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<document ID-CLB-Dataset= "55415" ID-DOI= "http://dx.doi.org/10.3897/zookeys.1176.106224" ID-GBIF-Dataset= "7bb917e2-6ad1-4f3f-a1ae-df3feeb99862" ID-PMC= "PMC10468690" ID-Pensoft-Pub= "1313-2970-1176-165" ID-Pensoft-UUID= "C538BFA63A3455D089E0A07BC22A97C6" ID-PubMed= "37664866" ID-ZooBank= "91E50336528C43DE826642D12D0C7C1C" ModsDocID= "1313-2970-1176-165" checkinTime= "1692810056931" checkinUser= "pensoft" docAuthor= "Khan, Eid Muhammad, Kamran, Muhammad, Mirza, Jawwad Hassan & Alatawi, Fahad Jaber" docDate= "2023" docId= "4E0A4286D8715265AAD699D360ED0DEF" docLanguage= "en" docName= "ZooKeys 1176: 165-180" docOrigin= "ZooKeys 1176" docPubDate= "2023-08-23" docSource= "http://dx.doi.org/10.3897/zookeys.1176.106224" docTitle= "Raphignathus (Diraphignathus) neohecmatanaensis sp. nov." docType= "treatment" docVersion= "4" id= "C538BFA63A3455D089E0A07BC22A97C6" lastPageNumber= "165" masterDocId= "C538BFA63A3455D089E0A07BC22A97C6" masterDocTitle= "New subgenera and a new species of the genus Raphignathus Duges (Prostigmata, Raphignathidae), with taxonomic notes on the genus Neoraphignathus Smiley & Moser" masterLastPageNumber= "180" masterPageNumber= "165" pageNumber= "165" updateTime= "1732480068003" updateUser= "ExternalLinkService" >
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<mods:title id= "175644203169857B2838C54ECF696B19" > New subgenera and a new species of the genus Raphignathus Duges (Prostigmata, Raphignathidae), with taxonomic notes on the genus Neoraphignathus Smiley & Moser</mods:title>
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<mods:namePart id= "3B502A6554CB83B055762940E75F9EDF" > Khan, Eid Muhammad</mods:namePart>
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<mods:affiliation id= "DF53BCA3BA06F1CA1D222F5803A574DE" > Acarology Research Laboratory, Department of Plant Protection, College of Food and Agriculture Sciences, King Saud University, Riyadh, Saudi Arabia</mods:affiliation>
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<mods:namePart id= "19070C5E02EB36FAE94301254724C5F1" > Kamran, Muhammad</mods:namePart>
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<mods:affiliation id= "5F7459BDDE1D464186CE45FDA5F2875B" > Acarology Research Laboratory, Department of Plant Protection, College of Food and Agriculture Sciences, King Saud University, Riyadh, Saudi Arabia</mods:affiliation>
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<mods:namePart id= "03A5C8F0E1B3487FB56CC70F642873B7" > Mirza, Jawwad Hassan</mods:namePart>
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<mods:affiliation id= "36194B52E32C0406F06A446DB5ED4B1E" > Acarology Research Laboratory, Department of Plant Protection, College of Food and Agriculture Sciences, King Saud University, Riyadh, Saudi Arabia</mods:affiliation>
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<mods:namePart id= "E6B65850B0A34F94EF22058A0C2947BB" > Alatawi, Fahad Jaber</mods:namePart>
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<mods:nameIdentifier id= "06B76433C2F36100A5DE1D95A8588F37" type= "email" > falatawi@ksu.edu.sa</mods:nameIdentifier>
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<treatment id= "4E0A4286D8715265AAD699D360ED0DEF" ID-GBIF-Taxon= "212453373" LSID= "urn:lsid:zoobank.org:act:12E13CEE-478F-4A3F-A47E-B3C7F34C73FA" httpUri= "http://treatment.plazi.org/id/4E0A4286D8715265AAD699D360ED0DEF" lastPageNumber= "165" pageId= "0" pageNumber= "165" scope_family= "Raphignathidae" scope_subGenus= "Raphignathus" scope_subOrder= "Prostigmata" >
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<subSubSection id= "8CF8D0C57A6E27E5CA5C9B034A8FDA6E" pageId= "0" pageNumber= "165" type= "nomenclature" >
<paragraph id= "4922FD89CE8A0924410A159D05E3AF33" pageId= "0" pageNumber= "165" >
