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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1030.62544" ID-GBIF-Dataset="91fec855-7fa9-4ad6-84d7-9b9d35e23e8b" ID-PMC="PMC8060247" ID-Pensoft-Pub="1313-2970-1030-1" ID-Pensoft-UUID="875199397EEE5F7898EA1DB25DA62D25" ID-PubMed="33958904" ModsDocID="1313-2970-1030-1" checkinTime="1618371561543" checkinUser="pensoft" docAuthor="Means, Jackson C., Hennen, Derek A. &amp; Marek, Paul E." docDate="2021" docId="7992210F560656628C8EA9AA8A2A52D0" docLanguage="en" docName="ZooKeys 1030: 1-180" docOrigin="ZooKeys 1030" docPubDate="2021-04-13" docSource="http://dx.doi.org/10.3897/zookeys.1030.62544" docTitle="Nannaria solenas Means &amp; Hennen &amp; Marek 2021, sp. nov." docType="treatment" docVersion="5" id="875199397EEE5F7898EA1DB25DA62D25" lastPageNumber="1" masterDocId="875199397EEE5F7898EA1DB25DA62D25" masterDocTitle="A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae)" masterLastPageNumber="180" masterPageNumber="1" pageNumber="1" updateTime="1673961214978" updateUser="julia">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Means, Jackson C.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7377-0696</mods:nameIdentifier>
<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
<mods:nameIdentifier type="email">mjacks4@vt.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hennen, Derek A.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7005-1151</mods:nameIdentifier>
<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Marek, Paul E.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7048-2514</mods:nameIdentifier>
<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-04-13</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1030</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>180</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1030.62544</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1030.62544</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1030-1</mods:identifier>
<mods:identifier type="Pensoft-UUID">875199397EEE5F7898EA1DB25DA62D25</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="181004733" LSID="urn:lsid:zoobank.org:act:D0A9A027-576C-4BB4-9B51-0EE1178377A7" httpUri="http://treatment.plazi.org/id/7992210F560656628C8EA9AA8A2A52D0" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName ID-CoL="8RVXQ" LSID="http://zoobank.org/D0A9A027-576C-4BB4-9B51-0EE1178377A7" authority="Means &amp; Hennen &amp; Marek, 2021" authorityName="Means &amp; Hennen &amp; Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Nannaria solenas" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="solenas" status="sp. nov.">Nannaria solenas</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 82" captionStartId="F82" captionText="Figure 82. Nannaria solenas sp. nov. holotype ♂ (VTEC, MPE 00128) left gonopod A anterior view; red arrow upturned acropodite tip B medial view C posterior view; red triangle indicates pronounced prefemoral spine. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure82" httpUri="https://binary.pensoft.net/fig/531185" pageId="0" pageNumber="1">Figs 82</figureCitation>
<figureCitation captionStart="Figure 83" captionStartId="F83" captionText="Figure 83. Nannaria solenas sp. nov. non-type ♂ (VTEC, MPE 02420) coloration. Scale bar: 4.0 mm." figureDoi="10.3897/zookeys.1030.62544.figure83" httpUri="https://binary.pensoft.net/fig/531186" pageId="0" pageNumber="1">, 83</figureCitation>
<figureCitation captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Nannaria solenas sp. nov. non-type ♂ (VTEC, MPE 02420) from Crawfish Valley left gonopod; red triangle indicates reduced prefemoral spine." figureDoi="10.3897/zookeys.1030.62544.figure84" httpUri="https://binary.pensoft.net/fig/531187" pageId="0" pageNumber="1">, 84 Vernacular name: &quot;The Pipestem Twisted-Claw Millipede&quot;</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation ID-GBIF-Occurrence="3071355397" accessionNumber="MPE00128" collectingDate="2014-08-21" collectorName="J. Means, E. Francis" country="United States" county="Summers County" elevation="835" latitude="37.5278" location="Athens" longLatPrecision="6" longitude="-80.9889" municipality="Pipestem Resort State Park" specimenCount="♂" stateProvince="West Virginia" typeStatus="Holotype">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">
<typeStatus>Holotype</typeStatus>
</emphasis>
:
<collectingCountry name="United States of America">United States</collectingCountry>
-
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="United States of America" name="West Virginia">West Virginia</collectingRegion>
</emphasis>
•
<specimenCount>♂</specimenCount>
;
<collectingCounty>Summers County</collectingCounty>
,
<collectingMunicipality>Pipestem Resort State Park</collectingMunicipality>
, path to lake, ca.
