Newman" authorityYear="1838" box="[658,779,1420,1446]" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="2F9FEAC9B71EFFA66BA2FA5A40625FEB" box="[658,779,1420,1446]" italics="true" pageId="17" pageNumber="16">Onichodon</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
;
|
|||
|
|
<taxonomicName id="DAEB4D58B71EFFA66A29FA5A40FE5FEB" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[793,919,1420,1446]" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="confluentus">
|
|||
|
|
<emphasis id="2F9FEAC9B71EFFA66A29FA5A40FE5FEB" box="[793,919,1420,1446]" italics="true" pageId="17" pageNumber="16">confluentus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<specimenCount id="0BEDFD52B71EFFA66AAFFA4640DD5FEB" box="[927,948,1424,1446]" pageId="17" pageNumber="16" type="male">♂</specimenCount>
|
|||
|
|
; Otto, Muona & Córdoba-Alfaro; Det.
|
|||
|
|
<collectionCode id="7BFAAE1EB71EFFA66C42FA5B46EF5FEA" box="[1394,1414,1421,1447]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="16">R</collectionCode>
|
|||
|
|
.L. Otto; 2019” (yellow printed label) (
|
|||
|
|
<collectionCode id="7BFAAE1EB71EFFA66B79FA7841F85F85" box="[585,657,1454,1480]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="17" pageNumber="16">INBC</collectionCode>
|
|||
|
|
);
|
|||
|
|
<specimenCount id="0BEDFD52B71EFFA66B94FA7841A55F8A" box="[676,716,1454,1480]" pageId="17" pageNumber="16" type="male">1 ♂</specimenCount>
|
|||
|
|
, “
|
|||
|
|
<date id="6955101BB71EFFA66BD3FA78401A5F85" box="[739,883,1454,1480]" pageId="17" pageNumber="16" value="2001-03-21">21 Mar 2001</date>
|
|||
|
|
; INBio-OET-ALAS transect; COSTA RICA: Prov.
|
|||
|
|
<collectingRegion id="DF2FF839B71EFFA669F0FA0642765FA7" box="[192,287,1488,1514]" country="Costa Rica" name="Heredia" pageId="17" pageNumber="16">Heredia</collectingRegion>
|
|||
|
|
:;
|
|||
|
|
<quantity id="DA139B3EB71EFFA6681DFA06421D5FA7" box="[301,372,1488,1514]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="17" pageNumber="16" unit="km" value="16.0">16 km</quantity>
|
|||
|
|
SSE La Virgen, 1050–;
|
|||
|
|
<quantity id="DA139B3EB71EFFA66B48FA1941A75FA7" box="[632,718,1487,1514]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="17" pageNumber="16" unit="m" value="1150.0">1150 m</quantity>
|
|||
|
|
,
|
|||
|
|
<geoCoordinate id="78DF501CB71EFFA66BE9FA06405B5FA7" box="[729,818,1488,1514]" degrees="10" direction="north" minutes="16" orientation="latitude" pageId="17" pageNumber="16" precision="925" value="10.266666">10°16′N</geoCoordinate>
|
|||
|
|
<geoCoordinate id="78DF501CB71EFFA66A09FA1940F05FA7" box="[825,921,1487,1514]" degrees="84" direction="west" minutes="05" orientation="longitude" pageId="17" pageNumber="16" precision="925" value="-84.083336">84°05′W</geoCoordinate>
|
|||
|
|
(“21 Mar” and “1” were handwritten on label) /
|
|||
|
|
<date id="6955101BB71EFFA669E1FA2742135C46" box="[209,378,1521,1548]" pageId="17" pageNumber="16" value="2001-03-21">21 Marzo 2001</date>
|
|||
|
|
; Transect; 11/M/07/067” / “
|
|||
|
|
<emphasis id="2F9FEAC9B71EFFA66B83FA27402C5C46" bold="true" box="[691,837,1521,1547]" pageId="17" pageNumber="16">
|
|||
|
|
<typeStatus id="C2508879B71EFFA66B83FA2740285C46" box="[691,833,1521,1547]" pageId="17" pageNumber="16" type="paratype">PARATYPE</typeStatus>
|
|||
|
|
:
|
|||
|
|
</emphasis>
|
|||
|
|
;
|
|||
|
|
<taxonomicName id="DAEB4D58B71EFFA66A63FA2740A55C46" authorityName="Newman" authorityYear="1838" box="[851,972,1521,1547]" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="2F9FEAC9B71EFFA66A63FA2740A55C46" box="[851,972,1521,1547]" italics="true" pageId="17" pageNumber="16">Onichodon</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
;
|
|||
|
|
<taxonomicName id="DAEB4D58B71EFFA66AEAFA2747315C46" authorityName="Otto, Muona and Cordoba-Alfaro" authorityYear="2023" box="[986,1112,1521,1547]" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="confluentus">
|
|||
|
|
<emphasis id="2F9FEAC9B71EFFA66AEAFA2747315C46" box="[986,1112,1521,1547]" italics="true" pageId="17" pageNumber="16">confluentus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<specimenCount id="0BEDFD52B71EFFA66D6EFA23471A5C46" box="[1118,1139,1525,1547]" pageId="17" pageNumber="16" type="male">♂</specimenCount>
|
|||
|
|
; Otto, Muona & Córdoba-Alfaro; Det.