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<taxonomicName id= "C3CFF774450E2C6530C5D05A1D53B9B9" LSID= "https://zoobank.org/12E13CEE-478F-4A3F-A47E-B3C7F34C73FA" authority= "sp. nov." authorityName= "sp. nov." family= "Raphignathidae" genus= "Raphignathus" higherTaxonomySource= "treatment-meta" kingdom= "Animalia" lsidName= "Raphignathus (Diraphignathus) neohecmatanaensis" order= "Prostigmata" pageId= "0" pageNumber= "165" rank= "species" species= "neohecmatanaensis" subGenus= "Diraphignathus" > Raphignathus (Diraphignathus) neohecmatanaensis sp. nov.</taxonomicName>
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</paragraph>
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<subSubSection id= "2BB2961A89646967D0D8DCB132D1507E" pageId= "0" pageNumber= "165" type= "description" >
<paragraph id= "7B6077DF8D0AD3F408BF764FEBCC930D" pageId= "0" pageNumber= "165" >
<figureCitation id= "D17D1D727BBE4A6DA0FCF3989E113C1D" captionStart= "Figure 1" captionStartId= "F1" captionText= "Figure 1. Raphignathus (Diraphignathus) neohecmatanaensis sp. nov. (female), dorsum. Scale bar: 100 µm." figureDoi= "10.3897/zookeys.1176.106224.figure1" httpUri= "https://binary.pensoft.net/fig/895996" pageId= "0" pageNumber= "165" > Figs 1</figureCitation>
<figureCitation id= "B80DE894D2387CB16AFE67A3F979CD90" captionStart= "Figure 2" captionStartId= "F2" captionText= "Figure 2. Raphignathus (Diraphignathus) neohecmatanaensis sp. nov. (female), venter. Scale bar: 100 µm." figureDoi= "10.3897/zookeys.1176.106224.figure2" httpUri= "https://binary.pensoft.net/fig/895997" pageId= "0" pageNumber= "165" > , 2</figureCitation>
<figureCitation id= "5478F47D6EEFE884045A7CD9317B82B4" captionStart= "Figure 3" captionStartId= "F3" captionText= "Figure 3. Raphignathus (Diraphignathus) neohecmatanaensis sp. nov. (female), gnathosoma. Scale bar: 50 µm." figureDoi= "10.3897/zookeys.1176.106224.figure3" httpUri= "https://binary.pensoft.net/fig/895998" pageId= "0" pageNumber= "165" > , 3</figureCitation>
<figureCitation id= "F07A3F508491F12A2C6847CA1DF83A2C" captionStart= "Figure 4" captionStartId= "F4" captionText= "Figure 4. Raphignathus (Diraphignathus) neohecmatanaensis sp. nov. (female), palp. Scale bar: 20 µm." figureDoi= "10.3897/zookeys.1176.106224.figure4" httpUri= "https://binary.pensoft.net/fig/895999" pageId= "0" pageNumber= "165" > , 4</figureCitation>
<figureCitation id= "A533C337FB02E9B01F869756CD2D858C" captionStart= "Figures 5– captionStartId= "F5" captionText= "Figures 5 - 8. Raphignathus (Diraphignathus) neohecmatanaensis sp. nov. (female), Legs I-IV. Scale bar: 20 µm." figureDoi= "10.3897/zookeys.1176.106224.figures5-8" httpUri= "https://binary.pensoft.net/fig/896000" pageId= "0" pageNumber= "165" > , 5-8</figureCitation>
</paragraph>
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<subSubSection id= "00BC80EA5C78C315A01B101751325AE4" pageId= "0" pageNumber= "165" type= "diagnosis" >
<paragraph id= "0314D918205F73BE2803145CA87F7A42" pageId= "0" pageNumber= "165" > Diagnosis.</paragraph>
<paragraph id= "6D2F4560C18FEB5EE75CB8F0363022F9" pageId= "0" pageNumber= "165" >
<emphasis id= "33213AA86DAEE1B52FC0143162E8A3AB" bold= "true" pageId= "0" pageNumber= "165" > Female</emphasis>
: endopodal shield absent between coxae I-IV; two small shields present posterolateral to median prodorsal shield; palp femora with two setae; femora 6-5-3-2; genua 5(+κ)-5(+κ)-4-4; tibiae 5(+φρ)-5(+φρ)-4(+φρ)-4 (+φρ); tarsi 21(1ω) -20(1ω)-15-14.
</paragraph>
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<subSubSection id= "40A79FFB43CC71D9ED60AA1C6B98E764" pageId= "0" pageNumber= "165" type= "description" >
<paragraph id= "A597CE30088B3EEFD9979567178F615A" pageId= "0" pageNumber= "165" > Description.</paragraph>
<paragraph id= "ADF27E13EE6CD27924DC52BE0B91A00C" pageId= "0" pageNumber= "165" >
<emphasis id= "5F9A63C89102FFAD7033C0C3B3DB5FD3" bold= "true" pageId= "0" pageNumber= "165" > Female</emphasis>
(
<emphasis id= "55FB77A38A345A78107EFC684B462C35" italics= "true" pageId= "0" pageNumber= "165" > n</emphasis>
= 4). Idiosoma oval, length of body (including gnathosoma) 533 (525-545); width of body 345 (338-353).