<quantity metricMagnitude="4" metricUnit="m" metricValue="1.2" unit="km" value="12.0">12 km</quantity>
northeast of
<location LSID="urn:lsid:plazi:treatment:7992210F560656628C8EA9AA8A2A52D0:50177720EAC1A94962B2EAE2C46CD535" country="United States" county="Summers County" latitude="37.5278" longLatPrecision="6" longitude="-80.9889" municipality="Pipestem Resort State Park" name="Athens" stateProvince="West Virginia">Athens</location>
;
<geoCoordinate degrees="37.5278" direction="north" orientation="latitude" precision="5" value="37.5278">37.5278°N</geoCoordinate>
, -
<geoCoordinate degrees="80.9889" direction="west" orientation="longitude" precision="5" value="-80.9889">80.9889°W</geoCoordinate>
;
<quantity metricMagnitude="2" metricUnit="m" metricValue="8.35" unit="m" value="835.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="8.35" unit="m" value="835.0">835 m</elevation>
</quantity>
;
<collectingDate value="2014-08-21">21 Aug. 2014</collectingDate>
; hand collected;
<collectorName>J. Means</collectorName>
,
<collectorName>E. Francis</collectorName>
leg.; VTEC
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE00128">MPE00128</accessionNumber>
.
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation ID-GBIF-Occurrence="3071355627" accessionNumber="MPE00130" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="J. Means &amp; E. Francis" country="United States" county="Summers County" elevation="835" latitude="37.5278" location="West Virginia" longLatPrecision="6" longitude="-80.9889" municipality="Pipestem Resort State Park" specimenCount="4" specimenCount-male="4" stateProvince="West Virginia" typeStatus="Paratypes">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">
<typeStatus>Paratypes</typeStatus>
</emphasis>
:
<collectingCountry name="United States of America">United States</collectingCountry>
-
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="United States of America" name="West Virginia">West Virginia</collectingRegion>
</emphasis>
•
<specimenCount type="male">4 ♂♂</specimenCount>
; same collection data as holotype; VTEC
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE00130">MPE00130</accessionNumber>
</materialsCitation>
, 132-134 •
<materialsCitation ID-GBIF-Occurrence="3071355398" accessionNumber="MPE00135" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="J. Means &amp; E. Francis" country="United States" county="Summers County" elevation="835" latitude="37.5278" location="Athens" longLatPrecision="6" longitude="-80.9889" municipality="Pipestem Resort State Park" specimenCount="4" specimenCount-male="4" stateProvince="West Virginia" typeStatus="paratype">
<specimenCount type="male">4 ♂♂</specimenCount>
; same collection data as holotype; VMNH,
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE00135">MPE00135</accessionNumber>
</materialsCitation>
, 137, 139, 142 •
<materialsCitation ID-GBIF-Occurrence="3071355330" accessionNumber="MPE00129" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="J. Means &amp; E. Francis" country="United States" county="Summers County" elevation="835" latitude="37.5278" location="Athens" longLatPrecision="6" longitude="-80.9889" municipality="Pipestem Resort State Park" specimenCount="3" specimenCount-female="3" stateProvince="West Virginia" typeStatus="paratype">
<specimenCount type="female">3 ♀♀</specimenCount>
; same collection data as holotype; VMNH
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE00129">MPE00129</accessionNumber>
</materialsCitation>
, 131, 136 •
<materialsCitation ID-GBIF-Occurrence="3071355763" accessionNumber="MPE00138" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="J. Means &amp; E. Francis" country="United States" county="Summers County" elevation="835" latitude="37.5278" location="Athens" longLatPrecision="6" longitude="-80.9889" municipality="Pipestem Resort State Park" specimenCount="3" specimenCount-female="3" stateProvince="West Virginia" typeStatus="paratype">
<specimenCount type="female">3 ♀♀</specimenCount>
; same collection data as holotype; VMNH
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE00138">MPE00138</accessionNumber>
</materialsCitation>
, 140, 141.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Other material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation ID-GBIF-Occurrence="3071355438" collectingDate="1967-04-11" collectorName="Herpetology Class" country="United States" latitude="37.1726" location="Hamilton's Cave" longLatPrecision="6" longitude="-80.878" specimenCount="2" specimenCount-male="2" stateProvince="Virginia">
<collectingCountry name="United States of America">United States</collectingCountry>
-
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="United States of America" name="Virginia">Virginia</collectingRegion>
</emphasis>
•
<specimenCount type="male">2 ♂♂</specimenCount>
;
<location LSID="urn:lsid:plazi:treatment:7992210F560656628C8EA9AA8A2A52D0:6BF789A0C061AC48438A09DA48CD4096" country="United States" latitude="37.1726" longLatPrecision="6" longitude="-80.878" name="Hamilton's Cave" stateProvince="Virginia">
<normalizedToken originalValue="Hamilton’s">Hamilton's</normalizedToken>
Cave
</location>
, ca.
