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<document id="A1B3081742A640C2E3BB603A56A520B8" ID-DOI="10.1649/0010-065x-63.sp7.1" ID-ISSN="1938-4394" ID-Zenodo-Dep="4912048" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623157183841" checkinUser="carolina" docAuthor="Seago, Ainsley E." docDate="2009" docId="039887BAFFD5C07992E083B8EFCE9655" docLanguage="en" docName="TheColeopBull.63.mo7.1-73.pdf.imf" docOrigin="The Coleopterists Bulletin (mo 7) 63" docStyle="DocumentStyle:A431C8F32F3D86E670C18C13DA9B5F2B.4:TheColeopBull.2000-2009.monograph" docStyleId="A431C8F32F3D86E670C18C13DA9B5F2B" docStyleName="TheColeopBull.2000-2009.monograph" docStyleVersion="4" docTitle="Agyrtodes lescheni Seago 2009, new species" docType="treatment" docVersion="5" lastPageNumber="53" masterDocId="FFA1FFC2FFE7C04D936D8227ED609758" masterDocTitle="Revision Of Agyrtodes Portevin (Coleoptera: Leiodidae)" masterLastPageNumber="73" masterPageNumber="1" pageNumber="51" updateTime="1698957853775" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="CFE635BE7A16375080D33D1BF197C754" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="31B8BAB6B6B0BD3240E73720D157DDD5">
<mods:title id="91054232B265A4357E8EF82017475A75">Revision Of Agyrtodes Portevin (Coleoptera: Leiodidae)</mods:title>
</mods:titleInfo>
<mods:name id="38834391469A241AC45E1739852ED1D5" type="personal">
<mods:role id="158166FCB4F554B26283FDF1B0DD22DC">
<mods:roleTerm id="B6A44ACF4A628329C60D60A6C5856B5B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5FE01D5A569DD6C5C4DE33EED8B56CF7">Seago, Ainsley E.</mods:namePart>
<mods:affiliation id="6B40ACABA3D731C1C45FDABBDE4745FC">ESPM- Div. of Insect Biology 137 Mulford Hall, University of California Berkeley, CA 94720 - 3114</mods:affiliation>
<mods:nameIdentifier id="D5474480B35F7AFE7C6B2374B3A46617" type="email">seago@nature.berkeley.edu</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="22EEF081055B85AA1B188C64902BF322">text</mods:typeOfResource>
<mods:relatedItem id="C6937F5F9AF39C2FB946321165A66677" type="host">
<mods:titleInfo id="C853F18DB3824BFA6321BC75610E0778">
<mods:title id="BDB0D4947EAD6FE073029A25C91C0C46">The Coleopterists Bulletin</mods:title>
</mods:titleInfo>
<mods:part id="0B6F6A4A1E789F50A04CDAD7ED5795D2">
<mods:date id="041D6172DDB01881EF4704E0B5A270B9">2009</mods:date>
<mods:detail id="B49B5372F047F44BA2065396A77C80A5" type="series">
<mods:title id="17A7313D6C7CF26965DDE6AAE375001D">mo 7</mods:title>
</mods:detail>
<mods:detail id="F84047A26CE975453BFF8CE3B28D4AFA" type="pubDate">
<mods:number id="0257897F74CD54920734FEE9B871BDF0">2009-04-22</mods:number>
</mods:detail>
<mods:detail id="603A5CAA413306543BCF90E4C4DF51E5" type="volume">
<mods:number id="CA933C832ACB3E1CEA589C318B9DFDF9">63</mods:number>
</mods:detail>
<mods:extent id="1FFC23761D8F8A91407F24793FC39786" unit="page">
<mods:start id="7079D2F5F492A995A75D85D42C54DB6E">1</mods:start>
<mods:end id="E85CBFA0B98E439CD9124BDEE956888B">73</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="2D629DA9E10E7F01A340ABD6B9F8F711">journal article</mods:classification>