</paragraph>
<paragraph id= "FA0E15B418CC3C17C0563EBB5A0F7E71" pageId= "0" pageNumber= "165" >
<emphasis id= "A3982EB227EF62761746C64322D6CF24" bold= "true" italics= "true" pageId= "0" pageNumber= "165" > Dorsum</emphasis>
(Fig.
<figureCitation id= "EDF99A8F8C328A8765CBF54326D559F7" captionStart= "Figure 1" captionStartId= "F1" captionText= "Figure 1. Raphignathus (Diraphignathus) neohecmatanaensis sp. nov. (female), dorsum. Scale bar: 100 µm." figureDoi= "10.3897/zookeys.1176.106224.figure1" httpUri= "https://binary.pensoft.net/fig/895996" pageId= "0" pageNumber= "165" > 1</figureCitation>
). Propodosoma with one medial and two lateral shields each containing three setae; medial sclerite with setae
<emphasis id= "6BA17EFE03731755083A3EB18126DB23" italics= "true" pageId= "0" pageNumber= "165" > vi</emphasis>
,
<emphasis id= "52B2DBE155665D3DDD55067CFC4EB7AB" italics= "true" pageId= "0" pageNumber= "165" > si</emphasis>
and
<emphasis id= "88DA99B2BC137374D40CDB7AE25D3F2E" italics= "true" pageId= "0" pageNumber= "165" > c1</emphasis>
; paired ovoid lateral shields each with an eye, one cupule (
<emphasis id= "787D7690F516FE9E51EC3080104AD05A" italics= "true" pageId= "0" pageNumber= "165" > ia</emphasis>
) and
<emphasis id= "345F126FA8629D0B9DD93E77376EA307" italics= "true" pageId= "0" pageNumber= "165" > sci</emphasis>
,
<emphasis id= "AB890E78D57CCC11983F40D47BD3F8B5" italics= "true" pageId= "0" pageNumber= "165" > sce</emphasis>
and seta
<emphasis id= "626ADD6358F688D543D96E078C4C727C" italics= "true" pageId= "0" pageNumber= "165" > c2</emphasis>
seta; opisthosomal setal pairs
<emphasis id= "047E03FA445712422474027C3D8B62A1" italics= "true" pageId= "0" pageNumber= "165" > d1</emphasis>
,
<emphasis id= "87F79207329C7D0E03481246E5A4AF6E" italics= "true" pageId= "0" pageNumber= "165" > e1</emphasis>
and the pair of cupuli (
<emphasis id= "8B218D5DFC063DF63AD99A9731B54415" italics= "true" pageId= "0" pageNumber= "165" > im</emphasis>
); posterior opisthosomal shield large, rectangular, bearing four pairs of setae (
<emphasis id= "099CE1C72BF469A1466C7990F3941B9D" italics= "true" pageId= "0" pageNumber= "165" > f1</emphasis>
,
<emphasis id= "859229FF63E12CD2F382881271D0A92C" italics= "true" pageId= "0" pageNumber= "165" > h1-3</emphasis>
) and one pair of the cupule (
<emphasis id= "477F4F0266D3EACC3DFB0EA0185EC037" italics= "true" pageId= "0" pageNumber= "165" > ip</emphasis>
); all dorsal shields finely punctate; dorsal body setae setiform, smooth and acute; pseudanal setae
<emphasis id= "653F8C60E0DB831D91B9336F04B02660" italics= "true" pageId= "0" pageNumber= "165" > ps1</emphasis>
dorsally located. Lengths of dorsal setae:
<emphasis id= "D773E15E1E627B35E6EDCD3FCC28A80F" italics= "true" pageId= "0" pageNumber= "165" > vi</emphasis>
25 (23-27),
<emphasis id= "026D5802CF682B6DF0693B9B0817E007" italics= "true" pageId= "0" pageNumber= "165" > ve</emphasis>
28 (26-30),
<emphasis id= "BE37D46DA3029A775DA88288A6B9E192" italics= "true" pageId= "0" pageNumber= "165" > sci</emphasis>