<locationDeviation location="Mechanicsburg">
<quantity metricMagnitude="3" metricUnit="m" metricValue="6.437376" unit="mi" value="4.0">4 mi.</quantity>
E Mechanicsburg
</locationDeviation>
;
<geoCoordinate degrees="37.1726" direction="north" orientation="latitude" precision="5" value="37.1726">37.1726°N</geoCoordinate>
, -
<geoCoordinate degrees="80.8780" direction="west" orientation="longitude" precision="5" value="-80.878">80.8780°W</geoCoordinate>
;
<collectingDate value="1967-04-11">11 Apr. 1967</collectingDate>
;
<collectorName>Herpetology Class</collectorName>
leg.; VMNH NAN0151
</materialsCitation>
•
<materialsCitation ID-GBIF-Occurrence="3071355362" collectingDate="1956-04-29" collectorName="R. Hoffman" country="United States" latitude="37.1726" location="Hamilton's Cave" longLatPrecision="6" longitude="-80.878" specimenCount="1" specimenCount-male="1" stateProvince="Virginia">
<specimenCount type="male">1 ♂</specimenCount>
; same collection data as preceding;
<collectingDate value="1956-04-29">29 Apr. 1956</collectingDate>
;
<collectorName>R. Hoffman</collectorName>
leg.; VMNH NAN0177
</materialsCitation>
•
<materialsCitation ID-GBIF-Occurrence="3071355593" collectingDate="1967-09-20" collectorName="Knight, Rushin, Liscombe, R. Hoffman" country="United States" county="Bland County" latitude="37.1834" location="Hamilton's Cave" longLatPrecision="6" longitude="-80.8624" specimenCount="19" specimenCount-male="19" stateProvince="Virginia">
<specimenCount type="male">19 ♂♂</specimenCount>
;
<collectingCounty>Bland County</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:7992210F560656628C8EA9AA8A2A52D0:BECE19259A04ECC1B80DEDF97D063DEA" country="United States" county="Bland County" latitude="37.1834" longLatPrecision="6" longitude="-80.8624" name="Hamilton's Cave" stateProvince="Virginia">
<normalizedToken originalValue="Hamilton’s">Hamilton's</normalizedToken>
Cave
</location>
,
<locationDeviation location="Mechanicsburg">
<quantity metricMagnitude="3" metricUnit="m" metricValue="8.04672" unit="mi" value="5.0">5 mi.</quantity>
ENE of Mechanicsburg
</locationDeviation>
;
<geoCoordinate degrees="37.1834" direction="north" orientation="latitude" precision="5" value="37.1834">37.1834°N</geoCoordinate>
, -
<geoCoordinate degrees="80.8624" direction="west" orientation="longitude" precision="5" value="-80.8624">80.8624°W</geoCoordinate>
;
<collectingDate value="1967-09-20">20 Sep. 1967</collectingDate>
;
<collectorName>Knight</collectorName>
,
<collectorName>Rushin</collectorName>
,
<collectorName>Liscombe</collectorName>
,
<collectorName>R. Hoffman</collectorName>
leg.; VMNH NAN0322
</materialsCitation>
•
<materialsCitation ID-GBIF-Occurrence="3071355604" collectingDate="1980-05-16" collectorName="R. Hoffman" country="United States" county="Bland County" latitude="37.1468" location="Mechanicsville" longLatPrecision="6" longitude="-80.8727" municipality="Hamilton's Cave" specimenCount="5" specimenCount-male="5" stateProvince="Virginia">
<specimenCount type="male">5 ♂♂</specimenCount>
;
<collectingCounty>Bland County</collectingCounty>
, hillside outside
<collectingMunicipality>
<normalizedToken originalValue="Hamilton’s">Hamilton's</normalizedToken>
Cave
</collectingMunicipality>
, ca.
