treatments-xml/data/60/65/87/606587D11C00FF8BFE1106693CFDB972.xml

<document ID-DOI="10.17109/AZH.68.1.85.2022" ID-ISSN="2064-2474" ID-Zenodo-Dep="7160782" approvalRequired="9" approvalRequired_for_textStreams="9" checkinTime="1665198500562" checkinUser="felipe" docAuthor="Csorba, Gábor &amp; Furey, Neil M." docDate="2022" docId="606587D11C00FF8BFE1106693CFDB972" docLanguage="en" docName="ActaZoolAcadSciH.68.1.85-97.pdf" docOrigin="Acta Zoologica Academiae Scientiarum Hungaricae 68 (1)" docSource="http://dx.doi.org/10.17109/azh.68.1.85.2022" docStyle="DocumentStyle:183DC955716D8EEDE76F04077FE39880.1:ActaZoolAcadSciH.2015-.journal_article" docStyleId="183DC955716D8EEDE76F04077FE39880" docStyleName="ActaZoolAcadSciH.2015-.journal_article" docStyleVersion="1" docTitle="Myotis hayesi Csorba &amp; Furey 2022, sp. n." docType="treatment" docVersion="2" lastPageNumber="94" masterDocId="9C5CFFA91C04FF82FFF1043E3D4CB835" masterDocTitle="From Greener Times: A New Species Of Thick-Thumbed Myotis From Phnom Penh, Cambodia" masterLastPageNumber="97" masterPageNumber="85" pageNumber="89" updateTime="1665200060917" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title>From Greener Times: A New Species Of Thick-Thumbed Myotis From Phnom Penh, Cambodia</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Csorba, Gábor</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Furey, Neil M.</mods:namePart>
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<mods:title>Acta Zoologica Academiae Scientiarum Hungaricae</mods:title>
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<mods:date>2022</mods:date>
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<mods:number>2022-02-14</mods:number>
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<mods:number>68</mods:number>
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<mods:url>http://dx.doi.org/10.17109/azh.68.1.85.2022</mods:url>
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<mods:identifier type="DOI">10.17109/AZH.68.1.85.2022</mods:identifier>
<mods:identifier type="ISSN">2064-2474</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:606587D11C00FF8BFE1106693CFDB972" httpUri="http://treatment.plazi.org/id/606587D11C00FF8BFE1106693CFDB972" lastPageId="9" lastPageNumber="94" pageId="4" pageNumber="89">
<subSubSection box="[480,745,599,626]" pageId="4" pageNumber="89" type="nomenclature">
<paragraph blockId="4.[480,745,599,626]" box="[480,745,599,626]" pageId="4" pageNumber="89">
<heading box="[480,745,599,626]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="89" reason="2">
<taxonomicName authority="Csorba &amp; Furey, 2022" authorityName="Csorba &amp; Furey" authorityYear="2022" box="[480,668,599,625]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="4" pageNumber="89" phylum="Chordata" rank="species" species="hayesi" status="sp. n.">
<emphasis bold="true" box="[480,668,599,625]" pageId="4" pageNumber="89">Myotis hayesi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[675,745,600,626]" pageId="4" pageNumber="89" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection lastPageId="9" lastPageNumber="94" pageId="4" pageNumber="89" type="description">
<paragraph blockId="4.[550,674,634,660]" box="[550,674,634,660]" pageId="4" pageNumber="89">
<heading box="[550,674,634,660]" centered="true" fontSize="11" level="3" pageId="4" pageNumber="89" reason="3">
(
<figureCitation box="[559,666,634,660]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStart-4="Fig" captionStart-5="Fig" captionStartId-0="5.[174,210,567,589]" captionStartId-1="5.[174,210,1524,1546]" captionStartId-2="6.[136,172,889,911]" captionStartId-3="6.[136,172,1525,1547]" captionStartId-4="7.[174,210,1458,1480]" captionStartId-5="10.[136,172,1467,1489]" captionTargetBox-0="[176,1123,194,533]" captionTargetBox-1="[174,1122,1132,1497]" captionTargetBox-2="[139,1084,192,847]" captionTargetBox-3="[140,1085,1260,1501]" captionTargetBox-4="[176,1122,1232,1440]" captionTargetBox-5="[136,1088,940,1442]" captionTargetId-0="figure-211@5.[174,1126,189,537]" captionTargetId-1="figure-276@5.[174,1126,1130,1497]" captionTargetId-2="figure-106@6.[136,1088,189,849]" captionTargetId-3="figure-180@6.[136,1088,1259,1503]" captionTargetId-4="figure-588@7.[174,1126,1230,1441]" captionTargetId-5="figure-241@10.[136,1089,940,1442]" captionTargetPageId-0="5" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="10" captionText-0="Fig. 2. Right thumb viewed from the leading edge of wing of A) Myotis hayesi sp.n. (holo- type, HNHM 2005.82.32.) from Cambodia, B) M. rosseti (HNHM 2869.51.b.1) from Thailand, and C) M. muricola (HNHM 2005.82.20.) from Cambodia. Note the shape of the claw and the relative length of the distal phalanges in relation to the base of the thumb. Scale = 5 mm" captionText-1="Fig. 3. Plantar view of left hindfoot of A = Myotis hayesi sp. n. (holotype, HNHM 2005.82.32.) from Cambodia, B = M. rosseti (HNHM 2869.51.b.2) from Thailand, and C = M. muricola" captionText-2="Fig. 4. Lateral view of skulls of Myotis species: A = M. hayesi sp. n. (holotype, HNHM 2005.82.32.) from Cambodia, B = M. rosseti (holotype, RMNH 19629) from Cambodia, C = M. muricola (HNHM 2005.82.20.) from Cambodia, D = M. ater (HZM 10.34188) from Cam- bodia, E = M. nipalensis (BM(NH) 16.7.29.38) from West Bengal, India, and F = M. alticrania-" captionText-3="Fig. 5. Lateral view of the mandible (scale = 5 mm), and occlusal views of the lower (right) and upper (left) toothrows (scale = 3 mm) of Myotis hayesi sp. n. (holotype, HNHM" captionText-4="Fig. 6. Bacula (dorsal and right lateral views) of selected Myotis species: A = M. hayesi sp. n. (holotype, HNHM 2005.82.32.), B = M. annatessae (Vietnam, ZMMU S-164989), C = M. ater (Vietnam, ZMMU S-172604), D = M. alticraniatus (Vietnam, ZMMU S-167188), E = M. muricola (Vietnam, ZMMU S-172616). B–E (D as M. siligorensis) after KRUSKOP and BORISSENKO" captionText-5="Fig. 7. Landcover in Phnom Penh city in February 2003 (left) and February 2021 (right). The central square symbol represents the available coordinates (11°35’N, 104°55’E) for the holotype of Myotis hayesi sp. n., whereas the dashed line encloses the area this was likely captured within (NW to SE corner: 11°36’N, 104°54’E to 11°34’N, 104°56’E). Imagery from Google-Earth Pro (Google Inc.)" figureDoi-0="http://doi.org/10.5281/zenodo.7160788" figureDoi-1="http://doi.org/10.5281/zenodo.7160792" figureDoi-2="http://doi.org/10.5281/zenodo.7160794" figureDoi-3="http://doi.org/10.5281/zenodo.7160796" figureDoi-4="http://doi.org/10.5281/zenodo.7160802" figureDoi-5="http://doi.org/10.5281/zenodo.7160804" httpUri-0="https://zenodo.org/record/7160788/files/figure.png" httpUri-1="https://zenodo.org/record/7160792/files/figure.png" httpUri-2="https://zenodo.org/record/7160794/files/figure.png" httpUri-3="https://zenodo.org/record/7160796/files/figure.png" httpUri-4="https://zenodo.org/record/7160802/files/figure.png" httpUri-5="https://zenodo.org/record/7160804/files/figure.png" pageId="4" pageNumber="89">Figs 2–7</figureCitation>
)
</heading>
</paragraph>
<paragraph blockId="4.[136,1089,690,1058]" pageId="4" pageNumber="89">
<materialsCitation collectingDate="2000-02-05" collectionCode="HNHM" collectorName="Joe L. Walston" pageId="4" pageNumber="89" specimenCode="HNHM 2005.82" specimenCount="2" typeStatus="holotype">
<typeStatus box="[197,297,690,712]" pageId="4" pageNumber="89">Holotype</typeStatus>
<emphasis box="[305,317,691,712]" italics="true" pageId="4" pageNumber="89">–</emphasis>
<specimenCode box="[325,499,690,712]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="4" pageNumber="89">HNHM 2005.82</specimenCode>
.32. (field number # JLW 02.07), adult male, body in alcohol, skull extracted. Collected by 
<collectorName box="[517,662,719,741]" pageId="4" pageNumber="89">Joe L. Walston</collectorName>
on 
<date box="[703,877,719,741]" pageId="4" pageNumber="89" value="2000-02-05">
<collectingDate box="[703,877,719,741]" pageId="4" pageNumber="89" value="2000-02-05">5 February 2000</collectingDate>
</date>
. Measurements for the 
<typeStatus box="[175,269,748,770]" pageId="4" pageNumber="89">holotype</typeStatus>
(in mm) are as follows: FA = 33.8; thumb length (excluding claw) = 3.2; TAIL = 37.1; HF (excluding claws) = 6.5; longest digit of HF (excluding claw) = 2.2; greatest width of HF (at base of digits) = 3.3; TIB = 15.7; EAR= 12.6; TRAGUS = 5.3; GTL = 13.64; CCL = 12.09; CCW = 3.56; M 
<superScript attach="right" box="[361,368,834,846]" fontSize="5" pageId="4" pageNumber="89">3</superScript>
M 
<superScript attach="both" box="[390,397,834,846]" fontSize="5" pageId="4" pageNumber="89">3</superScript>
W = 5.56; ZYW = 8.41; MAW = 7.13; IOW = 3.50; BCW = 6.53; BCH
</materialsCitation>
</paragraph>
<paragraph blockId="4.[136,1089,690,1058]" box="[136,858,862,890]" pageId="4" pageNumber="89">
= 4.72; CP 
<superScript attach="both" box="[240,247,862,874]" fontSize="5" pageId="4" pageNumber="89">4</superScript>
L = 2.30; CM 
<superScript attach="both" box="[379,386,862,874]" fontSize="5" pageId="4" pageNumber="89">3</superScript>
L = 5.09; ML = 9.93; CM 
<subScript attach="both" box="[634,641,878,890]" fontSize="5" pageId="4" pageNumber="89">3</subScript>
L = 5.45; CPH = 2.92.