27 (26-29),
<emphasis id= "17695013FFA2654F0458ECDFBCDD9BF9" italics= "true" pageId= "0" pageNumber= "165" > sce</emphasis>
28 (26-29),
<emphasis id= "B8DA7DAB5D00ED239F9AE5BF983C1D67" italics= "true" pageId= "0" pageNumber= "165" > c1</emphasis>
23 (17-19),
<emphasis id= "2C63D3F9CCBA07DEB5528EE11E92D270" italics= "true" pageId= "0" pageNumber= "165" > c2</emphasis>
25 (19-23),
<emphasis id= "E745E6C70EF89B301C2BD6BC1A0C7ECC" italics= "true" pageId= "0" pageNumber= "165" > d1</emphasis>
21 (20-23),
<emphasis id= "64E4C6A428A5DD548F3276BAEFA7F31A" italics= "true" pageId= "0" pageNumber= "165" > e1</emphasis>
22 (21-23),
<emphasis id= "68E6D94BD11171E5B6234EF06079729F" italics= "true" pageId= "0" pageNumber= "165" > f1</emphasis>
21 (19-22),
<emphasis id= "DFEE3DC235123C860BD60A1F631C5361" italics= "true" pageId= "0" pageNumber= "165" > h1</emphasis>
24 (23-25),
<emphasis id= "BC7E58937DED8E301665D1F0D69EA21C" italics= "true" pageId= "0" pageNumber= "165" > h2</emphasis>
25 (23-25),
<emphasis id= "9BBC95B942A2455E76EC20A1CCB67AE1" italics= "true" pageId= "0" pageNumber= "165" > h3</emphasis>
22 (21-24),
<emphasis id= "F41AC7128A68947D71136AA76D2E565A" italics= "true" pageId= "0" pageNumber= "165" > ps1</emphasis>
23 (21-25); distances between dorsal 195 setae:
<emphasis id= "0E2CC7804F676544ACAC6FFA00ACC31B" italics= "true" pageId= "0" pageNumber= "165" > vi-vi</emphasis>
27 (29-31),
<emphasis id= "0E2533F98113BAF11E1D82078552E169" italics= "true" pageId= "0" pageNumber= "165" > sci-sci</emphasis>
123 (118-128),
<emphasis id= "E2C31D465DA108783472F0794BE87F56" italics= "true" pageId= "0" pageNumber= "165" > vi- sci</emphasis>
70 (66-72),
<emphasis id= "04A88E229A7B2B28A62A1C64807FE235" italics= "true" pageId= "0" pageNumber= "165" > sce-c2</emphasis>
68 (65-72),
<emphasis id= "8F62A2F13BC39D64783F66782DC121B2" italics= "true" pageId= "0" pageNumber= "165" > c1-c1</emphasis>
30 (28-32),
<emphasis id= "63AAB95C2B2A805B24E84A4D3E5C7814" italics= "true" pageId= "0" pageNumber= "165" > d1-d1</emphasis>
99 (92-101),
<emphasis id= "B093EF6D96BE4EE61A1A7DE93C3F1278" italics= "true" pageId= "0" pageNumber= "165" > c1-d1</emphasis>
53 (49-55),
<emphasis id= "77B2C305E076670321C6DA0D119EBB0B" italics= "true" pageId= "0" pageNumber= "165" > d1-e1</emphasis>
22 (21-24),
<emphasis id= "5E8023CE6F8E1A83A85924FBDBDE2E93" italics= "true" pageId= "0" pageNumber= "165" > f1-f1</emphasis>
88 (83-90),
<emphasis id= "6572B2F6050EA2EB400696B1F4F40DC6" italics= "true" pageId= "0" pageNumber= "165" > e1-f1</emphasis>
65 (61-68),
<emphasis id= "EB903D514D05279931C912CFB3D72246" italics= "true" pageId= "0" pageNumber= "165" > h1-h1</emphasis>
45 (42-48),
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43 (41-46),
<emphasis id= "BDA11720C2E7847D7AC0A73F07F11510" italics= "true" pageId= "0" pageNumber= "165" > h2-h2</emphasis>
77 (72-80),
<emphasis id= "4F949D6FB34FF7DE4542B401A407AA35" italics= "true" pageId= "0" pageNumber= "165" > h3-h3</emphasis>
101 (98-106).