<quantity metricMagnitude="3" metricUnit="m" metricValue="6.0" unit="km" value="6.0">6 km</quantity>
east of
<location LSID="urn:lsid:plazi:treatment:7992210F560656628C8EA9AA8A2A52D0:87CE1F7141D52BAC37DC343456A53238" country="United States" county="Bland County" latitude="37.1468" longLatPrecision="6" longitude="-80.8727" municipality="Hamilton's Cave" name="Mechanicsville" stateProvince="Virginia">Mechanicsville</location>
(presumably a typo of
<location LSID="urn:lsid:plazi:treatment:7992210F560656628C8EA9AA8A2A52D0:4B61643C6AAAD8171018B79D07DD293F" country="United States" county="Bland County" latitude="37.1468" longLatPrecision="6" longitude="-80.8727" municipality="Hamilton's Cave" name="Mechanicsburg" stateProvince="Virginia">Mechanicsburg</location>
);
<geoCoordinate degrees="37.1468" direction="north" orientation="latitude" precision="5" value="37.1468">37.1468°N</geoCoordinate>
, -
<geoCoordinate degrees="80.8727" direction="west" orientation="longitude" precision="5" value="-80.8727">80.8727°W</geoCoordinate>
;
<collectingDate value="1980-05-16">16 May 1980</collectingDate>
;
<collectorName>R. Hoffman</collectorName>
leg.; VMNH NAN0323
</materialsCitation>
•
<materialsCitation ID-GBIF-Occurrence="3071355393" collectingDate="1978-10-21" collectorName="R. Hoffman" country="United States" county="Bland County" latitude="37.1307" location="Mechanicsburg" longLatPrecision="6" longitude="-80.8916" municipality="Big Walker Mountain" specimenCount="10" specimenCount-male="10" stateProvince="Virginia">
<specimenCount type="male">10 ♂♂</specimenCount>
;
<collectingCounty>Bland County</collectingCounty>
,
<collectingMunicipality>Big Walker Mountain</collectingMunicipality>
, on
<quantity metricMagnitude="3" metricUnit="m" metricValue="4.828032" unit="mi" value="3.0">3 mi</quantity>
/
<locationDeviation location="Mechanicsville">SE of Mechanicsville</locationDeviation>
(presumably a typo of
<location LSID="urn:lsid:plazi:treatment:7992210F560656628C8EA9AA8A2A52D0:3168D173F105BE1FCB3422BC9BAFB556" country="United States" county="Bland County" latitude="37.1307" longLatPrecision="6" longitude="-80.8916" municipality="Big Walker Mountain" name="Mechanicsburg" stateProvince="Virginia">Mechanicsburg</location>
);
<geoCoordinate degrees="37.1307" direction="north" orientation="latitude" precision="5" value="37.1307">37.1307°N</geoCoordinate>
, -
<geoCoordinate degrees="80.8916" direction="west" orientation="longitude" precision="5" value="-80.8916">80.8916°W</geoCoordinate>
;
<collectingDate value="1978-10-21">21 Oct. 1978</collectingDate>
;
<collectorName>R. Hoffman</collectorName>
leg.; VMNH NAN0220
</materialsCitation>
•
<materialsCitation ID-GBIF-Occurrence="3071355519" collectingDate="2006-08-20" collectorName="P. Marek, C. Spruill" country="United States" county="Russell County" elevation="625" latitude="36.9533" location="Pinnacles Nature Preserve" longLatPrecision="6" longitude="-82.055" specimenCount="10" specimenCount-male="10" stateProvince="Virginia">
<specimenCount type="male">10 ♂</specimenCount>
;
<collectingCounty>Russell County</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:7992210F560656628C8EA9AA8A2A52D0:0B3114664808C842BF1B15286183E16A" country="United States" county="Russell County" latitude="36.9533" longLatPrecision="6" longitude="-82.055" name="Pinnacles Nature Preserve" stateProvince="Virginia">Pinnacles Nature Preserve</location>
, nr preserve tr.;
<geoCoordinate degrees="36.9533" direction="north" orientation="latitude" precision="5" value="36.9533">36.9533°N</geoCoordinate>
, -
<geoCoordinate degrees="82.0550" direction="west" orientation="longitude" precision="5" value="-82.055">82.0550°W</geoCoordinate>
; elev.
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.25" unit="m" value="625.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.25" unit="m" value="625.0">625 m</elevation>
</quantity>
;
<collectingDate value="2006-08-20">20 Aug. 2006</collectingDate>
; hand collected;
<collectorName>P. Marek</collectorName>
,
<collectorName>C. Spruill</collectorName>
leg.; VTEC MMC0201, 203-207, 209, 212, 214, 215
</materialsCitation>
•
<materialsCitation ID-GBIF-Occurrence="3071355766" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="P. Marek &amp; C. Spruill" country="United States" county="Russell County" elevation="625" latitude="36.9533" location="Pinnacles Nature Preserve" longLatPrecision="6" longitude="-82.055" specimenCount="5" specimenCount-female="5" stateProvince="Virginia">