</paragraph>
<paragraph blockId="4.[136,1089,690,1058]" pageId="4" pageNumber="89">
<materialsCitation country="Cambodia" latitude="11.583333" location="Phnom Penh" longLatPrecision="1295" longitude="104.916664" pageId="4" pageNumber="89" specimenCount="1" typeStatus="holotype">
Type locality – 
<location LSID="urn:lsid:plazi:treatment:606587D11C00FF8BFE1106693CFDB972:ED13601C1C00FF86FE8107423CB1BBA7" box="[368,509,892,914]" country="Cambodia" latitude="11.583333" longLatPrecision="1295" longitude="104.916664" name="Phnom Penh" pageId="4" pageNumber="89">Phnom Penh</location>
, 
<collectingCountry box="[522,634,892,914]" name="Cambodia" pageId="4" pageNumber="89">Cambodia</collectingCountry>
, 
<geoCoordinate box="[647,731,892,914]" degrees="11" direction="north" minutes="35" orientation="latitude" pageId="4" pageNumber="89" precision="925" value="11.583333">11°35’N</geoCoordinate>
<geoCoordinate box="[740,834,892,914]" degrees="104" direction="east" minutes="55" orientation="longitude" pageId="4" pageNumber="89" precision="925" value="104.916664">104°55’E</geoCoordinate>
. The 
<typeStatus box="[896,990,892,914]" pageId="4" pageNumber="89">holotype</typeStatus>
was collected in an urban area with scattered gardens planted with various species of palms and broad-leaved trees
</materialsCitation>
.
</paragraph>
<paragraph blockId="4.[136,1089,690,1058]" pageId="4" pageNumber="89">
Etymology – Named after Benjamin Hayes, in recognition of his exceptional contri- butions to improving understanding and conservation of bats in 
<collectingCountry box="[828,918,1007,1029]" name="Vietnam" pageId="4" pageNumber="89">Vietnam</collectingCountry>
and 
<collectingCountry box="[972,1085,1007,1029]" name="Cambodia" pageId="4" pageNumber="89">Cambodia</collectingCountry>
. The proposed English name is ‘Hayes’ thick-thumbed myotis’.
</paragraph>
<paragraph blockId="4.[136,1089,1094,1390]" pageId="4" pageNumber="89">
Diagnosis – A small species of 
<taxonomicName box="[586,665,1095,1120]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="4" pageNumber="89" phylum="Chordata" rank="genus">
<emphasis box="[586,665,1095,1120]" italics="true" pageId="4" pageNumber="89">Myotis</emphasis>
</taxonomicName>
(
<tableCitation box="[682,773,1094,1121]" captionStart="Table 2" captionStartId="8.[138,192,406,428]" captionText="Table 2. External and craniodental measurements of selected Myotis species (in mm). Acronyms and definitions for measurements are given in the text. Values are given as mean±SD (for n ≥ 5) (min–max) n." pageId="4" pageNumber="89">Table 2</tableCitation>
) with the forearm length of 
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and a skull length of 
<quantity box="[558,682,1128,1155]" metricMagnitude="-2" metricUnit="m" metricValue="1.364" pageId="4" pageNumber="89" unit="mm" value="13.64">13.64 mm</quantity>
. The thumb, footpad, and calcar (including keel) are distinctly yellowish and contrast sharply with the overall brownish wing membranes. The thumb is shortened and thickened (
<figureCitation box="[956,1023,1195,1222]" captionStart="Fig" captionStartId="5.[174,210,567,589]" captionTargetBox="[176,1123,194,533]" captionTargetId="figure-211@5.[174,1126,189,537]" captionTargetPageId="5" captionText="Fig. 2. Right thumb viewed from the leading edge of wing of A) Myotis hayesi sp.n. (holo- type, HNHM 2005.82.32.) from Cambodia, B) M. rosseti (HNHM 2869.51.b.1) from Thailand, and C) M. muricola (HNHM 2005.82.20.) from Cambodia. Note the shape of the claw and the relative length of the distal phalanges in relation to the base of the thumb. Scale = 5 mm" figureDoi="http://doi.org/10.5281/zenodo.7160788" httpUri="https://zenodo.org/record/7160788/files/figure.png" pageId="4" pageNumber="89">Fig. 2</figureCitation>
). The sole of the hind foot is wide, smooth, and concave and its greatest width (meas- ured at the base of the digits) exceeds the length of the longest digit (
<figureCitation box="[955,1022,1262,1289]" captionStart="Fig" captionStartId="5.[174,210,1524,1546]" captionTargetBox="[174,1122,1132,1497]" captionTargetId="figure-276@5.[174,1126,1130,1497]" captionTargetPageId="5" captionText="Fig. 3. Plantar view of left hindfoot of A = Myotis hayesi sp. n. (holotype, HNHM 2005.82.32.) from Cambodia, B = M. rosseti (HNHM 2869.51.b.2) from Thailand, and C = M. muricola" figureDoi="http://doi.org/10.5281/zenodo.7160792" httpUri="https://zenodo.org/record/7160792/files/figure.png" pageId="4" pageNumber="89">Fig. 3</figureCitation>
). The rostrum is relatively short, and the frontal part of the skull is elevated, although the braincase is not especially globose (
<figureCitation box="[608,677,1330,1356]" captionStart="Fig" captionStartId="6.[136,172,889,911]" captionTargetBox="[139,1084,192,847]" captionTargetId="figure-106@6.[136,1088,189,849]" captionTargetPageId="6" captionText="Fig. 4. Lateral view of skulls of Myotis species: A = M. hayesi sp. n. (holotype, HNHM 2005.82.32.) from Cambodia, B = M. rosseti (holotype, RMNH 19629) from Cambodia, C = M. muricola (HNHM 2005.82.20.) from Cambodia, D = M. ater (HZM 10.34188) from Cam- bodia, E = M. nipalensis (BM(NH) 16.7.29.38) from West Bengal, India, and F = M. alticrania-" figureDoi="http://doi.org/10.5281/zenodo.7160794" httpUri="https://zenodo.org/record/7160794/files/figure.png" pageId="4" pageNumber="89">Fig. 4</figureCitation>
). There are three upper and lower premolars, and the middle premolars are situated within the toothrow (
<figureCitation box="[1003,1072,1363,1390]" captionStart="Fig" captionStartId="6.[136,172,1525,1547]" captionTargetBox="[140,1085,1260,1501]" captionTargetId="figure-180@6.[136,1088,1259,1503]" captionTargetPageId="6" captionText="Fig. 5. Lateral view of the mandible (scale = 5 mm), and occlusal views of the lower (right) and upper (left) toothrows (scale = 3 mm) of Myotis hayesi sp. n. (holotype, HNHM" figureDoi="http://doi.org/10.5281/zenodo.7160796" httpUri="https://zenodo.org/record/7160796/files/figure.png" pageId="4" pageNumber="89">Fig. 5</figureCitation>
).
</paragraph>
<paragraph blockId="4.[136,1089,1425,1591]" lastBlockId="5.[174,1127,718,1086]" lastPageId="5" lastPageNumber="90" pageId="4" pageNumber="89">
Description – This is a small species of 
<taxonomicName box="[632,701,1425,1446]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="4" pageNumber="89" phylum="Chordata" rank="genus">
<emphasis box="[632,701,1425,1446]" italics="true" pageId="4" pageNumber="89">Myotis</emphasis>
</taxonomicName>
, with a forearm length of 
<quantity box="[995,1088,1425,1446]" metricMagnitude="-2" metricUnit="m" metricValue="3.38" pageId="4" pageNumber="89" unit="mm" value="33.8">33.8 mm</quantity>
(
<tableCitation box="[143,224,1453,1475]" captionStart="Table 2" captionStartId="8.[138,192,406,428]" captionText="Table 2. External and craniodental measurements of selected Myotis species (in mm). Acronyms and definitions for measurements are given in the text. Values are given as mean±SD (for n ≥ 5) (min–max) n." pageId="4" pageNumber="89">Table 2</tableCitation>
). The description is based on a wet specimen whose colours might have been influenced by preservation in alcohol. On the dorsal surface, individual hairs in the middorsum are 
<quantity box="[263,383,1511,1533]" metricMagnitude="-3" metricUnit="m" metricValue="6.949999999999999" metricValueMax="7.1" metricValueMin="6.8" pageId="4" pageNumber="89" unit="mm" value="6.949999999999999" valueMax="7.1" valueMin="6.8">6.8–7.1 mm</quantity>
long and are medium brown for most of their length with a lighter tip, but without any obvious banding. The fur extends marginally onto the base of the uropatagium and hindlimbs. On the ventral surface, individual hairs are 5.8–6.0 mm long, medium-brown for approximately 60–70% of their length and then progressively lighten to pale brown in the remainder. Long hairs which are uniformly of the latter colours occur on and adjacent to the genitalia, whereas thin transverse rows of very short, off-white hairs sparsely occur over much of the uropatagium. The overall impression is of medium brown both above and below.