</paragraph>
<caption id= "7706F519EECD021F4A52541C7D5D5C17" doi= "10.3897/zookeys.1176.106224.figure1" httpUri= "https://binary.pensoft.net/fig/895996" pageId= "0" pageNumber= "165" start= "Figure 1" startId= "F1" >
<paragraph id= "32EBB8F7273DAE93D41090BF60D05BC9" pageId= "0" pageNumber= "165" >
<emphasis id= "C38BCF998EF48A55D4DCE54D2BCD25EB" bold= "true" pageId= "0" pageNumber= "165" > Figure 1.</emphasis>
2024-11-25 00:49:40 +01:00
<taxonomicName id= "ED9D257BCCBCED5C80DB4D20BBE655F4" family= "Raphignathidae" genus= "Raphignathus" kingdom= "Animalia" lsidName= "Raphignathus (Diraphignathus) neohecmatanaensis" order= "Prostigmata" pageId= "0" pageNumber= "165" rank= "species" species= "neohecmatanaensis" subGenus= "Diraphignathus" > Raphignathus (Diraphignathus) neohecmatanaensis</taxonomicName>
2024-06-21 12:36:04 +02:00
sp. nov. (female), dorsum. Scale bar: 100
<normalizedToken id= "E1B69706D66917DD077D9FC1CF3F5672" originalValue= "µm" > µm</normalizedToken>
.
</paragraph>
</caption>
<paragraph id= "9F537A7161FA6FA204610D4350FFE834" pageId= "0" pageNumber= "165" >
<emphasis id= "D48A040823374A3D3DE888681853A87C" bold= "true" italics= "true" pageId= "0" pageNumber= "165" > Venter</emphasis>
(Fig.
<figureCitation id= "764BB44EA7BF71EEA0B3D56742437928" captionStart= "Figure 2" captionStartId= "F2" captionText= "Figure 2. Raphignathus (Diraphignathus) neohecmatanaensis sp. nov. (female), venter. Scale bar: 100 µm." figureDoi= "10.3897/zookeys.1176.106224.figure2" httpUri= "https://binary.pensoft.net/fig/895997" pageId= "0" pageNumber= "165" > 2</figureCitation>
). Venter entirely striated, without punctations; coxisternal shields absent (Fig.
<figureCitation id= "E676F5E7B616AB2A0E98E72D8B86819A" captionStart= "Figure 2" captionStartId= "F2" captionText= "Figure 2. Raphignathus (Diraphignathus) neohecmatanaensis sp. nov. (female), venter. Scale bar: 100 µm." figureDoi= "10.3897/zookeys.1176.106224.figure2" httpUri= "https://binary.pensoft.net/fig/895997" pageId= "0" pageNumber= "165" > 2</figureCitation>
); ventral setae lengths:
<emphasis id= "D83F127E9B9B0809AE3893159DD2024B" italics= "true" pageId= "0" pageNumber= "165" > 1a</emphasis>
42 (41-44),
<emphasis id= "FE40C9594C7C5050F0F47D490704660F" italics= "true" pageId= "0" pageNumber= "165" > 1b</emphasis>
38 (35-39),
<emphasis id= "2FA577762AC667B6820CA2E663EF5410" italics= "true" pageId= "0" pageNumber= "165" > 1c</emphasis>
34 (33-36),
<emphasis id= "FD7F984AA8132BD50C598163D83033FD" italics= "true" pageId= "0" pageNumber= "165" > 2b</emphasis>
36 (30-34),
<emphasis id= "EC39C37CA7D5FE2837E5D6A71785741C" italics= "true" pageId= "0" pageNumber= "165" > 2c</emphasis>
32 (30-34),
<emphasis id= "5B36365DD116959B4778DC570A993958" italics= "true" pageId= "0" pageNumber= "165" > 3a</emphasis>
24 (23-26),
<emphasis id= "A2D7395346B5F63FE096F60CA56D587A" italics= "true" pageId= "0" pageNumber= "165" > 3b</emphasis>
18 (17-20),
<emphasis id= "F769DA7B4D80D918B278476A59DD5737" italics= "true" pageId= "0" pageNumber= "165" > 3c</emphasis>
36 (34-38),
<emphasis id= "2DC688253FAC612EE2E78C5BACEBBAC1" italics= "true" pageId= "0" pageNumber= "165" > 4a</emphasis>
22 (23-27),
<emphasis id= "949B0F07CD8FB64A622BDADAD9C218F3" italics= "true" pageId= "0" pageNumber= "165" > 4c</emphasis>
38 (35-40); two pairs of aggenital setae (
<emphasis id= "B3A9613BF39D01078955AB1E19A93867" italics= "true" pageId= "0" pageNumber= "165" > ag1-2</emphasis>
) with one cupule (
<emphasis id= "E86D54643E084F25B09FA0021C520AF0" italics= "true" pageId= "0" pageNumber= "165" > ih</emphasis>
) on each side of the genital shield; anal opening and genital shields separate; genital shield prominent with a few punctations, bearing three pairs of genital setae (