<specimenCount type="female">5 ♀♀</specimenCount>
; same collection data as preceding; VTEC MMC0200, 208, 210, 211, 213
</materialsCitation>
•
<materialsCitation ID-GBIF-Occurrence="3071355722" collectingDate="1971-03-29" collectorName="C. Chapman" country="United States" county="Tazewell County" latitude="37.206" location="Cove Creek" longLatPrecision="6" longitude="-81.307" municipality="East River Mtn." specimenCount="1" specimenCount-male="1" stateProvince="Virginia">
<specimenCount type="male">1 ♂</specimenCount>
;
<collectingCounty>Tazewell County</collectingCounty>
, SE slope
<collectingMunicipality>East River Mtn.</collectingMunicipality>
, near
<location LSID="urn:lsid:plazi:treatment:7992210F560656628C8EA9AA8A2A52D0:DE357B09BE3C7B3A85E07DE64EF5D5AA" country="United States" county="Tazewell County" latitude="37.206" longLatPrecision="6" longitude="-81.307" municipality="East River Mtn." name="Cove Creek" stateProvince="Virginia">Cove Creek</location>
;
<geoCoordinate degrees="37.2060" direction="north" orientation="latitude" precision="5" value="37.206">37.2060°N</geoCoordinate>
, -
<geoCoordinate degrees="81.3070" direction="west" orientation="longitude" precision="5" value="-81.307">81.3070°W</geoCoordinate>
;
<collectingDate value="1971-03-29">29 Mar. 1971</collectingDate>
;
<collectorName>C. Chapman</collectorName>
leg.; VMNH NAN0049
</materialsCitation>
•
<materialsCitation ID-GBIF-Occurrence="3071355740" collectingDate="1971-02-19" collectorName="W. Shear" country="United States" county="Tazewell County" elevation="1219" latitude="37.0114" location="Beartown Mtn." longLatPrecision="6" longitude="-81.699" municipality="Burkes Garden" specimenCount="3" specimenCount-male="3" stateProvince="Virginia">
<specimenCount type="male">3 ♂♂</specimenCount>
;
<collectingCounty>Tazewell County</collectingCounty>
,
<collectingMunicipality>Burkes Garden</collectingMunicipality>
, e. slope of
<location LSID="urn:lsid:plazi:treatment:7992210F560656628C8EA9AA8A2A52D0:C74EF7F098AC750C0A37E1D448989E25" country="United States" county="Tazewell County" latitude="37.0114" longLatPrecision="6" longitude="-81.699" municipality="Burkes Garden" name="Beartown Mtn." stateProvince="Virginia">Beartown Mtn.</location>
;
<geoCoordinate degrees="37.0114" direction="north" orientation="latitude" precision="5" value="37.0114">37.0114°N</geoCoordinate>
, -
<geoCoordinate degrees="81.6990" direction="west" orientation="longitude" precision="5" value="-81.699">81.6990°W</geoCoordinate>
; elev.
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.219" unit="m" value="1219.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.219" unit="m" value="1219.0">1219 m</elevation>
</quantity>
;
<collectingDate value="1971-02-19">19 Feb. 1971</collectingDate>
;
<collectorName>W. Shear</collectorName>
leg.; VMNH NAN0328
</materialsCitation>
•
<materialsCitation ID-GBIF-Occurrence="3071355372" accessionNumber="MPE02416" collectingDate="2017-03-24" collectorName="C. Harden" country="United States" county="Wythe County" elevation="772" latitude="36.9526" location="Channel Rock Hollow" longLatPrecision="6" longitude="-81.3247" municipality="Crawfish Valley" specimenCount="1" specimenCount-male="1" stateProvince="Virginia">
<specimenCount type="male">1 ♂</specimenCount>
;
<collectingCounty>Wythe County</collectingCounty>
,
<collectingMunicipality>Crawfish Valley</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:7992210F560656628C8EA9AA8A2A52D0:C073803D999240DE0E039AD5DB1AC44D" country="United States" county="Wythe County" latitude="36.9526" longLatPrecision="6" longitude="-81.3247" municipality="Crawfish Valley" name="Channel Rock Hollow" stateProvince="Virginia">Channel Rock Hollow</location>
trail
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.609344" unit="mi" value="1.0">1 mile</quantity>
from
<location LSID="urn:lsid:plazi:treatment:7992210F560656628C8EA9AA8A2A52D0:97792DB0DDDD07AE7B472499BC3D7DFA" country="United States" county="Wythe County" latitude="36.9526" longLatPrecision="6" longitude="-81.3247" municipality="Crawfish Valley" name="Strawberry Rd." stateProvince="Virginia">Strawberry Rd.</location>
end;
<geoCoordinate degrees="36.9526" direction="north" orientation="latitude" precision="5" value="36.9526">36.9526°N</geoCoordinate>
, -
<geoCoordinate degrees="81.3247" direction="west" orientation="longitude" precision="5" value="-81.3247">81.3247°W</geoCoordinate>
; elev.