</paragraph>
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<paragraph blockId="5.[174,1127,567,675]" pageId="5" pageNumber="90">
<emphasis bold="true" box="[174,240,567,589]" pageId="5" pageNumber="90">Fig. 2.</emphasis>
Right thumb viewed from the leading edge of wing of A) 
<taxonomicName authority="Csorba &amp; Furey, 2022" authorityName="Csorba &amp; Furey" authorityYear="2022" box="[864,999,567,589]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="5" pageNumber="90" phylum="Chordata" rank="species" species="hayesi" status="sp.n.">
<emphasis box="[864,999,567,589]" italics="true" pageId="5" pageNumber="90">Myotis hayesi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1007,1057,567,588]" pageId="5" pageNumber="90" rank="species">sp.n.</taxonomicNameLabel>
(holotype, HNHM 2005.82.32.) from Cambodia, B) 
<taxonomicName baseAuthorityName="Oey" baseAuthorityYear="1951" box="[648,743,596,617]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="5" pageNumber="90" phylum="Chordata" rank="species" species="rosseti">
<emphasis box="[648,743,596,617]" italics="true" pageId="5" pageNumber="90">M. rosseti</emphasis>
</taxonomicName>
(HNHM 2869.51.b.1) from Thailand, and C) 
<taxonomicName baseAuthorityName="Gray" baseAuthorityYear="1846" box="[254,375,624,646]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="5" pageNumber="90" phylum="Chordata" rank="species" species="muricola">
<emphasis box="[254,375,624,646]" italics="true" pageId="5" pageNumber="90">M. muricola</emphasis>
</taxonomicName>
(HNHM 2005.82.20.) from Cambodia. Note the shape of the claw and the relative length of the distal phalanges in relation to the base of the thumb. Scale = 5 mm
</paragraph>
</caption>
<paragraph blockId="5.[174,1127,718,1086]" lastBlockId="5.[174,1127,1524,1604]" pageId="5" pageNumber="90">
The nostrils are typical for the genus, as are the ears which are 
<quantity box="[891,981,862,884]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="5" pageNumber="90" unit="mm" value="12.6">12.6 mm</quantity>
in length and just reach the tip of the nose when laid forward. The anterior border of each ear is smoothly convex, and the tip is narrowly rounded and darker in colour than the rest of the yellowish pinna. The distal portion of the posterior margin has a small concavity, whereas the mid-portion is essentially straight. The proximal portion is broadly convex but includes a small flap of skin which renders it strongly convex prior to the base. Short transverse cartilaginous ribs occur on the inner surface of the ear conch. The tragus is narrow for most of its length (
<quantity box="[311,390,1064,1085]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="5" pageNumber="90" unit="mm" value="1.7">1.7 mm</quantity>
wide near the base), slightly less than half the height of the pinna and (HNHM 2005.82.20.) from 
<collectingCountry box="[661,774,1582,1604]" name="Cambodia" pageId="5" pageNumber="90">Cambodia</collectingCountry>
. Scale = 
<quantity box="[861,921,1582,1604]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="90" unit="mm" value="5.0">5 mm</quantity>
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7160792" ID-Zenodo-Dep="7160792" httpUri="https://zenodo.org/record/7160792/files/figure.png" pageId="5" pageNumber="90" startId="5.[174,210,1524,1546]" targetBox="[174,1122,1132,1497]" targetPageId="5">
<paragraph blockId="5.[174,1127,1524,1604]" pageId="5" pageNumber="90">
<emphasis bold="true" box="[174,237,1524,1546]" pageId="5" pageNumber="90">Fig. 3.</emphasis>
Plantar view of left hindfoot of A = 
<taxonomicName authority="Csorba &amp; Furey, 2022" authorityName="Csorba &amp; Furey" authorityYear="2022" box="[606,740,1524,1546]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="5" pageNumber="90" phylum="Chordata" rank="species" species="hayesi" status="sp. n.">
<emphasis box="[606,740,1524,1546]" italics="true" pageId="5" pageNumber="90">Myotis hayesi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[744,800,1525,1546]" pageId="5" pageNumber="90" rank="species">sp. n.</taxonomicNameLabel>
(holotype, HNHM 2005.82.32.) from Cambodia, B = 
<taxonomicName baseAuthorityName="Oey" baseAuthorityYear="1951" box="[402,501,1554,1575]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="5" pageNumber="90" phylum="Chordata" rank="species" species="rosseti">
<emphasis box="[402,501,1554,1575]" italics="true" pageId="5" pageNumber="90">M. rosseti</emphasis>
</taxonomicName>
(HNHM 2869.51.b.2) from Thailand, and C = 
<taxonomicName baseAuthorityName="Gray" baseAuthorityYear="1846" box="[1004,1127,1553,1575]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="5" pageNumber="90" phylum="Chordata" rank="species" species="muricola">M. muricola</taxonomicName>
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7160794" ID-Zenodo-Dep="7160794" httpUri="https://zenodo.org/record/7160794/files/figure.png" pageId="6" pageNumber="91" startId="6.[136,172,889,911]" targetBox="[139,1084,192,847]" targetPageId="6">
<paragraph blockId="6.[136,1089,889,1027]" pageId="6" pageNumber="91">
<emphasis bold="true" box="[136,204,889,911]" pageId="6" pageNumber="91">Fig. 4.</emphasis>
Lateral view of skulls of 
<taxonomicName box="[492,649,889,911]" pageId="6" pageNumber="91">
<emphasis box="[492,561,890,911]" italics="true" pageId="6" pageNumber="91">Myotis</emphasis>
species
</taxonomicName>
: A = 
<taxonomicName authority="Csorba &amp; Furey, 2022" authorityName="Csorba &amp; Furey" authorityYear="2022" box="[711,809,889,911]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="hayesi" status="sp. n.">
<emphasis box="[711,809,889,911]" italics="true" pageId="6" pageNumber="91">M. hayesi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[818,879,890,911]" pageId="6" pageNumber="91" rank="species">sp. n.</taxonomicNameLabel>
(holotype, HNHM 2005.82.32.) from Cambodia, B = 
<taxonomicName baseAuthorityName="Oey" baseAuthorityYear="1951" box="[488,587,919,940]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="rosseti">
<emphasis box="[488,587,919,940]" italics="true" pageId="6" pageNumber="91">M. rosseti</emphasis>
</taxonomicName>
(holotype, RMNH 19629) from Cambodia, C = 
<taxonomicName baseAuthorityName="Gray" baseAuthorityYear="1846" box="[136,257,947,969]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="muricola">
<emphasis box="[136,257,947,969]" italics="true" pageId="6" pageNumber="91">M. muricola</emphasis>
</taxonomicName>
(HNHM 2005.82.20.) from Cambodia, D = 
<taxonomicName baseAuthorityName="Peters" baseAuthorityYear="1866" box="[715,787,948,969]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="ater">
<emphasis box="[715,787,948,969]" italics="true" pageId="6" pageNumber="91">M. ater</emphasis>
</taxonomicName>
(HZM 10.34188) from Cambodia, E = 
<taxonomicName authorityName="Dobson" authorityYear="1871" box="[246,377,976,997]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="nipalensis">
<emphasis box="[246,377,976,997]" italics="true" pageId="6" pageNumber="91">M. nipalensis</emphasis>
</taxonomicName>
(BM(NH) 16.7.29.38) from West Bengal, India, and F = M. alticrania-
</paragraph>
</caption>
<paragraph blockId="6.[136,1089,889,1027]" box="[343,881,1004,1027]" pageId="6" pageNumber="91">
<emphasis box="[343,374,1005,1026]" italics="true" pageId="6" pageNumber="91">tus</emphasis>
(MNHG 1956.090) from Lao PDR. Scale = 
<quantity box="[821,881,1004,1026]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="91" unit="mm" value="5.0">5 mm</quantity>
</paragraph>
<paragraph blockId="6.[136,1089,1073,1239]" pageId="6" pageNumber="91">terminates in a pointed tip, which is again darker than the rest of the tragus. Its anterior margin is essentially straight, while the posterior margin is convex and has a small concavity immediately above the base. The face around the eyes is nearly naked, and the skin is light yellowish-brown.</paragraph>
<paragraph blockId="6.[136,1089,1073,1239]" lastBlockId="6.[136,1089,1525,1605]" pageId="6" pageNumber="91">
The wings are uniformly brown in colour. The thumb is light brown above and yellowish below and possesses a fleshy pad on its ventral base (
<figureCitation box="[792,854,1217,1239]" captionStart="Fig" captionStartId="5.[174,210,567,589]" captionTargetBox="[176,1123,194,533]" captionTargetId="figure-211@5.[174,1126,189,537]" captionTargetPageId="5" captionText="Fig. 2. Right thumb viewed from the leading edge of wing of A) Myotis hayesi sp.n. (holo- type, HNHM 2005.82.32.) from Cambodia, B) M. rosseti (HNHM 2869.51.b.1) from Thailand, and C) M. muricola (HNHM 2005.82.20.) from Cambodia. Note the shape of the claw and the relative length of the distal phalanges in relation to the base of the thumb. Scale = 5 mm" figureDoi="http://doi.org/10.5281/zenodo.7160788" httpUri="https://zenodo.org/record/7160788/files/figure.png" pageId="6" pageNumber="91">Fig. 2</figureCitation>
). The hind limbs (TIB 2005.82.32.) from 
<collectingCountry box="[650,760,1583,1605]" name="Cambodia" pageId="6" pageNumber="91">Cambodia</collectingCountry>
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7160796" ID-Zenodo-Dep="7160796" httpUri="https://zenodo.org/record/7160796/files/figure.png" pageId="6" pageNumber="91" startId="6.[136,172,1525,1547]" targetBox="[140,1085,1260,1501]" targetPageId="6">
<paragraph blockId="6.[136,1089,1525,1605]" pageId="6" pageNumber="91">
<emphasis bold="true" box="[136,205,1525,1547]" pageId="6" pageNumber="91">Fig. 5.</emphasis>
Lateral view of the mandible (scale = 5 mm), and occlusal views of the lower (right) and upper (left) toothrows (scale = 3 mm) of 
<taxonomicName authority="Csorba &amp; Furey, 2022" authorityName="Csorba &amp; Furey" authorityYear="2022" box="[686,822,1554,1576]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="6" pageNumber="91" phylum="Chordata" rank="species" species="hayesi" status="sp. n.">
<emphasis box="[686,822,1554,1576]" italics="true" pageId="6" pageNumber="91">Myotis hayesi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[828,885,1555,1576]" pageId="6" pageNumber="91" rank="species">sp. n.</taxonomicNameLabel>
(holotype, HNHM
</paragraph>
</caption>
<paragraph blockId="7.[174,1127,190,1191]" pageId="7" pageNumber="92">
= 
<quantity box="[193,285,190,212]" metricMagnitude="-2" metricUnit="m" metricValue="1.5699999999999998" pageId="7" pageNumber="92" unit="mm" value="15.7">15.7 mm</quantity>
) and feet (HF = 
<quantity box="[460,540,190,212]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="7" pageNumber="92" unit="mm" value="6.5">6.5 mm</quantity>
) are similarly light brown above and yellowish below. Tail length is 
<quantity box="[316,410,219,240]" metricMagnitude="-2" metricUnit="m" metricValue="3.71" pageId="7" pageNumber="92" unit="mm" value="37.1">37.1 mm</quantity>
. The calcar has a distinct and elongated keel and is reasonably well developed, occupying approximately half of the trailing edge of the uropatagium. The plagiopatagium is attached to the base of the first toe. The sole of the hind foot is wide, smooth, and concave, and its greatest width (
<quantity box="[657,737,306,327]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="7" pageNumber="92" unit="mm" value="3.3">3.3 mm</quantity>
) exceeds the length of the digits (
<quantity metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="7" pageNumber="92" unit="mm" value="2.2">2.2 mm</quantity>
excluding the claws) (
<figureCitation box="[454,513,334,356]" captionStart="Fig" captionStartId="5.[174,210,1524,1546]" captionTargetBox="[174,1122,1132,1497]" captionTargetId="figure-276@5.[174,1126,1130,1497]" captionTargetPageId="5" captionText="Fig. 3. Plantar view of left hindfoot of A = Myotis hayesi sp. n. (holotype, HNHM 2005.82.32.) from Cambodia, B = M. rosseti (HNHM 2869.51.b.2) from Thailand, and C = M. muricola" figureDoi="http://doi.org/10.5281/zenodo.7160792" httpUri="https://zenodo.org/record/7160792/files/figure.png" pageId="7" pageNumber="92">Fig. 3</figureCitation>
).