<mods:identifier id="901447AB1D4E646E8EFE7E33F5606531" type="DOI">10.1649/0010-065x-63.sp7.1</mods:identifier>
<mods:identifier id="EAA70081CB2A5DAC5237FE481940455A" type="ISSN">1938-4394</mods:identifier>
<mods:identifier id="2602DED84DB33066C19908E34A4B36D4" type="Zenodo-Dep">4912048</mods:identifier>
<mods:identifier id="2490877AE64FB6B9F57D0D019419F5A6" type="ZooBank">B58B2216-0CFA-41C6-8141-7024E65ECF85</mods:identifier>
</mods:mods>
<treatment id="039887BAFFD5C07992E083B8EFCE9655" ID-DOI="http://doi.org/10.5281/zenodo.4912305" ID-Zenodo-Dep="4912305" LSID="urn:lsid:plazi:treatment:039887BAFFD5C07992E083B8EFCE9655" httpUri="http://treatment.plazi.org/id/039887BAFFD5C07992E083B8EFCE9655" lastPageId="52" lastPageNumber="53" pageId="50" pageNumber="51">
<subSubSection id="C32B6527FFD5C07F92E083B8EFB19689" pageId="50" pageNumber="51" type="nomenclature">
<paragraph id="8B8E36ACFFD5C07F92E083B8EE6E96ED" blockId="50.[397,782,415,465]" box="[397,782,415,437]" pageId="50" pageNumber="51">
<heading id="D0C681C0FFD5C07F92E083B8EE6E96ED" box="[397,782,415,437]" centered="true" fontSize="8" level="2" pageId="50" pageNumber="51" reason="2">
<emphasis id="B945EABEFFD5C07F92E083B8EE6E96ED" bold="true" box="[397,782,415,437]" pageId="50" pageNumber="51">
<taxonomicName id="4C314D2FFFD5C07F92E083B8EFEE96ED" ID-CoL="BBVV" authority="Seago" authorityName="Seago" authorityYear="2009" box="[397,654,415,437]" class="Insecta" family="Leiodidae" genus="Agyrtodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="lescheni" status="sp. nov.">
<emphasis id="B945EABEFFD5C07F92E083B8EF2896ED" bold="true" box="[397,584,415,437]" italics="true" pageId="50" pageNumber="51">Agyrtodes lescheni</emphasis>
Seago
</taxonomicName>
,
<taxonomicNameLabel id="A27657C5FFD5C07F91F183B8EE6E96ED" box="[668,782,415,437]" pageId="50" pageNumber="51" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8E36ACFFD5C07F92A7839CEFB19689" blockId="50.[397,782,415,465]" box="[458,721,443,465]" pageId="50" pageNumber="51">(Figs. 16, 53, 73, 93, 103)</paragraph>
</subSubSection>
<subSubSection id="C32B6527FFD5C07F93AA83D6EFD19500" pageId="50" pageNumber="51" type="materials_examined">
<paragraph id="8B8E36ACFFD5C07F93AA83D6EFB79565" blockId="50.[175,1005,497,1570]" pageId="50" pageNumber="51">
<materialsCitation id="3B593CF1FFD5C07F93AA83D6EFB79565" collectionCode="V, FMNH" collectorName="N. Zeal. &amp; S. Isl. &amp; S &amp; J Peck" elevation="250" location="Reefton" pageId="50" pageNumber="51" specimenCount="1" typeStatus="holotype">
<emphasis id="B945EABEFFD5C07F93AA83D6EC43955F" bold="true" box="[199,291,497,519]" pageId="50" pageNumber="51">
<typeStatus id="548A880EFFD5C07F93AA83D6EC43955F" box="[199,291,497,519]" pageId="50" pageNumber="51" type="holotype">Holotype</typeStatus>
</emphasis>
: male, point-mounted with genitalia mounted separately on acetate, labeled ‘‘
<collectorName id="26C4537AFFD5C07F9267802BEC04957A" box="[266,356,524,546]" pageId="50" pageNumber="51">N. Zeal.</collectorName>