|
|||
|
|
<collectionCode id="7BFAAE1EB71EFFA6687AF9C242375C63" box="[330,350,1556,1582]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="17" pageNumber="16">R</collectionCode>
|
|||
|
|
.L. Otto; 2019” (yellow printed label) (
|
|||
|
|
<collectionCode id="7BFAAE1EB71EFFA66A33F9C540225C60" box="[771,843,1555,1581]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="17" pageNumber="16">INBC</collectionCode>
|
|||
|
|
).
|
|||
|
|
<typeStatus id="C2508879B71EFFA66A6DF9C240A05C63" box="[861,969,1556,1582]" pageId="17" pageNumber="16" type="paratype">Paratypes</typeStatus>
|
|||
|
|
are to be deposited in
|
|||
|
|
<collectionCode id="7BFAAE1EB71EFFA66DF5F9C5466E5C60" box="[1221,1287,1555,1581]" country="Costa Rica" httpUri="http://grbio.org/cool/y350-djvc" name="Instituto Nacional de Biodiversidad (INBio)" pageId="17" pageNumber="16">INBC</collectionCode>
|
|||
|
|
and
|
|||
|
|
<collectionCode id="7BFAAE1EB71EFFA66C0EF9C546EA5C60" box="[1342,1411,1555,1581]" pageId="17" pageNumber="16">GERP</collectionCode>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="55F16550B71EFFA569F0F9EA41D25EB4" lastPageId="18" lastPageNumber="17" pageId="17" pageNumber="16" type="description">
|
|||
|
|
<paragraph id="1D5436DBB71EFFA569F0F9EA428B5E4B" blockId="17.[192,1443,1596,1927]" lastBlockId="18.[191,1442,802,1131]" lastPageId="18" lastPageNumber="17" pageId="17" pageNumber="16">
|
|||
|
|
<emphasis id="2F9FEAC9B71EFFA669F0F9EA416D5C1A" bold="true" box="[192,516,1596,1623]" pageId="17" pageNumber="16">
|
|||
|
|
Description. Male
|
|||
|
|
<typeStatus id="C2508879B71EFFA668ADF9EB416D5C1A" box="[413,516,1597,1623]" pageId="17" pageNumber="16" type="holotype">holotype</typeStatus>
|
|||
|
|
</emphasis>
|
|||
|
|
: Length, 8.0 mm. Width,
|
|||
|
|
<quantity id="DA139B3EB71EFFA66A00F9EB40E55C1A" box="[816,908,1597,1624]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="17" pageNumber="16" unit="mm" value="2.5">2.5 mm</quantity>
|
|||
|
|
. Body elongate; dark reddish-brown; antennae and legs dark reddish-brown; head, pronotum and elytra clothed with short, recumbent yellowish setae (
|
|||
|
|
<figureCitation id="85D02A5EB71EFFA66C65F98843895CD7" captionStart="Figures 27–30" captionStartId="16.[192,279,631,657]" captionTargetBox="[378,1254,213,601]" captionTargetId="figure-67@16.[378,1254,213,601]" captionTargetPageId="16" captionText="Figures 27–30. Costa Rican macraulacine eucnemids. 27) Maelodrus costaricensis sp. nov., dorsal habitus. 28) Maelodrus costaricensis sp. nov., lateral habitus. 29) Maelodrus costaricensis sp. nov., ventral habitus. 30) Onichodon confluentus sp. nov., male holotype, dorsal habitus. (Scale: 27–29 = 1.0 mm; 30 = 5.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.10831097" httpUri="https://zenodo.org/record/10831097/files/figure.png" pageId="17" pageNumber="16">Figure 30</figureCitation>
|
|||
|
|
).