</paragraph>
<paragraph blockId="7.[174,1127,190,1191]" pageId="7" pageNumber="92">
The penis is light yellowish and ca. 
<quantity box="[610,670,363,385]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="92" unit="mm" value="5.0">5 mm</quantity>
long. It slightly widens distally and is gen- erally hairless but possesses scattered, long whitish hairs around the glans.
</paragraph>
<paragraph blockId="7.[174,1127,190,1191]" pageId="7" pageNumber="92">
The skull is small with a greatest skull length (GTL) of 
<quantity box="[848,953,421,443]" metricMagnitude="-2" metricUnit="m" metricValue="1.364" pageId="7" pageNumber="92" unit="mm" value="13.64">13.64 mm</quantity>
and a condylocanine length (CCL) of 
<quantity box="[418,520,450,471]" metricMagnitude="-2" metricUnit="m" metricValue="1.209" pageId="7" pageNumber="92" unit="mm" value="12.09">12.09 mm</quantity>
(
<tableCitation box="[533,611,449,471]" captionStart="Table 2" captionStartId="8.[138,192,406,428]" captionText="Table 2. External and craniodental measurements of selected Myotis species (in mm). Acronyms and definitions for measurements are given in the text. Values are given as mean±SD (for n ≥ 5) (min–max) n." pageId="7" pageNumber="92">Table 2</tableCitation>
). In the lateral profile, the anterior portion of the skull ascends relatively steeply and subsequently very gradually, such that the highest point is attained at the rearmost portion of the braincase (
<figureCitation box="[785,845,507,529]" captionStart="Fig" captionStartId="6.[136,172,889,911]" captionTargetBox="[139,1084,192,847]" captionTargetId="figure-106@6.[136,1088,189,849]" captionTargetPageId="6" captionText="Fig. 4. Lateral view of skulls of Myotis species: A = M. hayesi sp. n. (holotype, HNHM 2005.82.32.) from Cambodia, B = M. rosseti (holotype, RMNH 19629) from Cambodia, C = M. muricola (HNHM 2005.82.20.) from Cambodia, D = M. ater (HZM 10.34188) from Cam- bodia, E = M. nipalensis (BM(NH) 16.7.29.38) from West Bengal, India, and F = M. alticrania-" figureDoi="http://doi.org/10.5281/zenodo.7160794" httpUri="https://zenodo.org/record/7160794/files/figure.png" pageId="7" pageNumber="92">Fig. 4</figureCitation>
). There is no sagittal crest, but the lambdoid crests, although weak, are distinct. The rostrum is relatively short and is not inflated. The posterior border of the palatal emargination extends to the middle of the upper canine. The zygoma are moderately developed and widest posteriorly.
</paragraph>
<paragraph blockId="7.[174,1127,190,1191]" pageId="7" pageNumber="92">
Upper toothrow length (CM 
<superScript attach="both" box="[543,550,622,634]" fontSize="5" pageId="7" pageNumber="92">3</superScript>
L) is 
<quantity box="[606,698,622,644]" metricMagnitude="-3" metricUnit="m" metricValue="5.09" pageId="7" pageNumber="92" unit="mm" value="5.09">5.09 mm</quantity>
(
<tableCitation box="[714,795,622,644]" captionStart="Table 2" captionStartId="8.[138,192,406,428]" captionText="Table 2. External and craniodental measurements of selected Myotis species (in mm). Acronyms and definitions for measurements are given in the text. Values are given as mean±SD (for n ≥ 5) (min–max) n." pageId="7" pageNumber="92">Table 2</tableCitation>
). The first upper incisor (I 
<superScript attach="left" box="[1085,1092,622,634]" fontSize="5" pageId="7" pageNumber="92">2</superScript>
) is bicuspid, with no sign of a third, cingular cusp. Its posterior external face is in contact with the anterior internal face of the second incisor (I 
<superScript attach="left" box="[681,688,679,691]" fontSize="5" pageId="7" pageNumber="92">3</superScript>
), such that I 
<superScript attach="left" box="[819,826,679,691]" fontSize="5" pageId="7" pageNumber="92">2</superScript>
is situated anterior to I 
<superScript attach="left" box="[1073,1080,679,691]" fontSize="5" pageId="7" pageNumber="92">3</superScript>
and clearly visible in the lateral view (
<figureCitation box="[534,595,708,731]" captionStart="Fig" captionStartId="6.[136,172,1525,1547]" captionTargetBox="[140,1085,1260,1501]" captionTargetId="figure-180@6.[136,1088,1259,1503]" captionTargetPageId="6" captionText="Fig. 5. Lateral view of the mandible (scale = 5 mm), and occlusal views of the lower (right) and upper (left) toothrows (scale = 3 mm) of Myotis hayesi sp. n. (holotype, HNHM" figureDoi="http://doi.org/10.5281/zenodo.7160796" httpUri="https://zenodo.org/record/7160796/files/figure.png" pageId="7" pageNumber="92">Fig. 5</figureCitation>
). I 
<superScript attach="left" box="[622,629,708,720]" fontSize="5" pageId="7" pageNumber="92">2</superScript>
possesses a secondary cusp which is posterior to and lower than the principal cusp. I 
<superScript attach="left" box="[576,583,737,749]" fontSize="5" pageId="7" pageNumber="92">3</superScript>
is equal in height to I 
<superScript attach="left" box="[810,817,737,749]" fontSize="5" pageId="7" pageNumber="92">2</superScript>
, slightly larger in crown area (
<figureCitation box="[181,268,766,788]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="6.[136,172,889,911]" captionStartId-1="6.[136,172,1525,1547]" captionTargetBox-0="[139,1084,192,847]" captionTargetBox-1="[140,1085,1260,1501]" captionTargetId-0="figure-106@6.[136,1088,189,849]" captionTargetId-1="figure-180@6.[136,1088,1259,1503]" captionTargetPageId-0="6" captionTargetPageId-1="6" captionText-0="Fig. 4. Lateral view of skulls of Myotis species: A = M. hayesi sp. n. (holotype, HNHM 2005.82.32.) from Cambodia, B = M. rosseti (holotype, RMNH 19629) from Cambodia, C = M. muricola (HNHM 2005.82.20.) from Cambodia, D = M. ater (HZM 10.34188) from Cam- bodia, E = M. nipalensis (BM(NH) 16.7.29.38) from West Bengal, India, and F = M. alticrania-" captionText-1="Fig. 5. Lateral view of the mandible (scale = 5 mm), and occlusal views of the lower (right) and upper (left) toothrows (scale = 3 mm) of Myotis hayesi sp. n. (holotype, HNHM" figureDoi-0="http://doi.org/10.5281/zenodo.7160794" figureDoi-1="http://doi.org/10.5281/zenodo.7160796" httpUri-0="https://zenodo.org/record/7160794/files/figure.png" httpUri-1="https://zenodo.org/record/7160796/files/figure.png" pageId="7" pageNumber="92">Figs 4–5</figureCitation>
) and has a tiny cingular cusp on the labial side. It is separated by a brief diastema from the upper canine (C 
<superScript attach="left" box="[447,454,794,806]" fontSize="5" pageId="7" pageNumber="92">1</superScript>
) which is robust and has a well-defined cingulum. The height of C 
<superScript attach="left" box="[218,225,823,835]" fontSize="5" pageId="7" pageNumber="92">1</superScript>
distinctly exceeds that of the posterior premolar (P 
<superScript attach="left" box="[766,773,823,835]" fontSize="5" pageId="7" pageNumber="92">4</superScript>
), while its crown area is approximately three-quarters. Both the first and second premolars (P 
<superScript attach="left" box="[818,825,852,864]" fontSize="5" pageId="7" pageNumber="92">2</superScript>
, P 
<superScript attach="left" box="[851,858,852,864]" fontSize="5" pageId="7" pageNumber="92">3</superScript>
) are in the toothrow, and P 
<superScript attach="left" box="[188,195,881,893]" fontSize="5" pageId="7" pageNumber="92">3</superScript>
is visible in the lateral view and occupies half of P 
<quantity box="[733,768,881,903]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="7" pageNumber="92" unit="in" value="2.0">
<superScript attach="left" box="[733,740,881,893]" fontSize="5" pageId="7" pageNumber="92">2</superScript>
in
</quantity>
crown area. A faint anterolingual cusp is present on the third premolar (P 
<superScript attach="left" box="[612,619,910,922]" fontSize="5" pageId="7" pageNumber="92">4</superScript>
). The first and second upper molars possess a faint paraconule and paraloph, a metaloph and a well-separated metaconule.