<quantity metricMagnitude="2" metricUnit="m" metricValue="7.72" unit="m" value="772.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="7.72" unit="m" value="772.0">772 m</elevation>
</quantity>
;
<collectingDate value="2017-03-24">24 Mar. 2017</collectingDate>
; hand collected;
<collectorName>C. Harden</collectorName>
leg.; VTEC
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE02416">MPE02416</accessionNumber>
</materialsCitation>
•
<materialsCitation ID-GBIF-Occurrence="3071355515" accessionNumber="MPE02428" collectingDate="2017-03-24" collectorName="C. Harden" country="United States" county="Wythe County" elevation="770" latitude="36.9585" location="Channel Rock Hollow" longLatPrecision="6" longitude="-81.3189" municipality="Crawfish Valley" specimenCount="1" specimenCount-male="1" stateProvince="Virginia">
<specimenCount type="male">1 ♂</specimenCount>
;
<collectingCounty>Wythe County</collectingCounty>
,
<collectingMunicipality>Crawfish Valley</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:7992210F560656628C8EA9AA8A2A52D0:1FE0950E471C6895BFA62F36102FFEF2" country="United States" county="Wythe County" latitude="36.9585" longLatPrecision="6" longitude="-81.3189" municipality="Crawfish Valley" name="Channel Rock Hollow" stateProvince="Virginia">Channel Rock Hollow</location>
trail
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.414016" unit="mi" value="1.5">1.5 mile</quantity>
from
<location LSID="urn:lsid:plazi:treatment:7992210F560656628C8EA9AA8A2A52D0:AD94EB0FFBC37E5C6555A3DEDE9D95B0" country="United States" county="Wythe County" latitude="36.9585" longLatPrecision="6" longitude="-81.3189" municipality="Crawfish Valley" name="Strawberry Rd." stateProvince="Virginia">Strawberry Rd.</location>
end;
<geoCoordinate degrees="36.9585" direction="north" orientation="latitude" precision="5" value="36.9585">36.9585°N</geoCoordinate>
, -
<geoCoordinate degrees="81.3189" direction="west" orientation="longitude" precision="5" value="-81.3189">81.3189°W</geoCoordinate>
; elev.
<quantity metricMagnitude="2" metricUnit="m" metricValue="7.7" unit="m" value="770.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="7.7" unit="m" value="770.0">770 m</elevation>
</quantity>
;
<collectingDate value="2017-03-24">24 Mar. 2017</collectingDate>
; hand collected;
<collectorName>C. Harden</collectorName>
leg.; VTEC
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/MPE02428">MPE02428</accessionNumber>
</materialsCitation>
• SCAU -
<materialsCitation ID-GBIF-Occurrence="3071355588" collectingDate="1989-10-04" collectorName="W. Arnold" country="United States" county="Fayette County" latitude="38.0672" location="Fayette Station" longLatPrecision="6" longitude="-81.0834" specimenCount="1" specimenCount-male="1" stateProvince="West Virginia">
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="United States of America" name="West Virginia">West Virginia</collectingRegion>
</emphasis>
•
<specimenCount type="male">1 ♂</specimenCount>
;
<collectingCounty>Fayette County</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:7992210F560656628C8EA9AA8A2A52D0:7CE1E652DFF663FA3528E8D28BA00698" country="United States" county="Fayette County" latitude="38.0672" longLatPrecision="6" longitude="-81.0834" name="Fayette Station" stateProvince="West Virginia">Fayette Station</location>
;
<geoCoordinate degrees="38.0672" direction="north" orientation="latitude" precision="5" value="38.0672">38.0672°N</geoCoordinate>
, -
<geoCoordinate degrees="81.0834" direction="west" orientation="longitude" precision="5" value="-81.0834">81.0834°W</geoCoordinate>
;
<collectingDate value="1989-10-04">4 Oct. 1989</collectingDate>
;
<collectorName>W. Arnold</collectorName>
leg.; NCSM NAN0455
</materialsCitation>
•
<materialsCitation ID-GBIF-Occurrence="3071355742" collectingDate="1966-10-02" collectorName="W. Shear, For, Suppl." country="United States" county="Mercer County" latitude="37.4267" location="Jackson's Park" longLatPrecision="6" longitude="-81.0402" municipality="Athens" specimenCount="1" specimenCount-male="1" stateProvince="West Virginia">
<specimenCount type="male">1 ♂</specimenCount>
;
<collectingCounty>Mercer County</collectingCounty>
,
<collectingMunicipality>Athens</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:7992210F560656628C8EA9AA8A2A52D0:41A956669D783219D6F71DDF403E409E" country="United States" county="Mercer County" latitude="37.4267" longLatPrecision="6" longitude="-81.0402" municipality="Athens" name="Jackson's Park" stateProvince="West Virginia">
<normalizedToken originalValue="Jackson’s">Jackson's</normalizedToken>
Park
</location>
,
<location LSID="urn:lsid:plazi:treatment:7992210F560656628C8EA9AA8A2A52D0:8AC1954C18203C1213CA600D9D1E024E" country="United States" county="Mercer County" latitude="37.4267" longLatPrecision="6" longitude="-81.0402" municipality="Athens" name="Unity Rd." stateProvince="West Virginia">Unity Rd.</location>
;
<geoCoordinate degrees="37.4267" direction="north" orientation="latitude" precision="5" value="37.4267">37.4267°N</geoCoordinate>
, -
<geoCoordinate degrees="81.0402" direction="west" orientation="longitude" precision="5" value="-81.0402">81.0402°W</geoCoordinate>
;
<collectingDate value="1966-10-02">2 Oct. 1966</collectingDate>
;
<collectorName>W. Shear</collectorName>
leg.; VMNH NAN0031.