|
|||
|
|
<emphasis id="2F9FEAC9B71EFFA669C2F95642595CD7" bold="true" box="[242,304,1664,1690]" pageId="17" pageNumber="16">Head</emphasis>
|
|||
|
|
: Surface dullish, subspherical, confluently rugose; frons convex, simple without impression or carina; apical margin of frontoclypeal region weakly trilobed, about 2 times wider than base; mandibles stout, bidentate, densely punctate.
|
|||
|
|
<emphasis id="2F9FEAC9B71EFFA668BEF915429A5C90" bold="true" box="[398,499,1731,1757]" pageId="17" pageNumber="16">Antenna</emphasis>
|
|||
|
|
: Filiform from flagellomeres I–IX, attaining at least 1/3 the length of the body, just beyond pronotal hind angles; flagellomere I longer than wide, slightly longer than II; flagellomeres II–VIII each longer than wide, sub-equal; flagellomere IX slightly longer than VIII.
|
|||
|
|
<emphasis id="2F9FEAC9B71EFFA66AE0F8D147215D6C" bold="true" box="[976,1096,1799,1825]" pageId="17" pageNumber="16">Pronotum</emphasis>
|
|||
|
|
: Surface somewhat shiny, with closely spaced punctures; quadrate, with well-developed, sharp hind angles; sides subparallel-sided at basal 1/2, arcuate at apical 1/2, slightly sinuate above pronotal hind angles; disc convex with shallow fovea above scutellar shield; base sinuous, with circular pit on each side above scutellar shield.
|
|||
|
|
<emphasis id="2F9FEAC9B71EFFA66D3BF8BA47A85DCB" bold="true" box="[1035,1217,1900,1926]" pageId="17" pageNumber="16">Scutellar shield</emphasis>
|
|||
|
|
: Longer than wide, sub-triangular, shallowly punctate, setose and distally rounded.
|
|||
|
|
<emphasis id="2F9FEAC9B71DFFA56ABAFCF440B85971" bold="true" box="[906,977,802,828]" pageId="18" pageNumber="17">Elytra</emphasis>
|
|||
|
|
: Very shallow striae indicated; interstices slightly elevated; surfaces shiny, transversely rugose at basal 1/4, closely punctate elsewhere.
|
|||
|
|
<emphasis id="2F9FEAC9B71DFFA56D84FC9247805913" bold="true" box="[1204,1257,836,862]" pageId="18" pageNumber="17">Legs</emphasis>
|
|||
|
|
: First tarsomere as long as the combined lengths of the remaining four on meso- and metatarsi; tibiae rounded in cross section; metatarsomeres I–III simple; metatarsomere IV excavated; metatarsomere V elongate with basally toothed claws.
|
|||
|
|
<emphasis id="2F9FEAC9B71DFFA5698FFC7F4264598E" bold="true" box="[191,269,937,963]" pageId="18" pageNumber="17">Venter</emphasis>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="85D02A5EB71DFFA5682AFC7E42E1598E" box="[282,392,936,963]" captionStart="Figures 31–34" captionStartId="18.[192,279,656,682]" captionTargetBox="[353,1279,213,626]" captionTargetId="figure-451@18.[353,1279,213,626]" captionTargetPageId="18" captionText="Figures 31–34. Costa Rican macraulacine eucnemids. 31) Onichodon confluentus sp. nov., male holotype, ventral habitus. 32) Onichodon confluentus sp. nov., female allotype, dorsal habitus. 33) Onichodon rufus sp. nov., dorsal habitus. 34) Onichodon rufus sp. nov., ventral habitus. (Scale: 31–32 = 5.0 mm; 33–34 = 1.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.10831099" httpUri="https://zenodo.org/record/10831099/files/figure.png" pageId="18" pageNumber="17">Figure 31</figureCitation>
|
|||
|
|
): Closely punctate, with short, recumbent yellowish setae; hypomeron with basally opened, lateral antennal grooves; metepisterna parallel-sided; elytral epipleura punctate; metacoxal plates medially 3.0–6.0 times wider than laterally.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="49946653B71DFFA569F0FD46479A58A0" ID-DOI="http://doi.org/10.5281/zenodo.10831099" ID-Zenodo-Dep="10831099" httpUri="https://zenodo.org/record/10831099/files/figure.png" pageId="18" pageNumber="17" startId="18.[192,279,656,682]" targetBox="[353,1279,213,626]" targetPageId="18" targetType="figure">
|
|||
|
|
<paragraph id="1D5436DBB71DFFA569F0FD46479A58A0" blockId="18.[191,1440,655,750]" pageId="18" pageNumber="17">
|
|||
|
|
<emphasis id="2F9FEAC9B71DFFA569F0FD46420058E7" bold="true" box="[192,361,656,682]" pageId="18" pageNumber="17">Figures 31–34.</emphasis>
|
|||
|
|
Costa Rican macraulacine eucnemids.