</paragraph>
<paragraph blockId="7.[174,1127,190,1191]" pageId="7" pageNumber="92">
Lower toothrow length (CM 
<subScript attach="both" box="[542,549,983,995]" fontSize="5" pageId="7" pageNumber="92">3</subScript>
L) is 
<quantity box="[604,696,968,990]" metricMagnitude="-3" metricUnit="m" metricValue="5.45" pageId="7" pageNumber="92" unit="mm" value="5.45">5.45 mm</quantity>
(
<tableCitation box="[711,792,968,990]" captionStart="Table 2" captionStartId="8.[138,192,406,428]" captionText="Table 2. External and craniodental measurements of selected Myotis species (in mm). Acronyms and definitions for measurements are given in the text. Values are given as mean±SD (for n ≥ 5) (min–max) n." pageId="7" pageNumber="92">Table 2</tableCitation>
). The lower canine (C 
<subScript attach="left" box="[1032,1039,983,995]" fontSize="5" pageId="7" pageNumber="92">1</subScript>
) has an anterior cingular cusp and a reduced posterior platform. It exceeds the last lower premolar (P4) in height but is slightly smaller in crown area. The middle lower premolar (P3) is small and attains no more than half the height and one-third of crown area of the first premolar (P 
<subScript attach="left" box="[196,203,1098,1110]" fontSize="5" pageId="7" pageNumber="92">2</subScript>
). The lower molars are myotodont. The talonids of the first (M 
<subScript attach="left" box="[858,865,1098,1110]" fontSize="5" pageId="7" pageNumber="92">1</subScript>
) and second (M 
<subScript attach="left" box="[1033,1040,1098,1110]" fontSize="5" pageId="7" pageNumber="92">2</subScript>
) molars are well defined and exceed their respective trigonids in crown area. In the lateral view, the entoconids of both teeth distinctly exceed their respective hypoconids in height. The talonid of the third lower molar (M
</paragraph>
<paragraph box="[547,554,1185,1197]" pageId="7" pageNumber="92">3</paragraph>
<paragraph blockId="7.[174,1127,190,1191]" box="[554,843,1169,1191]" pageId="7" pageNumber="92">) is reduced in comparison.</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7160802" ID-Zenodo-Dep="7160802" httpUri="https://zenodo.org/record/7160802/files/figure.png" pageId="7" pageNumber="92" startId="7.[174,210,1458,1480]" targetBox="[176,1122,1232,1440]" targetPageId="7">
<paragraph blockId="7.[174,1127,1458,1595]" pageId="7" pageNumber="92">
<emphasis bold="true" box="[174,238,1458,1480]" pageId="7" pageNumber="92">Fig. 6.</emphasis>
Bacula (dorsal and right lateral views) of selected 
<taxonomicName box="[768,921,1458,1480]" pageId="7" pageNumber="92">
<emphasis box="[768,837,1459,1480]" italics="true" pageId="7" pageNumber="92">Myotis</emphasis>
species
</taxonomicName>
: A = 
<taxonomicName authority="Csorba &amp; Furey, 2022" authorityName="Csorba &amp; Furey" authorityYear="2022" box="[971,1065,1458,1480]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="7" pageNumber="92" phylum="Chordata" rank="species" species="hayesi" status="sp. n.">
<emphasis box="[971,1065,1458,1480]" italics="true" pageId="7" pageNumber="92">M. hayesi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1070,1126,1459,1480]" pageId="7" pageNumber="92" rank="species">sp. n.</taxonomicNameLabel>
(holotype, HNHM 2005.82.32.), B = 
<taxonomicName authorityName="Kruskop &amp; Borisenko" authorityYear="2013" box="[553,693,1488,1509]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="7" pageNumber="92" phylum="Chordata" rank="species" species="annatessae">
<emphasis box="[553,693,1488,1509]" italics="true" pageId="7" pageNumber="92">M. annatessae</emphasis>
</taxonomicName>
(Vietnam, ZMMU S-164989), C = 
<taxonomicName baseAuthorityName="Peters" baseAuthorityYear="1866" box="[1054,1126,1488,1509]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="7" pageNumber="92" phylum="Chordata" rank="species" species="ater">
<emphasis box="[1054,1126,1488,1509]" italics="true" pageId="7" pageNumber="92">M. ater</emphasis>
</taxonomicName>
(Vietnam, ZMMU S-172604), D = 
<taxonomicName box="[525,683,1516,1538]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="7" pageNumber="92" phylum="Chordata" rank="species" species="alticraniatus">
<emphasis box="[525,683,1516,1538]" italics="true" pageId="7" pageNumber="92">M. alticraniatus</emphasis>
</taxonomicName>
(Vietnam, ZMMU S-167188), E = 
<taxonomicName baseAuthorityName="Gray" baseAuthorityYear="1846" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="7" pageNumber="92" phylum="Chordata" rank="species" species="muricola">
<emphasis italics="true" pageId="7" pageNumber="92">M. muricola</emphasis>
</taxonomicName>
(Vietnam, ZMMU S-172616). B–E (D as 
<taxonomicName baseAuthorityName="Horsfield" baseAuthorityYear="1855" box="[642,787,1545,1566]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="7" pageNumber="92" phylum="Chordata" rank="species" species="siligorensis">
<emphasis box="[642,787,1545,1566]" italics="true" pageId="7" pageNumber="92">M. siligorensis</emphasis>
</taxonomicName>
) after KRUSKOP and BORISSENKO
</paragraph>
</caption>
<footnote box="[523,777,1573,1595]" pageId="7" pageNumber="92">
<paragraph blockId="7.[174,1127,1458,1595]" box="[523,777,1573,1595]" pageId="7" pageNumber="92">(2013). Scale bar = 1 mm</paragraph>
</footnote>
<caption pageId="8" pageNumber="93" startId="8.[138,192,406,428]">
<paragraph blockId="8.[138,1515,406,454]" pageId="8" pageNumber="93">
<emphasis bold="true" box="[138,213,406,428]" pageId="8" pageNumber="93">Table 2.</emphasis>
External and craniodental measurements of selected 
<taxonomicName box="[716,850,406,428]" pageId="8" pageNumber="93">
<emphasis box="[716,778,406,427]" italics="true" pageId="8" pageNumber="93">Myotis</emphasis>
species
</taxonomicName>
(in mm). Acronyms and definitions for measurements are given in the text. Values are given as mean±SD (for 
<emphasis box="[918,930,432,453]" italics="true" pageId="8" pageNumber="93">n</emphasis>
≥ 5) (min–max) 
<emphasis box="[1083,1095,432,453]" italics="true" pageId="8" pageNumber="93">n</emphasis>
.
</paragraph>
</caption>
<paragraph pageId="8" pageNumber="93">
<table box="[136,1506,466,1365]" gridcols="8" gridrows="17" pageId="8" pageNumber="93">
<tr box="[136,1506,466,485]" gridrow="0" pageId="8" pageNumber="93" rowspan-0="1">
<th box="[224,306,466,485]" gridcol="1" gridrow="0" pageId="8" pageNumber="93">
<taxonomicName authorityName="Csorba &amp; Furey" authorityYear="2022" box="[229,302,466,485]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="93" phylum="Chordata" rank="species" species="hayesi">
<emphasis box="[229,302,466,485]" italics="true" pageId="8" pageNumber="93">M. hayesi</emphasis>
</taxonomicName>
</th>
<th box="[341,479,466,485]" gridcol="2" gridrow="0" pageId="8" pageNumber="93">
<taxonomicName baseAuthorityName="Oey" baseAuthorityYear="1951" box="[372,448,466,485]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="93" phylum="Chordata" rank="species" species="rosseti">
<emphasis box="[372,448,466,485]" italics="true" pageId="8" pageNumber="93">M. rosseti</emphasis>
</taxonomicName>
</th>
<th box="[524,662,466,485]" gridcol="3" gridrow="0" pageId="8" pageNumber="93">
<taxonomicName baseAuthorityName="Thomas" baseAuthorityYear="1898" box="[556,632,466,485]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="93" phylum="Chordata" rank="species" species="ridleyi">
<emphasis box="[556,632,466,485]" italics="true" pageId="8" pageNumber="93">M. ridleyi</emphasis>
</taxonomicName>
</th>
<th box="[700,838,466,485]" gridcol="4" gridrow="0" pageId="8" pageNumber="93">
<taxonomicName baseAuthorityName="Gray" baseAuthorityYear="1846" box="[722,816,466,485]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="93" phylum="Chordata" rank="species" species="muricola">
<emphasis box="[722,816,466,485]" italics="true" pageId="8" pageNumber="93">M. muricola</emphasis>
</taxonomicName>
</th>
<th box="[870,1060,466,485]" gridcol="5" gridrow="0" pageId="8" pageNumber="93">
<taxonomicName baseAuthorityName="Peters" baseAuthorityYear="1866" box="[936,992,466,485]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="93" phylum="Chordata" rank="species" species="ater">
<emphasis box="[936,992,466,485]" italics="true" pageId="8" pageNumber="93">M. ater</emphasis>
</taxonomicName>
</th>
<th box="[1102,1240,466,485]" gridcol="6" gridrow="0" pageId="8" pageNumber="93">
<taxonomicName authorityName="Dobson" authorityYear="1871" box="[1120,1222,466,485]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="8" pageNumber="93" phylum="Chordata" rank="species" species="nipalensis">
<emphasis box="[1120,1222,466,485]" italics="true" pageId="8" pageNumber="93">M. nipalensis</emphasis>
</taxonomicName>
</th>