;
<collectorName id="26C4537AFFD5C07F9203802BECC9957A" box="[366,425,524,546]" pageId="50" pageNumber="51">S. Isl.</collectorName>
<quantity id="4CC99B49FFD5C07F92DD802BEC91957A" box="[432,497,524,546]" metricMagnitude="4" metricUnit="m" metricValue="1.4" pageId="50" pageNumber="51" unit="km" value="14.0">14 km</quantity>
SE
<location id="8EEE6077FFD5C07F9170802BEF11957A" LSID="urn:lsid:plazi:treatment:039887BAFFD5C07992E083B8EFCE9655:8EEE6077FFD5C07F9170802BEF11957A" box="[541,625,524,546]" name="Reefton" pageId="50" pageNumber="51">Reefton</location>
,
<quantity id="4CC99B49FFD5C07F9111802BEFDE957A" box="[636,702,524,546]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="50" pageNumber="51" unit="m" value="250.0">
<elevation id="001CD19FFFD5C07F9111802BEFDE957A" box="[636,702,524,546]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="50" pageNumber="51" unit="m" value="250.0">250 m</elevation>
</quantity>
, 29.
<collectionCode id="ED20AE69FFD5C07F9182802BEE63957A" box="[751,771,524,546]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="50" pageNumber="51" type="Museum">V</collectionCode>
. 82,
<collectorName id="26C4537AFFD5C07F9059802BEEC7957A" box="[820,935,524,546]" pageId="50" pageNumber="51">S &amp; J Peck</collectorName>
, beech forest litter/
<taxonomicName id="4C314D2FFFD5C07F925C8000ECF79564" box="[305,407,551,572]" class="Insecta" family="Leiodidae" genus="Agyrtodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B945EABEFFD5C07F925C8000ECF79564" box="[305,407,551,572]" italics="true" pageId="50" pageNumber="51">Agyrtodes</emphasis>
</taxonomicName>
det.
<bibRefCitation id="EFA04B5DFFD5C07F92A18000EF3C9565" author="Newton, A. F., Jr." box="[460,604,551,573]" pageId="50" pageNumber="51" pagination="41 - 178" refId="ref34526" refString="Newton, A. F., Jr. 1998. Phylogenetic problems, current classification and generic catalog of world Leiodidae (including Cholevidae). Phylogeny and Evolution of Subterranean and Endogean Cholevidae (5 Leiodidae Cholevinae) [pp. 41 - 178]. In: Proceedings of a Symposium (30 August, 1996, Florence, Italy), XX. International Congress of Entomology. 178 pp." type="book chapter" year="1998">Newton 1998</bibRefCitation>
.’’ (
<collectionCode id="ED20AE69FFD5C07F91148000EFAC9565" box="[633,716,551,573]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="50" pageNumber="51" type="Museum">FMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8B8E36ACFFD5C07F93AA8065EFD19500" blockId="50.[175,1005,497,1570]" box="[199,689,578,600]" pageId="50" pageNumber="51">
<emphasis id="B945EABEFFD5C07F93AA8065EC4C9500" bold="true" box="[199,300,578,600]" pageId="50" pageNumber="51">
<typeStatus id="548A880EFFD5C07F93AA8065EC4C9500" box="[199,300,578,600]" pageId="50" pageNumber="51" type="paratype">Paratypes</typeStatus>
</emphasis>
: listed in Appendix 1 (
<specimenCount id="9D37FD25FFD5C07F91768065EFC69500" box="[539,678,578,600]" pageId="50" pageNumber="51" type="generic">80 specimens</specimenCount>
).