|
|||
|
|
<emphasis id="2F9FEAC9B71DFFA56A29FD59405658E4" bold="true" box="[793,831,655,681]" pageId="18" pageNumber="17">31)</emphasis>
|
|||
|
|
<taxonomicName id="DAEB4D58B71DFFA56A75FD46472858E7" authority="Otto & Muona & Córdoba-Alfaro, 2023" authorityName="Otto & Muona & Córdoba-Alfaro" authorityYear="2023" box="[837,1089,656,682]" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="confluentus" status="sp. nov.">
|
|||
|
|
<emphasis id="2F9FEAC9B71DFFA56A75FD46472858E7" box="[837,1089,656,682]" italics="true" pageId="18" pageNumber="17">Onichodon confluentus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="2F9FEAC9B71DFFA56D77FD4647F458E7" bold="true" box="[1095,1181,656,682]" pageId="18" pageNumber="17">
|
|||
|
|
<taxonomicNameLabel id="34AC57B2B71DFFA56D77FD4647F458E7" box="[1095,1181,656,682]" pageId="18" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, male holotype, ventral habitus.
|
|||
|
|
<emphasis id="2F9FEAC9B71DFFA5682EFD67422D5886" bold="true" box="[286,324,689,715]" pageId="18" pageNumber="17">32)</emphasis>
|
|||
|
|
<taxonomicName id="DAEB4D58B71DFFA5687AFD67412F5886" authority="Otto & Muona & Córdoba-Alfaro, 2023" authorityName="Otto & Muona & Córdoba-Alfaro" authorityYear="2023" box="[330,582,689,715]" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="confluentus" status="sp. nov.">
|
|||
|
|
<emphasis id="2F9FEAC9B71DFFA5687AFD67412F5886" box="[330,582,689,715]" italics="true" pageId="18" pageNumber="17">Onichodon confluentus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="2F9FEAC9B71DFFA56B7CFD6441CA5881" bold="true" box="[588,675,690,716]" pageId="18" pageNumber="17">
|
|||
|
|
<taxonomicNameLabel id="34AC57B2B71DFFA56B7CFD6441CA5881" box="[588,675,690,716]" pageId="18" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, female allotype, dorsal habitus.
|
|||
|
|
<emphasis id="2F9FEAC9B71DFFA56D3DFD67475A5886" bold="true" box="[1037,1075,689,715]" pageId="18" pageNumber="17">33)</emphasis>
|
|||
|
|
<taxonomicName id="DAEB4D58B71DFFA56D09FD67479B5881" authority="Otto & Muona & Córdoba-Alfaro, 2023" authorityName="Otto & Muona & Córdoba-Alfaro" authorityYear="2023" box="[1081,1266,689,716]" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="rufus" status="sp. nov.">
|
|||
|
|
<emphasis id="2F9FEAC9B71DFFA56D09FD67479B5881" box="[1081,1266,689,716]" italics="true" pageId="18" pageNumber="17">Onichodon rufus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="2F9FEAC9B71DFFA56DC8FD6446265881" bold="true" box="[1272,1359,690,716]" pageId="18" pageNumber="17">
|
|||
|
|
<taxonomicNameLabel id="34AC57B2B71DFFA56DC8FD6446265881" box="[1272,1359,690,716]" pageId="18" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, dorsal habitus.