<collectorName>For</collectorName>
detailed collection data see
<collectorName>Suppl.</collectorName>
material 7
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
Adult males of
<taxonomicName authorityName="Means &amp; Hennen &amp; Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria solenas" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="solenas">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria solenas</emphasis>
</taxonomicName>
sp. nov. are distinct from other
<taxonomicName authorityName="Means &amp; Hennen &amp; Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria</emphasis>
</taxonomicName>
and the sympatric
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. asta" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="asta">
<emphasis italics="true" pageId="0" pageNumber="1">N. asta</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. aenigma" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="aenigma">
<emphasis italics="true" pageId="0" pageNumber="1">N. aenigma</emphasis>
</taxonomicName>
, based on the following combination of characters:
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Gonopods</emphasis>
.
</emphasis>
Gonopodal acropodite straight, not gently curving throughout as in
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. asta" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="asta">
<emphasis italics="true" pageId="0" pageNumber="1">N. asta</emphasis>
</taxonomicName>
sp. nov. or
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. aenigma" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="aenigma">
<emphasis italics="true" pageId="0" pageNumber="1">N. aenigma</emphasis>
</taxonomicName>
. Distal zone short, rectangular, bent medially at 90° angle with acropodite with slight cephalically-directed upturn at terminal edge (Fig.
<figureCitation captionStart="Figure 82" captionStartId="F82" captionText="Figure 82. Nannaria solenas sp. nov. holotype ♂ (VTEC, MPE 00128) left gonopod A anterior view; red arrow upturned acropodite tip B medial view C posterior view; red triangle indicates pronounced prefemoral spine. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure82" httpUri="https://binary.pensoft.net/fig/531185" pageId="0" pageNumber="1">82A</figureCitation>
, red arrow), not rounded, directed caudally with lateral flange as in
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. asta" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="asta">
<emphasis italics="true" pageId="0" pageNumber="1">N. asta</emphasis>
</taxonomicName>
sp. nov., or sinuous as in
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. aenigma" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="aenigma">
<emphasis italics="true" pageId="0" pageNumber="1">N. aenigma</emphasis>
</taxonomicName>
. Acropodite simple with slight swelling on inner margin but lacking dimple on outer margin as in
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. asta" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="asta">
<emphasis italics="true" pageId="0" pageNumber="1">N. asta</emphasis>
</taxonomicName>
sp. nov., or hooked lateral flange as in
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. aenigma" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="aenigma">
<emphasis italics="true" pageId="0" pageNumber="1">N. aenigma</emphasis>
</taxonomicName>
. Prefemur with long, acicular prefemoral process, not stout as in
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. asta" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="asta">
<emphasis italics="true" pageId="0" pageNumber="1">N. asta</emphasis>
</taxonomicName>
sp. nov., or laterally curving as in
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. aenigma" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="aenigma">
<emphasis italics="true" pageId="0" pageNumber="1">N. aenigma</emphasis>
</taxonomicName>
. Prefemoral spine pronounced and tooth-like (Fig.
<figureCitation captionStart="Figure 82" captionStartId="F82" captionText="Figure 82. Nannaria solenas sp. nov. holotype ♂ (VTEC, MPE 00128) left gonopod A anterior view; red arrow upturned acropodite tip B medial view C posterior view; red triangle indicates pronounced prefemoral spine. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure82" httpUri="https://binary.pensoft.net/fig/531185" pageId="0" pageNumber="1">82C</figureCitation>
, red triangle), not cephalically-curving as in
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. asta" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="asta">
<emphasis italics="true" pageId="0" pageNumber="1">N. asta</emphasis>
</taxonomicName>
sp. nov. or lacking as in
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. aenigma" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="aenigma">
<emphasis italics="true" pageId="0" pageNumber="1">N. aenigma</emphasis>
</taxonomicName>
. Telopodite basal zone simple, ca. 1/2 length of acropodite, not with lateral bulge, &lt;
<normalizedToken originalValue="½">1/2</normalizedToken>
length of acropodite as in
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. asta" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="asta">
<emphasis italics="true" pageId="0" pageNumber="1">N. asta</emphasis>
</taxonomicName>
sp. nov., or &lt;1/6 length of acropodite as in
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. aenigma" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="aenigma">
<emphasis italics="true" pageId="0" pageNumber="1">N. aenigma</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Color</emphasis>
.
</emphasis>
Tergites with orange paranotal spots (Fig.
<figureCitation captionStart="Figure 83" captionStartId="F83" captionText="Figure 83. Nannaria solenas sp. nov. non-type ♂ (VTEC, MPE 02420) coloration. Scale bar: 4.0 mm." figureDoi="10.3897/zookeys.1030.62544.figure83" httpUri="https://binary.pensoft.net/fig/531186" pageId="0" pageNumber="1">83</figureCitation>
). Black background. Dorsum of collum smooth with orange and white margin.