</paragraph>
</subSubSection>
<subSubSection id="C32B6527FFD5C07F93AA8079EC7B95D7" pageId="50" pageNumber="51" type="distribution">
<paragraph id="8B8E36ACFFD5C07F93AA8079EC7B95D7" blockId="50.[175,1005,497,1570]" pageId="50" pageNumber="51">
<emphasis id="B945EABEFFD5C07F93AA8079EC26952C" bold="true" box="[199,326,606,628]" pageId="50" pageNumber="51">Distribution.</emphasis>
<collectingCountry id="F326763CFFD5C07F92218079ECB9952C" box="[332,473,606,628]" name="New Zealand" pageId="50" pageNumber="51">New Zealand</collectingCountry>
: South Island,
<collectingRegion id="49F5F84EFFD5C07F911F8079EFDD952C" box="[626,701,606,628]" country="New Zealand" name="Nelson" pageId="50" pageNumber="51">Nelson</collectingRegion>
,
<collectingRegion id="49F5F84EFFD5C07F91AA8079EE34952C" box="[711,852,606,628]" country="New Zealand" name="Marlborough" pageId="50" pageNumber="51">Marlborough</collectingRegion>
, Buller regions (
<figureCitation id="130A2A29FFD5C07F93DA805EEC6F95D7" box="[183,271,633,655]" captionStart="Fig" captionStartId="47.[167,198,1096,1115]" captionTargetBox="[180,940,194,1074]" captionTargetId="figure-145@47.[180,940,194,1074]" captionTargetPageId="47" captionText="Fig. 103. Geographic distribution of Agyrtodes spp. in New Zealand, part 2. A. nebulosus, A. hunuensis, A. lescheni, new species, A. nemoralis." figureDoi="http://doi.org/10.5281/zenodo.4912082" httpUri="https://zenodo.org/record/4912082/files/figure.png" pageId="50" pageNumber="51">Fig. 103</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32B6527FFD5C07F93AA80B2EE61943E" pageId="50" pageNumber="51" type="diagnosis">
<paragraph id="8B8E36ACFFD5C07F93AA80B2EE61943E" blockId="50.[175,1005,497,1570]" pageId="50" pageNumber="51">
<emphasis id="B945EABEFFD5C07F93AA80B2EC4F95F3" bold="true" box="[199,303,661,683]" pageId="50" pageNumber="51">Diagnosis.</emphasis>
This species can be recognized by the transverse pale gold markings formed by the dorsal pubescence, and by the short mesofemoral and long metafemoral spurs.
<taxonomicName id="4C314D2FFFD5C07F92E980EDEF349587" baseAuthorityName="Broun" baseAuthorityYear="1880" box="[388,596,714,735]" class="Insecta" family="Leiodidae" genus="Agyrtodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="nebulosus">
<emphasis id="B945EABEFFD5C07F92E980EDEF349587" box="[388,596,714,735]" italics="true" pageId="50" pageNumber="51">Agyrtodes nebulosus</emphasis>
</taxonomicName>
, another
<collectingCountry id="F326763CFFD5C07F91AC80EDEE2F95B8" box="[705,847,714,736]" name="New Zealand" pageId="50" pageNumber="51">New Zealand</collectingCountry>
species, has a similarly colored dorsum but lacks mesofemoral spurs; the parameres of
<taxonomicName id="4C314D2FFFD5C07F90B880C2EC6E944D" baseAuthorityName="Broun" baseAuthorityYear="1880" class="Insecta" family="Leiodidae" genus="Agyrtodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="nebulosus">
<emphasis id="B945EABEFFD5C07F90B880C2EC6E944D" italics="true" pageId="50" pageNumber="51">A. nebulosus</emphasis>
</taxonomicName>
are also broadly expanded, abruptly truncate and twisted at the apices, whereas the parameres of