|
|||
|
|
<emphasis id="2F9FEAC9B71DFFA5682EFD05422D58A0" bold="true" box="[286,324,723,749]" pageId="18" pageNumber="17">34)</emphasis>
|
|||
|
|
<taxonomicName id="DAEB4D58B71DFFA5687AFD05416A58A0" authority="Otto & Muona & Córdoba-Alfaro, 2023" authorityName="Otto & Muona & Córdoba-Alfaro" authorityYear="2023" box="[330,515,723,749]" class="Insecta" family="Eucnemidae" genus="Onichodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="rufus" status="sp. nov.">
|
|||
|
|
<emphasis id="2F9FEAC9B71DFFA5687AFD05416A58A0" box="[330,515,723,749]" italics="true" pageId="18" pageNumber="17">Onichodon rufus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="2F9FEAC9B71DFFA56B3AFD05410858A0" bold="true" box="[522,609,723,749]" pageId="18" pageNumber="17">
|
|||
|
|
<taxonomicNameLabel id="34AC57B2B71DFFA56B3AFD05410858A0" box="[522,609,723,749]" pageId="18" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, ventral habitus. (Scale: 31–32 = 5.0 mm; 33–34 = 1.0 mm)
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="1D5436DBB71DFFA56830FBDB47BD5E27" blockId="18.[191,1442,802,1131]" pageId="18" pageNumber="17">
|
|||
|
|
<emphasis id="2F9FEAC9B71DFFA56830FBDB42D15E6A" bold="true" box="[256,440,1037,1063]" pageId="18" pageNumber="17">
|
|||
|
|
Female
|
|||
|
|
<typeStatus id="C2508879B71DFFA5686AFBDB42D15E6A" box="[346,440,1037,1063]" pageId="18" pageNumber="17" type="allotype">allotype</typeStatus>
|
|||
|
|
</emphasis>
|
|||
|
|
(
|
|||
|
|
<figureCitation id="85D02A5EB71DFFA568F6FBDB415F5E6A" box="[454,566,1037,1063]" captionStart="Figures 31–34" captionStartId="18.[192,279,656,682]" captionTargetBox="[353,1279,213,626]" captionTargetId="figure-451@18.[353,1279,213,626]" captionTargetPageId="18" captionText="Figures 31–34. Costa Rican macraulacine eucnemids. 31) Onichodon confluentus sp. nov., male holotype, ventral habitus. 32) Onichodon confluentus sp. nov., female allotype, dorsal habitus. 33) Onichodon rufus sp. nov., dorsal habitus. 34) Onichodon rufus sp. nov., ventral habitus. (Scale: 31–32 = 5.0 mm; 33–34 = 1.0 mm)" figureDoi="http://doi.org/10.5281/zenodo.10831099" httpUri="https://zenodo.org/record/10831099/files/figure.png" pageId="18" pageNumber="17">Figure 32</figureCitation>
|
|||
|
|
): Length, 12.0 mm. Width, 4.0 mm. Antennae weakly and asymmetrically serriform, about 1/3 the length of the body; flagellomere I longer than wide, slightly longer than II; flagellomere II slightly longer than wide, shorter than III; flagellomeres III–VIII longer than wide, sub-equal.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="1D5436DBB71DFFA569F0FBAC41D25EB4" blockId="18.[192,1441,1146,1273]" pageId="18" pageNumber="17">
|
|||
|
|
<emphasis id="2F9FEAC9B71DFFA569F0FBAC42285ED9" bold="true" box="[192,321,1146,1172]" pageId="18" pageNumber="17">Variations.</emphasis>
|
|||
|
|
Seven male
|
|||
|
|
<typeStatus id="C2508879B71DFFA568E0FBAD41555ED8" box="[464,572,1147,1173]" pageId="18" pageNumber="17" type="paratype">paratypes</typeStatus>
|
|||
|
|
measured 6.0–
|
|||
|
|
<quantity id="DA139B3EB71DFFA56BDAFBAC40275ED8" box="[746,846,1146,1173]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="18" pageNumber="17" unit="mm" value="10.5">10.5 mm</quantity>
|
|||
|
|
long and
|
|||
|
|
<quantity id="DA139B3EB71DFFA56AF1FBAC47215ED8" box="[961,1096,1146,1173]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="18" pageNumber="17" unit="mm" value="2.0" valueMax="2.5" valueMin="1.5">1.5–2.5 mm</quantity>
|
|||
|
|
wide. Two male
|
|||
|
|