</paragraph>
<caption doi="10.3897/zookeys.1030.62544.figure82" httpUri="https://binary.pensoft.net/fig/531185" pageId="0" pageNumber="1" start="Figure 82" startId="F82">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 82.</emphasis>
<taxonomicName authorityName="Means &amp; Hennen &amp; Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria solenas" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="solenas">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria solenas</emphasis>
</taxonomicName>
sp. nov. holotype ♂ (VTEC, MPE00128) left gonopod
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
anterior view; red arrow upturned acropodite tip
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
medial view
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
posterior view; red triangle indicates pronounced prefemoral spine. Scale bar: 0.5 mm.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1030.62544.figure83" httpUri="https://binary.pensoft.net/fig/531186" pageId="0" pageNumber="1" start="Figure 83" startId="F83">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 83.</emphasis>
<taxonomicName authorityName="Means &amp; Hennen &amp; Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria solenas" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="solenas">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria solenas</emphasis>
</taxonomicName>
sp. nov. non-type ♂ (VTEC, MPE02420) coloration. Scale bar: 4.0 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Measurements.</paragraph>
<paragraph pageId="0" pageNumber="1">♂ holotype (VTEC, MPE00128): BL = 30.8, CW = 4.1, IW = 2.1, ISW = 1.0, B11W = 5.0, B11H = 3.3; ♀ paratype (VMNH, MPE00140): BL = 34.7, CW = 4.3, IW = 2.5, ISW = 1.1, B11W = 5.5, B11H = 3.1.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="variation">
<paragraph pageId="0" pageNumber="1">Variation.</paragraph>
<paragraph pageId="0" pageNumber="1">
Individuals from Crawfish Valley, Wythe County, Virginia, have reduced, shelf-like prefemoral spines (Fig.
<figureCitation captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Nannaria solenas sp. nov. non-type ♂ (VTEC, MPE 02420) from Crawfish Valley left gonopod; red triangle indicates reduced prefemoral spine." figureDoi="10.3897/zookeys.1030.62544.figure84" httpUri="https://binary.pensoft.net/fig/531187" pageId="0" pageNumber="1">84</figureCitation>
, red triangle).
</paragraph>
<caption doi="10.3897/zookeys.1030.62544.figure84" httpUri="https://binary.pensoft.net/fig/531187" pageId="0" pageNumber="1" start="Figure 84" startId="F84">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 84.</emphasis>
<taxonomicName authorityName="Means &amp; Hennen &amp; Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria solenas" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="solenas">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria solenas</emphasis>
</taxonomicName>
sp. nov. non-type ♂ (VTEC, MPE02420) from Crawfish Valley left gonopod; red triangle indicates reduced prefemoral spine.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
Known from southwestern Virginia and southeastern West Virginia, with an individual from central West Virginia (West Virginia: Fayette, Mercer, and Summers counties; Virginia: Bland, Giles, Russell, Tazewell, and Wythe counties, Suppl. material 7; Fig.
<figureCitation captionStart="Figure 126" captionStartId="F126" captionText="Figure 126. Distribution map of Nannaria minor species group collection localities in the central Appalachians." figureDoi="10.3897/zookeys.1030.62544.figure126" httpUri="https://binary.pensoft.net/fig/531229" pageId="0" pageNumber="1">126</figureCitation>
). Distribution area: 3,009 km2; status: SRE.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="ecology">
<paragraph pageId="0" pageNumber="1">Ecology.</paragraph>
<paragraph pageId="0" pageNumber="1">
Individuals of
<taxonomicName authorityName="Means &amp; Hennen &amp; Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria solenas" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="solenas">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria solenas</emphasis>
</taxonomicName>
sp. nov. have been collected from mesic hardwood forests, dominated by oak, maple, and pine. Specimens from Crawfish Valley were collected at night while walking on top of grass in an overgrown road.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is named after its type locality, Pipestem Resort State Park in West Virginia. The specific name is a noun in apposition derived from the Greek
<emphasis italics="true" pageId="0" pageNumber="1">
<normalizedToken originalValue="solínas">solinas</normalizedToken>
</emphasis>
, meaning pipe.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type locality">
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">
United States, West Virginia, Summers County, Pipestem Resort State Park, path to lake, ca. 12 km northeast of Athens,
<geoCoordinate degrees="37.5278" direction="north" orientation="latitude" precision="5" value="37.5278">37.5278°N</geoCoordinate>
, -
<geoCoordinate degrees="80.9889" direction="west" orientation="longitude" precision="5" value="-80.9889">80.9889°W</geoCoordinate>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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