<th box="[1288,1506,466,485]" gridcol="7" gridrow="0" pageId="8" pageNumber="93">
<emphasis box="[1335,1458,466,485]" italics="true" pageId="8" pageNumber="93">M. alticraniatus</emphasis>
</th>
</tr>
<tr box="[136,1506,501,541]" gridrow="1" pageId="8" pageNumber="93">
<th box="[136,201,501,541]" gridcol="0" gridrow="1" pageId="8" pageNumber="93">FA</th>
<td box="[224,306,501,541]" gridcol="1" gridrow="1" pageId="8" pageNumber="93">
33.8 (–) 
<emphasis box="[293,302,512,531]" italics="true" pageId="8" pageNumber="93">1</emphasis>
</td>
<td box="[341,479,501,541]" gridcol="2" gridrow="1" pageId="8" pageNumber="93">
29.3±0.6 (28.5–30.1) 
<emphasis box="[447,466,522,541]" italics="true" pageId="8" pageNumber="93">15</emphasis>
</td>
<td box="[524,662,501,541]" gridcol="3" gridrow="1" pageId="8" pageNumber="93">
29.4 (28.2–30.3) 
<emphasis box="[653,662,512,531]" italics="true" pageId="8" pageNumber="93">5</emphasis>
</td>
<td box="[700,838,501,541]" gridcol="4" gridrow="1" pageId="8" pageNumber="93">
34.1±1.3 (31.7–36.4) 
<emphasis box="[805,824,522,541]" italics="true" pageId="8" pageNumber="93">14</emphasis>
</td>
<td box="[870,1060,501,541]" gridcol="5" gridrow="1" pageId="8" pageNumber="93">
36.9±1.1 (34.8–39.1) 
<emphasis box="[1001,1020,522,541]" italics="true" pageId="8" pageNumber="93">26</emphasis>
</td>
<td box="[1102,1240,501,541]" gridcol="6" gridrow="1" pageId="8" pageNumber="93">
34.9 (–) 
<emphasis box="[1198,1207,512,531]" italics="true" pageId="8" pageNumber="93">1</emphasis>
</td>
<td box="[1288,1506,501,541]" gridcol="7" gridrow="1" pageId="8" pageNumber="93">
33.6±1.3 (31.6–35.8) 
<emphasis box="[1473,1482,512,531]" italics="true" pageId="8" pageNumber="93">9</emphasis>
</td>
</tr>
<tr box="[136,1506,557,598]" gridrow="2" pageId="8" pageNumber="93">
<th box="[136,201,557,598]" gridcol="0" gridrow="2" pageId="8" pageNumber="93">TIB</th>
<td box="[224,306,557,598]" gridcol="1" gridrow="2" pageId="8" pageNumber="93">
15.7 (–) 
<emphasis box="[293,302,568,587]" italics="true" pageId="8" pageNumber="93">1</emphasis>
</td>
<td box="[341,479,557,598]" gridcol="2" gridrow="2" pageId="8" pageNumber="93">
14.2 (13.8–14.6) 
<emphasis box="[470,479,568,587]" italics="true" pageId="8" pageNumber="93">4</emphasis>
</td>
<td box="[524,662,557,598]" gridcol="3" gridrow="2" pageId="8" pageNumber="93">-</td>
<td box="[700,838,557,598]" gridcol="4" gridrow="2" pageId="8" pageNumber="93">
15.9 (15.7–16.0) 
<emphasis box="[829,838,568,587]" italics="true" pageId="8" pageNumber="93">2</emphasis>
</td>
<td box="[870,1060,557,598]" gridcol="5" gridrow="2" pageId="8" pageNumber="93">
15.8±0.8 (14.9–17.4) 
<emphasis box="[1001,1020,579,598]" italics="true" pageId="8" pageNumber="93">19</emphasis>
</td>
<td box="[1102,1240,557,598]" gridcol="6" gridrow="2" pageId="8" pageNumber="93">-</td>
<td box="[1288,1506,557,598]" gridcol="7" gridrow="2" pageId="8" pageNumber="93">
13.6±0.7 (13.1–14.9) 
<emphasis box="[1473,1482,568,587]" italics="true" pageId="8" pageNumber="93">6</emphasis>
</td>
</tr>
<tr box="[136,1506,614,633]" gridrow="3" pageId="8" pageNumber="93">
<th box="[136,201,614,633]" gridcol="0" gridrow="3" pageId="8" pageNumber="93">HF</th>
<td box="[224,306,614,633]" gridcol="1" gridrow="3" pageId="8" pageNumber="93">
6.5 (–) 
<emphasis box="[288,297,614,633]" italics="true" pageId="8" pageNumber="93">1</emphasis>
</td>
<td box="[341,479,614,633]" gridcol="2" gridrow="3" pageId="8" pageNumber="93">
6.9 (6.1–7.5) 
<emphasis box="[456,465,614,633]" italics="true" pageId="8" pageNumber="93">4</emphasis>
</td>
<td box="[524,662,614,633]" gridcol="3" gridrow="3" pageId="8" pageNumber="93">-</td>
<td box="[700,838,614,633]" gridcol="4" gridrow="3" pageId="8" pageNumber="93">
5.5 (5.3–5.7) 
<emphasis box="[815,824,614,633]" italics="true" pageId="8" pageNumber="93">2</emphasis>
</td>
<td box="[870,1060,614,633]" gridcol="5" gridrow="3" pageId="8" pageNumber="93">
7.2±0.5 (6.0–8.0) 
<emphasis box="[1022,1041,614,633]" italics="true" pageId="8" pageNumber="93">19</emphasis>
</td>
<td box="[1102,1240,614,633]" gridcol="6" gridrow="3" pageId="8" pageNumber="93">-</td>
<td box="[1288,1506,614,633]" gridcol="7" gridrow="3" pageId="8" pageNumber="93">
6.7±0.8 (5.0–7.2) 
<emphasis box="[1459,1468,614,633]" italics="true" pageId="8" pageNumber="93">6</emphasis>
</td>
</tr>
<tr box="[136,1506,649,689]" gridrow="4" pageId="8" pageNumber="93">
<th box="[136,201,649,689]" gridcol="0" gridrow="4" pageId="8" pageNumber="93">GTL</th>
<td box="[224,306,649,689]" gridcol="1" gridrow="4" pageId="8" pageNumber="93">
13.64 (–) 
<emphasis box="[297,306,659,678]" italics="true" pageId="8" pageNumber="93">1</emphasis>
</td>
<td box="[341,479,649,689]" gridcol="2" gridrow="4" pageId="8" pageNumber="93">
12.66±0.27 (12.29–12.99) 
<emphasis box="[456,475,670,689]" italics="true" pageId="8" pageNumber="93">12</emphasis>
</td>
<td box="[524,662,649,689]" gridcol="3" gridrow="4" pageId="8" pageNumber="93">
12.40±0.22 (12.07–12.72) 
<emphasis box="[644,653,670,689]" italics="true" pageId="8" pageNumber="93">6</emphasis>
</td>
<td box="[700,838,649,689]" gridcol="4" gridrow="4" pageId="8" pageNumber="93">
13.69±0.30 (13.11–14.31) 
<emphasis box="[815,834,670,689]" italics="true" pageId="8" pageNumber="93">26</emphasis>
</td>
<td box="[870,1060,649,689]" gridcol="5" gridrow="4" pageId="8" pageNumber="93">
14.73±0.30 (14.28–15.41) 
<emphasis box="[1010,1029,670,689]" italics="true" pageId="8" pageNumber="93">35</emphasis>
</td>
<td box="[1102,1240,649,689]" gridcol="6" gridrow="4" pageId="8" pageNumber="93">
13.75±0.30 (13.14–14.10) 
<emphasis box="[1221,1230,670,689]" italics="true" pageId="8" pageNumber="93">9</emphasis>
</td>
<td box="[1288,1506,649,689]" gridcol="7" gridrow="4" pageId="8" pageNumber="93">
12.16±0.29 (11.77–12.68) 
<emphasis box="[1487,1506,659,678]" italics="true" pageId="8" pageNumber="93">11</emphasis>
</td>
</tr>
<tr box="[136,1506,705,745]" gridrow="5" pageId="8" pageNumber="93">
<th box="[136,201,705,745]" gridcol="0" gridrow="5" pageId="8" pageNumber="93">CCL</th>
<td box="[224,306,705,745]" gridcol="1" gridrow="5" pageId="8" pageNumber="93">
12.09 (–) 
<emphasis box="[297,306,716,735]" italics="true" pageId="8" pageNumber="93">1</emphasis>
</td>
<td box="[341,479,705,745]" gridcol="2" gridrow="5" pageId="8" pageNumber="93">
10.87±0.25 (10.42–11.30) 
<emphasis box="[456,475,726,745]" italics="true" pageId="8" pageNumber="93">13</emphasis>
</td>
<td box="[524,662,705,745]" gridcol="3" gridrow="5" pageId="8" pageNumber="93">
10.93±0.14 (10.72–11.14) 
<emphasis box="[644,653,726,745]" italics="true" pageId="8" pageNumber="93">7</emphasis>
</td>
<td box="[700,838,705,745]" gridcol="4" gridrow="5" pageId="8" pageNumber="93">
12.01±0.32 (11.38–12.57) 
<emphasis box="[815,834,726,745]" italics="true" pageId="8" pageNumber="93">26</emphasis>
</td>
<td box="[870,1060,705,745]" gridcol="5" gridrow="5" pageId="8" pageNumber="93">
12.99±0.29 (12.49–13.65) 
<emphasis box="[1010,1029,726,745]" italics="true" pageId="8" pageNumber="93">35</emphasis>
</td>
<td box="[1102,1240,705,745]" gridcol="6" gridrow="5" pageId="8" pageNumber="93">
12.04±0.33 (11.43–12.47) 
<emphasis box="[1217,1236,726,745]" italics="true" pageId="8" pageNumber="93">10</emphasis>
</td>
<td box="[1288,1506,705,745]" gridcol="7" gridrow="5" pageId="8" pageNumber="93">
10.52±0.21 (10.17–10.93) 
<emphasis box="[1487,1506,716,735]" italics="true" pageId="8" pageNumber="93">14</emphasis>
</td>
</tr>
<tr box="[136,1506,761,802]" gridrow="6" pageId="8" pageNumber="93">
<th box="[136,201,761,802]" gridcol="0" gridrow="6" pageId="8" pageNumber="93">CCW</th>
<td box="[224,306,761,802]" gridcol="1" gridrow="6" pageId="8" pageNumber="93">
3.56 (–) 
<emphasis box="[293,302,772,791]" italics="true" pageId="8" pageNumber="93">1</emphasis>
</td>
<td box="[341,479,761,802]" gridcol="2" gridrow="6" pageId="8" pageNumber="93">
3.43±0.13 (3.21–3.60) 
<emphasis box="[447,466,783,802]" italics="true" pageId="8" pageNumber="93">13</emphasis>
</td>
<td box="[524,662,761,802]" gridcol="3" gridrow="6" pageId="8" pageNumber="93">
3.36±0.09 (3.24–3.53) 
<emphasis box="[635,644,783,802]" italics="true" pageId="8" pageNumber="93">7</emphasis>
</td>
<td box="[700,838,761,802]" gridcol="4" gridrow="6" pageId="8" pageNumber="93">
3.48±0.12 (3.23–3.75) 
<emphasis box="[805,824,783,802]" italics="true" pageId="8" pageNumber="93">26</emphasis>
</td>
<td box="[870,1060,761,802]" gridcol="5" gridrow="6" pageId="8" pageNumber="93">
3.99±0.12 (3.79–4.22) 
<emphasis box="[1001,1020,783,802]" italics="true" pageId="8" pageNumber="93">35</emphasis>
</td>
<td box="[1102,1240,761,802]" gridcol="6" gridrow="6" pageId="8" pageNumber="93">