<emphasis id= "860B3695EA8F1A41BDB40EFC783A754F" italics= "true" pageId= "0" pageNumber= "165" > g1-3</emphasis>
); anal opening terminal, with three pairs of setae (
<emphasis id= "5461B2D23947866D040D34135F05A629" italics= "true" pageId= "0" pageNumber= "165" > ps1</emphasis>
-3),
<emphasis id= "089CFF49C57EF13333B7538EAE1D3585" italics= "true" pageId= "0" pageNumber= "165" > ps1</emphasis>
dorsal
<emphasis id= "97C4B898100515117D38C85C87E46563" italics= "true" pageId= "0" pageNumber= "165" > ps3</emphasis>
and
<emphasis id= "12071EBE7C68D87E835C2361FE4CA32E" italics= "true" pageId= "0" pageNumber= "165" > ps2</emphasis>
ventral; ventral setal lengths:
<emphasis id= "0BBBA18DE2F5DAED2E270D32FBB5D91B" italics= "true" pageId= "0" pageNumber= "165" > ag1</emphasis>
29 (27-32);
<emphasis id= "776FCBBA81A1F573F43890F5834335B5" italics= "true" pageId= "0" pageNumber= "165" > ag2</emphasis>
27 (25-28);
<emphasis id= "3A6FA4E986113005BA48D459962CCF2A" italics= "true" pageId= "0" pageNumber= "165" > g1</emphasis>
31 (28-33);
<emphasis id= "B3969441A36E1F65FAAE05E48798EF1E" italics= "true" pageId= "0" pageNumber= "165" > g2</emphasis>
25 (23-28);
<emphasis id= "E264A67426BC3B3980F578144D1FE311" italics= "true" pageId= "0" pageNumber= "165" > g3</emphasis>
20 (19-22);
<emphasis id= "DA9DDB48C4E07EF1A803750C6609E499" italics= "true" pageId= "0" pageNumber= "165" > ps2</emphasis>
22 (21-23);
<emphasis id= "B6F1C802ED999362B35853BE47BC0C09" italics= "true" pageId= "0" pageNumber= "165" > ps3</emphasis>
21 (20-22). Distances between ventral setae:
<emphasis id= "4B726D45C6F7D2418E5ABCDCDA1D4742" italics= "true" pageId= "0" pageNumber= "165" > 1a-1a</emphasis>
55 (53-58),
<emphasis id= "6EDB98491537AFEFF6C1DAF972C4FDBD" italics= "true" pageId= "0" pageNumber= "165" > 3a-3a</emphasis>
115 (97-109),
<emphasis id= "CFDF88823B027B37D99566A7EA3E621A" italics= "true" pageId= "0" pageNumber= "165" > 4a-4a</emphasis>
70 (68-73),
<emphasis id= "F216DC042E6EEDED74501C74DACE70B9" italics= "true" pageId= "0" pageNumber= "165" > ag2-ag2</emphasis>
42 (41-43),
<emphasis id= "1C37F655B74ABC969E143019D45A422E" italics= "true" pageId= "0" pageNumber= "165" > g1-g1</emphasis>
38 (35-40),
<emphasis id= "B7AA2E718B902E87770BBD415585A78F" italics= "true" pageId= "0" pageNumber= "165" > g2-g2</emphasis>
52 (48-55),
<emphasis id= "DFB21224E703CB1B9E5F0BB1A60BFDF6" italics= "true" pageId= "0" pageNumber= "165" > g3-g3</emphasis>
75 (74-75),
<emphasis id= "A4B7F61D0B5753FAF95CB9BF42F5C5AF" italics= "true" pageId= "0" pageNumber= "165" > 2b-2c</emphasis>
30 (25-29),
<emphasis id= "D662DB3AAE73978808B2855C693CD0EB" italics= "true" pageId= "0" pageNumber= "165" > 1a-3a</emphasis>
50 (49-53),
<emphasis id= "FDA56B09BC9321FBC2E779206FD5CCF6" italics= "true" pageId= "0" pageNumber= "165" > 3a-4a</emphasis>
72 (68-75),
<emphasis id= "8DD56E78FC6E76A9ED68B81F0E5E8047" italics= "true" pageId= "0" pageNumber= "165" > 4a-ag1</emphasis>
85 (82-88),
<emphasis id= "3A73AB2213A54695F7D0007761D456E8" italics= "true" pageId= "0" pageNumber= "165" > ag1-ag2</emphasis>
65 (63-68),
<emphasis id= "CD5484D2220E0C466F5241D37D11BE05" italics= "true" pageId= "0" pageNumber= "165" > ag2-g1</emphasis>
63 (60-65),
<emphasis id= "CF2C1A07AAB990743A5F526296F05EAD" italics= "true" pageId= "0" pageNumber= "165" > g1-g2</emphasis>
15 (14-17),
<emphasis id= "D2CCD3A8C9AC1E0EEEEDE1F574617F0D" italics= "true" pageId= "0" pageNumber= "165" > g2-g3</emphasis>
22 (21-25),
<emphasis id= "397D892DC5C297ACC007484580CBB0B2" italics= "true" pageId= "0" pageNumber= "165" > ag-g1</emphasis>
42 (40-45),
<emphasis id= "D5556BEBE6243373B17CB090237B7995" italics= "true" pageId= "0" pageNumber= "165" > g3-ps3</emphasis>
16 (15-18),
<emphasis id= "E470E54E13415DEFFAD7721B4508D508" italics= "true" pageId= "0" pageNumber= "165" > ps2- ps3</emphasis>
16 (14-18).