<taxonomicName id="4C314D2FFFD5C07F92D2813CEF4C9468" authorityName="Seago" authorityYear="2009" box="[447,556,795,816]" class="Insecta" family="Leiodidae" genus="Agyrtodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="lescheni">
<emphasis id="B945EABEFFD5C07F92D2813CEF4C9468" box="[447,556,795,816]" italics="true" pageId="50" pageNumber="51">A. lescheni</emphasis>
</taxonomicName>
are simple and slender. In males, the long, curved, ventrally spinose first mesotarsomere of
<taxonomicName id="4C314D2FFFD5C07F91F68111EE779413" baseAuthorityName="Broun" baseAuthorityYear="1880" box="[667,791,822,843]" class="Insecta" family="Leiodidae" genus="Agyrtodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="nebulosus">
<emphasis id="B945EABEFFD5C07F91F68111EE779413" box="[667,791,822,843]" italics="true" pageId="50" pageNumber="51">A. nebulosus</emphasis>
</taxonomicName>
distinguishes it from
<taxonomicName id="4C314D2FFFD5C07F93C28177EC7C943D" authorityName="Seago" authorityYear="2009" box="[175,284,848,869]" class="Insecta" family="Leiodidae" genus="Agyrtodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="lescheni">
<emphasis id="B945EABEFFD5C07F93C28177EC7C943D" box="[175,284,848,869]" italics="true" pageId="50" pageNumber="51">A. lescheni</emphasis>
</taxonomicName>
which has simple, unmodified male mesotarsi.
</paragraph>
</subSubSection>
<subSubSection id="C32B6527FFD5C07E93AA814BEC45917A" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51" type="description">
<paragraph id="8B8E36ACFFD5C07F93AA814BEC8194C5" blockId="50.[175,1005,497,1570]" pageId="50" pageNumber="51">
<emphasis id="B945EABEFFD5C07F93AA814BEC2094DA" bold="true" box="[199,320,876,898]" pageId="50" pageNumber="51">Description.</emphasis>
TBL
<emphasis id="B945EABEFFD5C07F9210814BECF194DA" bold="true" box="[381,401,876,898]" pageId="50" pageNumber="51">5</emphasis>
<quantity id="4CC99B49FFD5C07F92F5814BEC8894DA" box="[408,488,876,898]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="50" pageNumber="51" unit="mm" value="2.4">2.4 mm</quantity>
, EW
<emphasis id="B945EABEFFD5C07F914C814BEF5594DA" bold="true" box="[545,565,876,898]" pageId="50" pageNumber="51">5</emphasis>
<quantity id="4CC99B49FFD5C07F9156814BEFEC94DA" box="[571,652,876,898]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="50" pageNumber="51" unit="mm" value="1.4">1.4 mm</quantity>
, PNW
<emphasis id="B945EABEFFD5C07F91BB814BEF8A94DA" bold="true" box="[726,746,876,898]" pageId="50" pageNumber="51">5</emphasis>
<quantity id="4CC99B49FFD5C07F919C814BEE2194DA" box="[753,833,876,898]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="50" pageNumber="51" unit="mm" value="1.2">1.2 mm</quantity>
, HW
<emphasis id="B945EABEFFD5C07F9013814BEEF294DA" bold="true" box="[894,914,876,898]" pageId="50" pageNumber="51">5</emphasis>
<quantity id="4CC99B49FFD5C07F90F5814BEE8994DA" box="[920,1001,876,898]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="50" pageNumber="51" unit="mm" value="0.5">0.5 mm</quantity>
. (TBL variation:
<quantity id="4CC99B49FFD5C07F923781A0ECB694C5" box="[346,470,903,925]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.6" metricValueMin="2.4" pageId="50" pageNumber="51" unit="mm" value="2.5" valueMax="2.6" valueMin="2.4">2.4–2.6 mm</quantity>
).