3.55±0.15 (3.32–3.76) 
<emphasis box="[1212,1221,783,802]" italics="true" pageId="8" pageNumber="93">9</emphasis>
</td>
<td box="[1288,1506,761,802]" gridcol="7" gridrow="6" pageId="8" pageNumber="93">
3.00±0.08 (2.88–3.20) 
<emphasis box="[1473,1492,772,791]" italics="true" pageId="8" pageNumber="93">14</emphasis>
</td>
</tr>
<tr box="[136,1506,818,858]" gridrow="7" pageId="8" pageNumber="93">
<th box="[136,201,818,858]" gridcol="0" gridrow="7" pageId="8" pageNumber="93">M3M3W</th>
<td box="[224,306,818,858]" gridcol="1" gridrow="7" pageId="8" pageNumber="93">
5.56 (–) 
<emphasis box="[293,302,828,847]" italics="true" pageId="8" pageNumber="93">1</emphasis>
</td>
<td box="[341,479,818,858]" gridcol="2" gridrow="7" pageId="8" pageNumber="93">
5.17±0.13 (4.91–5.38) 
<emphasis box="[447,466,839,858]" italics="true" pageId="8" pageNumber="93">13</emphasis>
</td>
<td box="[524,662,818,858]" gridcol="3" gridrow="7" pageId="8" pageNumber="93">
5.25±0.22 (4.80–5.44) 
<emphasis box="[635,644,839,858]" italics="true" pageId="8" pageNumber="93">7</emphasis>
</td>
<td box="[700,838,818,858]" gridcol="4" gridrow="7" pageId="8" pageNumber="93">
5.57±0.19 (5.22–5.94) 
<emphasis box="[805,824,839,858]" italics="true" pageId="8" pageNumber="93">26</emphasis>
</td>
<td box="[870,1060,818,858]" gridcol="5" gridrow="7" pageId="8" pageNumber="93">
6.10±0.15 (5.75–6.36) 
<emphasis box="[1001,1020,839,858]" italics="true" pageId="8" pageNumber="93">35</emphasis>
</td>
<td box="[1102,1240,818,858]" gridcol="6" gridrow="7" pageId="8" pageNumber="93">
5.58±0.19 (5.33–5.85) 
<emphasis box="[1207,1226,839,858]" italics="true" pageId="8" pageNumber="93">10</emphasis>
</td>
<td box="[1288,1506,818,858]" gridcol="7" gridrow="7" pageId="8" pageNumber="93">
4.67±0.13 (4.40–4.87) 
<emphasis box="[1473,1492,828,847]" italics="true" pageId="8" pageNumber="93">13</emphasis>
</td>
</tr>
<tr box="[136,1506,874,914]" gridrow="8" pageId="8" pageNumber="93">
<th box="[136,201,874,914]" gridcol="0" gridrow="8" pageId="8" pageNumber="93">ZYW</th>
<td box="[224,306,874,914]" gridcol="1" gridrow="8" pageId="8" pageNumber="93">
8.41 (–) 
<emphasis box="[293,302,885,904]" italics="true" pageId="8" pageNumber="93">1</emphasis>
</td>
<td box="[341,479,874,914]" gridcol="2" gridrow="8" pageId="8" pageNumber="93">
8.52 (8.43–8.61) 
<emphasis box="[470,479,885,904]" italics="true" pageId="8" pageNumber="93">4</emphasis>
</td>
<td box="[524,662,874,914]" gridcol="3" gridrow="8" pageId="8" pageNumber="93">
8.08 (–) 
<emphasis box="[621,630,885,904]" italics="true" pageId="8" pageNumber="93">1</emphasis>
</td>
<td box="[700,838,874,914]" gridcol="4" gridrow="8" pageId="8" pageNumber="93">
8.63±0.31 (8.25–9.13) 
<emphasis box="[810,819,895,914]" italics="true" pageId="8" pageNumber="93">6</emphasis>
</td>
<td box="[870,1060,874,914]" gridcol="5" gridrow="8" pageId="8" pageNumber="93">
9.44±0.19 (8.94–9.71) 
<emphasis box="[1001,1020,895,914]" italics="true" pageId="8" pageNumber="93">26</emphasis>
</td>
<td box="[1102,1240,874,914]" gridcol="6" gridrow="8" pageId="8" pageNumber="93">
7.99 (–) 
<emphasis box="[1198,1207,885,904]" italics="true" pageId="8" pageNumber="93">1</emphasis>
</td>
<td box="[1288,1506,874,914]" gridcol="7" gridrow="8" pageId="8" pageNumber="93">
6.94±0.09 (6.82–7.07) 
<emphasis box="[1477,1486,885,904]" italics="true" pageId="8" pageNumber="93">6</emphasis>
</td>
</tr>
<tr box="[136,1506,930,971]" gridrow="9" pageId="8" pageNumber="93">
<th box="[136,201,930,971]" gridcol="0" gridrow="9" pageId="8" pageNumber="93">MAW</th>
<td box="[224,306,930,971]" gridcol="1" gridrow="9" pageId="8" pageNumber="93">
7.13 (–) 
<emphasis box="[293,302,941,960]" italics="true" pageId="8" pageNumber="93">1</emphasis>
</td>
<td box="[341,479,930,971]" gridcol="2" gridrow="9" pageId="8" pageNumber="93">
6.86±0.18 (6.60–7.13) 
<emphasis box="[447,466,952,971]" italics="true" pageId="8" pageNumber="93">10</emphasis>
</td>
<td box="[524,662,930,971]" gridcol="3" gridrow="9" pageId="8" pageNumber="93">
6.62 (6.59–6.64) 
<emphasis box="[653,662,941,960]" italics="true" pageId="8" pageNumber="93">2</emphasis>
</td>
<td box="[700,838,930,971]" gridcol="4" gridrow="9" pageId="8" pageNumber="93">
7.13±0.19 (6.76–7.31) 
<emphasis box="[810,819,952,971]" italics="true" pageId="8" pageNumber="93">7</emphasis>
</td>
<td box="[870,1060,930,971]" gridcol="5" gridrow="9" pageId="8" pageNumber="93">
7.46±0.13 (7.13–7.72) 
<emphasis box="[1001,1020,952,971]" italics="true" pageId="8" pageNumber="93">27</emphasis>
</td>
<td box="[1102,1240,930,971]" gridcol="6" gridrow="9" pageId="8" pageNumber="93">
6.94 (6.71–7.16) 
<emphasis box="[1231,1240,941,960]" italics="true" pageId="8" pageNumber="93">2</emphasis>
</td>
<td box="[1288,1506,930,971]" gridcol="7" gridrow="9" pageId="8" pageNumber="93">
6.36±0.11 (6.17–6.48) 
<emphasis box="[1473,1492,941,960]" italics="true" pageId="8" pageNumber="93">11</emphasis>
</td>
</tr>
<tr box="[136,1506,987,1027]" gridrow="10" pageId="8" pageNumber="93">
<th box="[136,201,987,1027]" gridcol="0" gridrow="10" pageId="8" pageNumber="93">IOW</th>
<td box="[224,306,987,1027]" gridcol="1" gridrow="10" pageId="8" pageNumber="93">
3.50 (–) 
<emphasis box="[293,302,997,1016]" italics="true" pageId="8" pageNumber="93">1</emphasis>
</td>
<td box="[341,479,987,1027]" gridcol="2" gridrow="10" pageId="8" pageNumber="93">
3.37±0.08 (3.24–3.51) 
<emphasis box="[447,466,1008,1027]" italics="true" pageId="8" pageNumber="93">13</emphasis>
</td>
<td box="[524,662,987,1027]" gridcol="3" gridrow="10" pageId="8" pageNumber="93">
3.02±0.14 (2.89–3.31) 
<emphasis box="[635,644,1008,1027]" italics="true" pageId="8" pageNumber="93">7</emphasis>
</td>
<td box="[700,838,987,1027]" gridcol="4" gridrow="10" pageId="8" pageNumber="93">
3.40±0.11 (3.21–3.57) 
<emphasis box="[805,824,1008,1027]" italics="true" pageId="8" pageNumber="93">17</emphasis>
</td>
<td box="[870,1060,987,1027]" gridcol="5" gridrow="10" pageId="8" pageNumber="93">
3.56±0.10 (3.36–3.74) 
<emphasis box="[1001,1020,1008,1027]" italics="true" pageId="8" pageNumber="93">19</emphasis>
</td>
<td box="[1102,1240,987,1027]" gridcol="6" gridrow="10" pageId="8" pageNumber="93">
3.63±0.19 (3.24–3.81) 
<emphasis box="[1212,1221,1008,1027]" italics="true" pageId="8" pageNumber="93">7</emphasis>
</td>
<td box="[1288,1506,987,1027]" gridcol="7" gridrow="10" pageId="8" pageNumber="93">
2.93±0.10 (2.76–3.06) 
<emphasis box="[1475,1494,997,1016]" italics="true" pageId="8" pageNumber="93">15</emphasis>
</td>
</tr>
<tr box="[136,1506,1043,1083]" gridrow="11" pageId="8" pageNumber="93">
<th box="[136,201,1043,1083]" gridcol="0" gridrow="11" pageId="8" pageNumber="93">BCW</th>
<td box="[224,306,1043,1083]" gridcol="1" gridrow="11" pageId="8" pageNumber="93">
6.53 (–) 
<emphasis box="[293,302,1054,1073]" italics="true" pageId="8" pageNumber="93">1</emphasis>
</td>
<td box="[341,479,1043,1083]" gridcol="2" gridrow="11" pageId="8" pageNumber="93">
6.43±0.13 (6.24–6.60) 
<emphasis box="[447,466,1064,1083]" italics="true" pageId="8" pageNumber="93">10</emphasis>
</td>
<td box="[524,662,1043,1083]" gridcol="3" gridrow="11" pageId="8" pageNumber="93">
6.36 (6.35–6.36) 
<emphasis box="[653,662,1054,1073]" italics="true" pageId="8" pageNumber="93">2</emphasis>
</td>
<td box="[700,838,1043,1083]" gridcol="4" gridrow="11" pageId="8" pageNumber="93">
6.52±0.18 (6.21–6.72) 
<emphasis box="[810,819,1064,1083]" italics="true" pageId="8" pageNumber="93">6</emphasis>
</td>
<td box="[870,1060,1043,1083]" gridcol="5" gridrow="11" pageId="8" pageNumber="93">
6.74±0.20 (6.38–7.05) 
<emphasis box="[1001,1020,1064,1083]" italics="true" pageId="8" pageNumber="93">11</emphasis>
</td>
<td box="[1102,1240,1043,1083]" gridcol="6" gridrow="11" pageId="8" pageNumber="93">
6.68 (6.62–6.73) 
<emphasis box="[1231,1240,1054,1073]" italics="true" pageId="8" pageNumber="93">2</emphasis>
</td>
<td box="[1288,1506,1043,1083]" gridcol="7" gridrow="11" pageId="8" pageNumber="93">
5.92±0.16 (5.70–6.17) 
<emphasis box="[1473,1492,1054,1073]" italics="true" pageId="8" pageNumber="93">12</emphasis>
</td>
</tr>
<tr box="[136,1506,1099,1140]" gridrow="12" pageId="8" pageNumber="93">
<th box="[136,201,1099,1140]" gridcol="0" gridrow="12" pageId="8" pageNumber="93">BCH</th>
<td box="[224,306,1099,1140]" gridcol="1" gridrow="12" pageId="8" pageNumber="93">
4.72 (–) 
<emphasis box="[293,302,1110,1129]" italics="true" pageId="8" pageNumber="93">1</emphasis>
</td>
<td box="[341,479,1099,1140]" gridcol="2" gridrow="12" pageId="8" pageNumber="93">
4.93±0.13 (4.73–5.15) 
<emphasis box="[447,466,1121,1140]" italics="true" pageId="8" pageNumber="93">10</emphasis>
</td>