</paragraph>
<caption id= "B353663262C7AB3AB847C9D98BAA8975" doi= "10.3897/zookeys.1176.106224.figure2" httpUri= "https://binary.pensoft.net/fig/895997" pageId= "0" pageNumber= "165" start= "Figure 2" startId= "F2" >
<paragraph id= "7E62751D3356277EC5071A35A3EB40BA" pageId= "0" pageNumber= "165" >
<emphasis id= "4A95E12020F3119122400E07BE60B249" bold= "true" pageId= "0" pageNumber= "165" > Figure 2.</emphasis>
2024-11-25 00:49:40 +01:00
<taxonomicName id= "44B27DC94A412BEFD6704767E4782039" family= "Raphignathidae" genus= "Raphignathus" kingdom= "Animalia" lsidName= "Raphignathus (Diraphignathus) neohecmatanaensis" order= "Prostigmata" pageId= "0" pageNumber= "165" rank= "species" species= "neohecmatanaensis" subGenus= "Diraphignathus" > Raphignathus (Diraphignathus) neohecmatanaensis</taxonomicName>
2024-06-21 12:36:04 +02:00
sp. nov. (female), venter. Scale bar: 100
<normalizedToken id= "C3A8AB46282211C29A6CBA8AC8B43674" originalValue= "µm" > µm</normalizedToken>
.
</paragraph>
</caption>
<paragraph id= "3485598787B3561DA464CD239BF9859F" pageId= "0" pageNumber= "165" >
<emphasis id= "2B110FCC382916035B63F9ED11F2C302" bold= "true" italics= "true" pageId= "0" pageNumber= "165" > Gnathosoma</emphasis>
(Figs
<figureCitation id= "DE2BD1519CC2757D6B5EA5A8C13DBFFD" captionStart= "Figure 3" captionStartId= "F3" captionText= "Figure 3. Raphignathus (Diraphignathus) neohecmatanaensis sp. nov. (female), gnathosoma. Scale bar: 50 µm." figureDoi= "10.3897/zookeys.1176.106224.figure3" httpUri= "https://binary.pensoft.net/fig/895998" pageId= "0" pageNumber= "165" > 3</figureCitation>
,
<figureCitation id= "6D1186D6258D8BFB6A7034A41BD0EA0A" captionStart= "Figure 4" captionStartId= "F4" captionText= "Figure 4. Raphignathus (Diraphignathus) neohecmatanaensis sp. nov. (female), palp. Scale bar: 20 µm." figureDoi= "10.3897/zookeys.1176.106224.figure4" httpUri= "https://binary.pensoft.net/fig/895999" pageId= "0" pageNumber= "165" > 4</figureCitation>
). Ventral infracapitular with two pairs of very long setae (
<emphasis id= "34B8F035CF5598EDECC65145AD7A2F90" italics= "true" pageId= "0" pageNumber= "165" > m</emphasis>
and
<emphasis id= "163C472C653C6F9CD933D67CDB18BEED" italics= "true" pageId= "0" pageNumber= "165" > n</emphasis>
),
<emphasis id= "D97E911FC5BFD56327C15CE0D8E1C2A1" italics= "true" pageId= "0" pageNumber= "165" > m</emphasis>
40 (39-42),
<emphasis id= "1C5714584ED08789DF317577EC03090D" italics= "true" pageId= "0" pageNumber= "165" > n</emphasis>
52 (49-54) and two pairs of pilose adoral setae (
<emphasis id= "DA20C8F7E8964061D545ABBA708456FD" italics= "true" pageId= "0" pageNumber= "165" > or1-2</emphasis>
),
<emphasis id= "6F096FE313351E7F8B7D04C4BAE6CC7E" italics= "true" pageId= "0" pageNumber= "165" > or1</emphasis>
23 (21-24),
<emphasis id= "290D2B5FD415843D5417B6E4A9376AC5" italics= "true" pageId= "0" pageNumber= "165" > or2</emphasis>
20 (19-22) (Fig.