</paragraph>
<paragraph id="8B8E36ACFFD5C07F93AA8185EFBF94B6" blockId="50.[175,1005,497,1570]" pageId="50" pageNumber="51">Body (Fig. 16) large, elongately ovate, dorsum dark reddish-brown with wavy, confused vestiture of long light brown and golden hairs, with transverse pale markings on elytra emphasized by pale gold vestiture.</paragraph>
<paragraph id="8B8E36ACFFD5C07F93AA81D4EEF592C3" blockId="50.[175,1005,497,1570]" pageId="50" pageNumber="51">Head quadrate, shallowly punctate, eyes of medium size, narrowly emarginate posteriorly. Epistomal suture present, with stem. Clypeus broad, ovate, excavate anteriorly; labrum transverse, broadly rectangular, with anterior margin weakly excavate. Mandibles large, outer edges strongly curved, with dense field of pores on ventral surface. Maxilla with slender galea bearing weakly sclerotized apical brush of short hairs, lacinia broader than galea, well-sclerotized, with large apical spore-brush of stout, curving, minute spines arranged in parallel rows; maxillary palpi with terminal segment slender, elongate, lanceolate, produced distally and truncate at apex. Penultimate segment broader than terminal segment, nearly twice as long as wide, sparsely armed with long, stiff setae; first segment triangular, constricted basally, apically as wide as base of penultimate segment. Labium with four large submarginal digitiform sensillae and two much shorter, stout, apically truncate digitiform sensillae at anterior margin of ligula. Labial palpi with apical segment short, slender, cylindrical, very weakly curved inward and bearing large patch of minute digitiform sensillae in basal half. Penultimate segment extremely short, transverse, wider than base of apical segment.</paragraph>
<paragraph id="8B8E36ACFFD5C07E93AA8786EC6C9607" blockId="50.[175,1005,497,1570]" lastBlockId="51.[146,975,194,1570]" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51">
Antennae (
<figureCitation id="130A2A29FFD5C07F92508786ECE792EF" box="[317,391,1441,1463]" captionStart="Figs" captionStartId="9.[167,206,1301,1320]" captionTargetBox="[197,877,194,1279]" captionTargetId="figure-57@9.[197,924,194,1279]" captionTargetPageId="9" captionText="Figs. 48–57. Antennae of Agyrtodes spp. 48) A. atropos; 49) A. koebelei; 50) A. variabilis, new species; 51) A. bicolor; 52) A. nebulosus; 53) A. lescheni, new species; 54) A. labralis; 55) A. monticola; 56) A. nemoralis; 57) A. hunuensis." figureDoi="http://doi.org/10.5281/zenodo.4912129" httpUri="https://zenodo.org/record/4912129/files/figure.png" pageId="50" pageNumber="51">Fig. 53</figureCitation>
) slender, basal segments light yellowish-brown, gradually darkening to reddish-brown at apex. Segment 1 cylindrical, elongate, gently curved; segment 2 wider than 1, fusiform with truncate base and apex. Segments 3–6 all slender, cylindrical, length.3
<emphasis id="B945EABEFFD5C07F915787D6EF2E915F" bold="true" box="[570,590,1521,1543]" pageId="50" pageNumber="51">3</emphasis>
width; segments 4 and 5 of equal size, segment 3 slightly longer than segment 6. Segment 7 barely longer than 6, approximately twice as wide as 6, broader distally. Segment 8 cylindrical, elongate, nearly 2
<emphasis id="B945EABEFFD4C07E922382FAEC0297AB" bold="true" box="[334,354,221,243]" pageId="51" pageNumber="52">3</emphasis>
as long as wide, of equivalent width to bases of 7 and 9. Segment 9 large, rounded basally, as broad and 7 and slightly shorter; segment 10 as broad as 9, shorter than 9 but still longer than wide. Segment 11 large, ovate, longer than each preceding segment, weakly produced at apex, bluntly papilliform.