<td box="[524,662,1099,1140]" gridcol="3" gridrow="12" pageId="8" pageNumber="93">
4.63 (4.62–4.64) 
<emphasis box="[653,662,1110,1129]" italics="true" pageId="8" pageNumber="93">2</emphasis>
</td>
<td box="[700,838,1099,1140]" gridcol="4" gridrow="12" pageId="8" pageNumber="93">
4.89±0.08 (4.73–4.94) 
<emphasis box="[810,819,1121,1140]" italics="true" pageId="8" pageNumber="93">6</emphasis>
</td>
<td box="[870,1060,1099,1140]" gridcol="5" gridrow="12" pageId="8" pageNumber="93">
5.25±0.16 (4.95–5.55) 
<emphasis box="[1001,1020,1121,1140]" italics="true" pageId="8" pageNumber="93">11</emphasis>
</td>
<td box="[1102,1240,1099,1140]" gridcol="6" gridrow="12" pageId="8" pageNumber="93">
4.96 (4.92–5.00) 
<emphasis box="[1231,1240,1110,1129]" italics="true" pageId="8" pageNumber="93">2</emphasis>
</td>
<td box="[1288,1506,1099,1140]" gridcol="7" gridrow="12" pageId="8" pageNumber="93">
4.51±0.17 (4.17–4.78) 
<emphasis box="[1475,1494,1110,1129]" italics="true" pageId="8" pageNumber="93">12</emphasis>
</td>
</tr>
<tr box="[136,1506,1156,1196]" gridrow="13" pageId="8" pageNumber="93">
<th box="[136,201,1156,1196]" gridcol="0" gridrow="13" pageId="8" pageNumber="93">CM3L</th>
<td box="[224,306,1156,1196]" gridcol="1" gridrow="13" pageId="8" pageNumber="93">
5.09 (–) 
<emphasis box="[293,302,1166,1185]" italics="true" pageId="8" pageNumber="93">1</emphasis>
</td>
<td box="[341,479,1156,1196]" gridcol="2" gridrow="13" pageId="8" pageNumber="93">
4.39±0.15 (4.16–4.62) 
<emphasis box="[447,466,1177,1196]" italics="true" pageId="8" pageNumber="93">13</emphasis>
</td>
<td box="[524,662,1156,1196]" gridcol="3" gridrow="13" pageId="8" pageNumber="93">
4.38±0.11 (4.20–4.56) 
<emphasis box="[635,644,1177,1196]" italics="true" pageId="8" pageNumber="93">7</emphasis>
</td>
<td box="[700,838,1156,1196]" gridcol="4" gridrow="13" pageId="8" pageNumber="93">
5.13±0.22 (4.62–5.47) 
<emphasis box="[805,824,1177,1196]" italics="true" pageId="8" pageNumber="93">27</emphasis>
</td>
<td box="[870,1060,1156,1196]" gridcol="5" gridrow="13" pageId="8" pageNumber="93">
5.63±0.14 (5.38–5.94) 
<emphasis box="[1001,1020,1177,1196]" italics="true" pageId="8" pageNumber="93">35</emphasis>
</td>
<td box="[1102,1240,1156,1196]" gridcol="6" gridrow="13" pageId="8" pageNumber="93">
5.05±0.16 (4.86–5.35) 
<emphasis box="[1212,1221,1177,1196]" italics="true" pageId="8" pageNumber="93">9</emphasis>
</td>
<td box="[1288,1506,1156,1196]" gridcol="7" gridrow="13" pageId="8" pageNumber="93">
4.44±0.11 (4.26–4.63) 
<emphasis box="[1473,1492,1166,1185]" italics="true" pageId="8" pageNumber="93">14</emphasis>
</td>
</tr>
<tr box="[136,1506,1212,1252]" gridrow="14" pageId="8" pageNumber="93">
<th box="[136,201,1212,1252]" gridcol="0" gridrow="14" pageId="8" pageNumber="93">ML</th>
<td box="[224,306,1212,1252]" gridcol="1" gridrow="14" pageId="8" pageNumber="93">
9.93 (–) 
<emphasis box="[293,302,1223,1242]" italics="true" pageId="8" pageNumber="93">1</emphasis>
</td>
<td box="[341,479,1212,1252]" gridcol="2" gridrow="14" pageId="8" pageNumber="93">
8.77±0.22 (8.33–9.03) 
<emphasis box="[447,466,1233,1252]" italics="true" pageId="8" pageNumber="93">13</emphasis>
</td>
<td box="[524,662,1212,1252]" gridcol="3" gridrow="14" pageId="8" pageNumber="93">
8.75±0.16 (8.52–8.98) 
<emphasis box="[635,644,1233,1252]" italics="true" pageId="8" pageNumber="93">7</emphasis>
</td>
<td box="[700,838,1212,1252]" gridcol="4" gridrow="14" pageId="8" pageNumber="93">
9.85±0.38 (9.07–10.49) 
<emphasis box="[810,829,1233,1252]" italics="true" pageId="8" pageNumber="93">26</emphasis>
</td>
<td box="[870,1060,1212,1252]" gridcol="5" gridrow="14" pageId="8" pageNumber="93">
10.91±0.26 (10.32–11.75) 
<emphasis box="[1010,1029,1233,1252]" italics="true" pageId="8" pageNumber="93">35</emphasis>
</td>
<td box="[1102,1240,1212,1252]" gridcol="6" gridrow="14" pageId="8" pageNumber="93">
9.85±0.26 (9.29–10.30) 
<emphasis box="[1212,1231,1233,1252]" italics="true" pageId="8" pageNumber="93">10</emphasis>
</td>
<td box="[1288,1506,1212,1252]" gridcol="7" gridrow="14" pageId="8" pageNumber="93">
8.41±0.21 (8.09–8.87) 
<emphasis box="[1477,1486,1223,1242]" italics="true" pageId="8" pageNumber="93">9</emphasis>
</td>
</tr>
<tr box="[136,1506,1268,1309]" gridrow="15" pageId="8" pageNumber="93">
<th box="[136,201,1268,1309]" gridcol="0" gridrow="15" pageId="8" pageNumber="93">CM3L</th>
<td box="[224,306,1268,1309]" gridcol="1" gridrow="15" pageId="8" pageNumber="93">
5.45 (–) 
<emphasis box="[293,302,1279,1298]" italics="true" pageId="8" pageNumber="93">1</emphasis>
</td>
<td box="[341,479,1268,1309]" gridcol="2" gridrow="15" pageId="8" pageNumber="93">
4.65±0.13 (4.42–4.84) 
<emphasis box="[447,466,1290,1309]" italics="true" pageId="8" pageNumber="93">13</emphasis>
</td>
<td box="[524,662,1268,1309]" gridcol="3" gridrow="15" pageId="8" pageNumber="93">
4.62±0.09 (4.47–4.74) 
<emphasis box="[635,644,1290,1309]" italics="true" pageId="8" pageNumber="93">7</emphasis>
</td>
<td box="[700,838,1268,1309]" gridcol="4" gridrow="15" pageId="8" pageNumber="93">
5.47±0.23 (4.95–5.77) 
<emphasis box="[805,824,1290,1309]" italics="true" pageId="8" pageNumber="93">27</emphasis>
</td>
<td box="[870,1060,1268,1309]" gridcol="5" gridrow="15" pageId="8" pageNumber="93">
5.98±0.13 (5.77–6.28) 
<emphasis box="[1041,1060,1279,1298]" italics="true" pageId="8" pageNumber="93">35</emphasis>
</td>
<td box="[1102,1240,1268,1309]" gridcol="6" gridrow="15" pageId="8" pageNumber="93">
5.43±0.22 (5.17–5.78) 
<emphasis box="[1212,1221,1290,1309]" italics="true" pageId="8" pageNumber="93">9</emphasis>
</td>
<td box="[1288,1506,1268,1309]" gridcol="7" gridrow="15" pageId="8" pageNumber="93">
4.61±0.10 (4.45–4.82) 
<emphasis box="[1473,1492,1279,1298]" italics="true" pageId="8" pageNumber="93">13</emphasis>
</td>
</tr>
<tr box="[136,1506,1325,1365]" gridrow="16" pageId="8" pageNumber="93">
<th box="[136,201,1325,1365]" gridcol="0" gridrow="16" pageId="8" pageNumber="93">CPH</th>
<td box="[224,306,1325,1365]" gridcol="1" gridrow="16" pageId="8" pageNumber="93">
2.92 (–) 
<emphasis box="[293,302,1335,1354]" italics="true" pageId="8" pageNumber="93">1</emphasis>
</td>
<td box="[341,479,1325,1365]" gridcol="2" gridrow="16" pageId="8" pageNumber="93">
2.60±0.15 (2.33–2.80) 
<emphasis box="[447,466,1346,1365]" italics="true" pageId="8" pageNumber="93">10</emphasis>
</td>
<td box="[524,662,1325,1365]" gridcol="3" gridrow="16" pageId="8" pageNumber="93">
2.57 (2.47–2.66) 
<emphasis box="[653,662,1335,1354]" italics="true" pageId="8" pageNumber="93">2</emphasis>
</td>
<td box="[700,838,1325,1365]" gridcol="4" gridrow="16" pageId="8" pageNumber="93">
2.90±0.20 (2.63–3.15) 
<emphasis box="[810,819,1346,1365]" italics="true" pageId="8" pageNumber="93">6</emphasis>
</td>
<td box="[870,1060,1325,1365]" gridcol="5" gridrow="16" pageId="8" pageNumber="93">
3.32±0.13 (3.08–3.52) 
<emphasis box="[1041,1060,1335,1354]" italics="true" pageId="8" pageNumber="93">27</emphasis>
</td>
<td box="[1102,1240,1325,1365]" gridcol="6" gridrow="16" pageId="8" pageNumber="93">
2.52 (2.44–2.59) 
<emphasis box="[1231,1240,1335,1354]" italics="true" pageId="8" pageNumber="93">2</emphasis>
</td>
<td box="[1288,1506,1325,1365]" gridcol="7" gridrow="16" pageId="8" pageNumber="93">2.11±0.08 (2.01–2.24) 9</td>
</tr>
</table>
</paragraph>
<paragraph blockId="9.[174,1127,190,327]" pageId="9" pageNumber="94">
The total length of the baculum is 
<quantity box="[602,696,190,212]" metricMagnitude="-4" metricUnit="m" metricValue="9.3" pageId="9" pageNumber="94" unit="mm" value="0.93">0.93 mm</quantity>
. Its basal portion is shallowly lobed and in dorsal outline is almost straight sided for the proximal half, then progressively narrows after the mid-point towards a widely rounded tip. The urethral groove is deep. In the lateral view, the distal portion of the baculum is dorsally directed only very slightly and the tip is not curved (
<figureCitation box="[360,421,305,327]" captionStart="Fig" captionStartId="7.[174,210,1458,1480]" captionTargetBox="[176,1122,1232,1440]" captionTargetId="figure-588@7.[174,1126,1230,1441]" captionTargetPageId="7" captionText="Fig. 6. Bacula (dorsal and right lateral views) of selected Myotis species: A = M. hayesi sp. n. (holotype, HNHM 2005.82.32.), B = M. annatessae (Vietnam, ZMMU S-164989), C = M. ater (Vietnam, ZMMU S-172604), D = M. alticraniatus (Vietnam, ZMMU S-167188), E = M. muricola (Vietnam, ZMMU S-172616). B–E (D as M. siligorensis) after KRUSKOP and BORISSENKO" figureDoi="http://doi.org/10.5281/zenodo.7160802" httpUri="https://zenodo.org/record/7160802/files/figure.png" pageId="9" pageNumber="94">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>