<typeStatus id="C2508879B71DFFA56C3AFBAD461F5ED8" box="[1290,1398,1147,1173]" pageId="18" pageNumber="17" type="paratype">paratypes</typeStatus>
|
|||
|
|
are larger than and just as wide as the
|
|||
|
|
<typeStatus id="C2508879B71DFFA56B66FB4A41D25EFB" box="[598,699,1180,1206]" pageId="18" pageNumber="17" type="holotype">holotype</typeStatus>
|
|||
|
|
. The remaining
|
|||
|
|
<specimenCount id="0BEDFD52B71DFFA56A46FB4A477A5EFB" box="[886,1043,1180,1206]" pageId="18" pageNumber="17" type="generic" typeStatus="paratypes">five paratypes</specimenCount>
|
|||
|
|
are shorter and narrower than the
|
|||
|
|
<typeStatus id="C2508879B71DFFA569F0FB6B424C5E9A" box="[192,293,1213,1239]" pageId="18" pageNumber="17" type="holotype">holotype</typeStatus>
|
|||
|
|
. Pronotal punctures are widely dispersed in
|
|||
|
|
<specimenCount id="0BEDFD52B71DFFA56A13FB6840D15E95" box="[803,952,1214,1240]" pageId="18" pageNumber="17" type="generic" typeStatus="paratype">one paratype</specimenCount>
|
|||
|
|
, whereas the remaining
|
|||
|
|
<specimenCount id="0BEDFD52B71DFFA56DE1FB6B460D5E95" box="[1233,1380,1213,1240]" pageId="18" pageNumber="17" type="generic" typeStatus="paratypes">six paratypes</specimenCount>
|
|||
|
|
have exoskeletal structures similar to the
|
|||
|
|
<typeStatus id="C2508879B71DFFA56B62FB0941DE5EB4" box="[594,695,1247,1273]" pageId="18" pageNumber="17" type="holotype">holotype</typeStatus>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="55F16550B71DFFA569F0FADE421B5F09" pageId="18" pageNumber="17" type="distribution">
|
|||
|
|
<paragraph id="1D5436DBB71DFFA569F0FADE421B5F09" blockId="18.[192,1440,1288,1348]" pageId="18" pageNumber="17">
|
|||
|
|
<emphasis id="2F9FEAC9B71DFFA569F0FADE42325F6F" bold="true" box="[192,347,1288,1314]" pageId="18" pageNumber="17">Distribution.</emphasis>
|
|||
|
|
This eucnemid species is known from
|
|||
|
|
<specimenCount id="0BEDFD52B71DFFA56A21FADE40A95F6F" box="[785,960,1288,1314]" pageId="18" pageNumber="17" type="generic">nine specimens</specimenCount>
|
|||
|
|
taken from two locations within the province of
|
|||
|
|
<collectingRegion id="DF2FF839B71DFFA56821FAFC42045F09" box="[273,365,1322,1348]" country="Costa Rica" name="Heredia" pageId="18" pageNumber="17">Heredia</collectingRegion>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="55F16550B71DFFA569F0FA8540185FC2" pageId="18" pageNumber="17" type="biology_ecology">
|
|||
|
|
<paragraph id="1D5436DBB71DFFA569F0FA8540185FC2" blockId="18.[192,1440,1363,1424]" pageId="18" pageNumber="17">
|
|||
|
|
<emphasis id="2F9FEAC9B71DFFA569F0FA8542495F20" bold="true" box="[192,288,1363,1389]" pageId="18" pageNumber="17">Biology.</emphasis>
|
|||
|
|
Two specimens were taken from a primary forest. Five specimens, including the
|
|||
|
|
<typeStatus id="C2508879B71DFFA56D8BFA8246745F23" box="[1211,1309,1364,1390]" pageId="18" pageNumber="17" type="holotype">holotype</typeStatus>
|
|||
|
|
were taken from a second growth forest. Larvae and pupae are unknown.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="55F16550B71DFFA569F0FA4946C85FF4" box="[192,1441,1439,1465]" pageId="18" pageNumber="17" type="etymology">
|
|||
|
|
<paragraph id="1D5436DBB71DFFA569F0FA4946C85FF4" blockId="18.[192,1441,1439,1465]" box="[192,1441,1439,1465]" pageId="18" pageNumber="17">
|
|||
|
|
<emphasis id="2F9FEAC9B71DFFA569F0FA49422D5FF4" bold="true" box="[192,324,1439,1465]" pageId="18" pageNumber="17">Etymology.</emphasis>
|
|||
|
|
The specific epithet is named from the presence of confluent surfaces of the frons on the new species.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|