</paragraph>
<paragraph id="8B8E36ACFFD4C07E93C78343EC829693" blockId="51.[146,975,194,1570]" pageId="51" pageNumber="52">Pronotum medium to dark reddish-brown, integument shining and finely punctate, vestiture of long, light golden setae. Lateral margins of pronotum gently curved; hind angles bluntly pointed, weakly produced posteriad. Hind margin of pronotum narrowly transparent.</paragraph>
<paragraph id="8B8E36ACFFD4C07E93C783F6EE7A9536" blockId="51.[146,975,194,1570]" pageId="51" pageNumber="52">Elytra with transverse strigae shallowly impressed, marked by puncture rows at setal insertions, without traces of longitudinal strial impressions. Each elytron with several short, transverse pale markings delimited by light golden setae: one at base, one immediately posterior to scutellum, adjacent to suture, one laterally, midlength, and one immediately anterior to elytral apex; vestiture dark brown on darker regions of elytra. Scutellum small, dark reddish-brown.</paragraph>
<paragraph id="8B8E36ACFFD4C07E93C78053EF0E9473" blockId="51.[146,975,194,1570]" pageId="51" pageNumber="52">Hypomeron and epipleura pale orange-brown, hypomeron strongly convex posterior to coxal insertions. Mesosternum and mestasternum dark reddishbrown, median carina elevated, obtusely pointed anterior to mesocoxal separation. Mesepimera extremely short, subquadrate, squarely truncate at coxal articulations. Metasternum with slightly produced setose median bulge, laterally convex and shallowly punctate. Metepisternum narrow, elongate, square anteriorly and tapering to a point posteriorly.</paragraph>
<paragraph id="8B8E36ACFFD4C07E93C78116EEDE943A" blockId="51.[146,975,194,1570]" pageId="51" pageNumber="52">Abdominal sterna III–VI normal, male sternum VII broadly, shallowly emarginate. Male sternum VIII deeply emarginate but not completely divided.</paragraph>
<paragraph id="8B8E36ACFFD4C07E93C7814FEC3B9378" blockId="51.[146,975,194,1570]" pageId="51" pageNumber="52">Legs of normal proportions, males with short, blunt spur arising from hind margin of mesofemur and longer, more acutely pointed spur arising from hind margin of metafemur. Male protarsi with first three segments expanded, cordate and bearing thick pads of tenent setae; first protarsomere subquadrate and slightly narrower than tibial apex. Male mesotarsi not expanded, first three segments bearing sparse pads of tenent setae. Claws and female tarsi simple.</paragraph>
<paragraph id="8B8E36ACFFD4C07E93C78602EE249211" blockId="51.[146,975,194,1570]" pageId="51" pageNumber="52">
Male genital segment (
<figureCitation id="130A2A29FFD4C07E92F18602EC869363" box="[412,486,1061,1083]" captionStart="Figs" captionStartId="12.[196,235,1299,1318]" captionTargetBox="[194,986,194,1276]" captionTargetId="figure-1@12.[194,986,194,1276]" captionTargetPageId="12" captionText="Figs. 68–77. Male genital segment of Agyrtodes spp. 68) A. atropos; 69) A. koebelei; 70) A. variabilis, new species; 71) A. bicolor; 72) A. nebulosus; 73) A. lescheni, new species; 74) A. labralis; 75) A. monticola; 76) A. nemoralis.; 77) A. hunuensis." figureDoi="http://doi.org/10.5281/zenodo.4912062" httpUri="https://zenodo.org/record/4912062/files/figure.png" pageId="51" pageNumber="52">Fig. 73</figureCitation>
) ovate, convex, nearly globular; pleurites and tergite bluntly pointed at apices, anterior apophysis short, not expanded anteriorly, weakly sclerotized, pleurites weakly fused along median line. Sternum X present as an elongate, tongue-shaped sclerite fused interiorly to junction of pleurites. Aedeagus (
<figureCitation id="130A2A29FFD4C07E920186B6ECD693FF" box="[364,438,1169,1191]" captionStart="Figs" captionStartId="14.[196,235,1332,1351]" captionTargetBox="[182,999,194,1309]" captionTargetId="figure-1@14.[180,1000,194,1309]" captionTargetPageId="14" captionText="Figs. 88–97. Aedeagus (median lobe and parameres) of Agyrtodes spp. 88) A. atropos; 89) A. koebelei; 90) A. variabilis, new species; 91) A. bicolor; 92) A. nebulosus; 93) A. lescheni, new species; 94) A. labralis; 95) A. monticola; 96) A. nemoralis; 97) A. hunuensis." figureDoi="http://doi.org/10.5281/zenodo.4912066" httpUri="https://zenodo.org/record/4912066/files/figure.png" pageId="51" pageNumber="52">Fig. 93</figureCitation>
) with median lobe simple, broad basally, narrowly triangular in distal half, bluntly pointed at apex; basal piece flared proximally, broader and slightly shorter than penis. Parameres straight, slender, weakly constricted at bases, faintly expanded and flattened at apices, barely extending past apex of penis; each paramere apex armed on interior face with sparse array of minute setae. Endophallus armed with several rows of minute, sharply pointed well-sclerotized teeth and four long, curving, fang-shaped sclerites.
</paragraph>
<paragraph id="8B8E36ACFFD4C07E93C78768EDB392EE" blockId="51.[146,975,194,1570]" pageId="51" pageNumber="52">
Female genitalia with styli long, slender, coxites bearing three long setae on lateral faces,,2.5
<emphasis id="B945EABEFFD4C07E9235874DEC0C92D8" bold="true" box="[344,364,1386,1408]" pageId="51" pageNumber="52">3</emphasis>
as long and as wide as styli; styli short, very slender, cylindrical, each with minute, subapical seta and terminal seta.2
<emphasis id="B945EABEFFD4C07E903F87A2EE0692C3" bold="true" box="[850,870,1413,1435]" pageId="51" pageNumber="52">3</emphasis>
length of stylus.
</paragraph>
<paragraph id="8B8E36ACFFD4C07E93C7879CEC45917A" blockId="51.[146,975,194,1570]" pageId="51" pageNumber="52">
<emphasis id="B945EABEFFD4C07E93C7879CEC379289" bold="true" box="[170,343,1467,1489]" pageId="51" pageNumber="52">Natural History.</emphasis>
Collected by hand on resupinate polypores under peeling bark, by pyrethrin-fogging fungusy logs, on decaying nikau palm fronds on ground, on
<taxonomicName id="4C314D2FFFD4C07E927887D5ECE8915F" authorityName="P.A.Karsten" authorityYear="1881" box="[277,392,1522,1543]" class="Agaricomycetes" family="Ganodermataceae" genus="Ganoderma" kingdom="Fungi" order="Polyporales" pageId="51" pageNumber="52" phylum="Basidiomycota" rank="genus">
<emphasis id="B945EABEFFD4C07E927887D5ECE8915F" box="[277,392,1522,1543]" italics="true" pageId="51" pageNumber="52">Ganoderma</emphasis>
</taxonomicName>
pore surface, in carrion traps, in sifted litter, and in window traps.
</paragraph>
</subSubSection>
<subSubSection id="C32B6527FFD3C07993AA82E5EFCE9655" pageId="52" pageNumber="53" type="etymology">
<paragraph id="8B8E36ACFFD3C07993AA82E5EFCE9655" blockId="52.[175,1004,194,269]" pageId="52" pageNumber="53">
<emphasis id="B945EABEFFD3C07993AA82E5EC5A9780" bold="true" box="[199,314,194,216]" pageId="52" pageNumber="53">Etymology.</emphasis>
This species is named for coleopterist and musician Richard Leschen, who provided many agyrtodine specimens, ecological data, and helpful comments to the benefit of the author’s research.
</paragraph>
</subSubSection>
</treatment>
</document>
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