<figureCitation id= "2E6E8E229F9A60EFC5854213BB87193F" captionStart= "Figure 3" captionStartId= "F3" captionText= "Figure 3. Raphignathus (Diraphignathus) neohecmatanaensis sp. nov. (female), gnathosoma. Scale bar: 50 µm." figureDoi= "10.3897/zookeys.1176.106224.figure3" httpUri= "https://binary.pensoft.net/fig/895998" pageId= "0" pageNumber= "165" > 3</figureCitation>
); stylophore conical and striated; palp chaetotaxy (femur-tarsus) as follows: 3-2-4+1 claw 4+1ω+4 eupathidia (ζ) (Fig.
<figureCitation id= "C01816F56444D87B64B6E704D7F07F07" captionStart= "Figure 4" captionStartId= "F4" captionText= "Figure 4. Raphignathus (Diraphignathus) neohecmatanaensis sp. nov. (female), palp. Scale bar: 20 µm." figureDoi= "10.3897/zookeys.1176.106224.figure4" httpUri= "https://binary.pensoft.net/fig/895999" pageId= "0" pageNumber= "165" > 4</figureCitation>
).
</paragraph>
<caption id= "D4B1DC67656D031CED23222AAEB19E99" doi= "10.3897/zookeys.1176.106224.figure3" httpUri= "https://binary.pensoft.net/fig/895998" pageId= "0" pageNumber= "165" start= "Figure 3" startId= "F3" >
<paragraph id= "A459F84CAC9B693158C9B7C372A3A553" pageId= "0" pageNumber= "165" >
<emphasis id= "5FC90FD43638FF2AA0F3E19B30795D8C" bold= "true" pageId= "0" pageNumber= "165" > Figure 3.</emphasis>
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<taxonomicName id= "CEC31D6A81CF87DD8593BABAC6F77E18" family= "Raphignathidae" genus= "Raphignathus" kingdom= "Animalia" lsidName= "Raphignathus (Diraphignathus) neohecmatanaensis" order= "Prostigmata" pageId= "0" pageNumber= "165" rank= "species" species= "neohecmatanaensis" subGenus= "Diraphignathus" > Raphignathus (Diraphignathus) neohecmatanaensis</taxonomicName>
2024-06-21 12:36:04 +02:00
sp. nov. (female), gnathosoma. Scale bar: 50
<normalizedToken id= "8D5D38FDEF4D23382FADB42137D94092" originalValue= "µm" > µm</normalizedToken>
.
</paragraph>
</caption>
<caption id= "763F214419CA5B0D4BAF7337CF7315ED" doi= "10.3897/zookeys.1176.106224.figure4" httpUri= "https://binary.pensoft.net/fig/895999" pageId= "0" pageNumber= "165" start= "Figure 4" startId= "F4" >
<paragraph id= "F3181648521A2898D4A1544C8C9E78CF" pageId= "0" pageNumber= "165" >
<emphasis id= "D0620E781959D161F94311060171B342" bold= "true" pageId= "0" pageNumber= "165" > Figure 4.</emphasis>
2024-11-25 00:49:40 +01:00
<taxonomicName id= "106CE6BFB6869F2CB1B4FA9ADEEEAE7C" family= "Raphignathidae" genus= "Raphignathus" kingdom= "Animalia" lsidName= "Raphignathus (Diraphignathus) neohecmatanaensis" order= "Prostigmata" pageId= "0" pageNumber= "165" rank= "species" species= "neohecmatanaensis" subGenus= "Diraphignathus" > Raphignathus (Diraphignathus) neohecmatanaensis</taxonomicName>
2024-06-21 12:36:04 +02:00
sp. nov. (female), palp. Scale bar: 20
<normalizedToken id= "2F6DB6DFB742B35FCC07CB20DEB0F6D2" originalValue= "µm" > µm</normalizedToken>
.
</paragraph>
</caption>
<paragraph id= "BCB309D92F600505E6BC1727981C902D" pageId= "0" pageNumber= "165" >
<emphasis id= "B0D8AD988A80E809488CA6D7FA794702" bold= "true" italics= "true" pageId= "0" pageNumber= "165" > Legs</emphasis>
(Figs
<figureCitation id= "818DDE4CA37DCC48FE6FF5740977EA44" captionStart= "Figures 5– captionStartId= "F5" captionText= "Figures 5 - 8. Raphignathus (Diraphignathus) neohecmatanaensis sp. nov. (female), Legs I-IV. Scale bar: 20 µm." figureDoi= "10.3897/zookeys.1176.106224.figures5-8" httpUri= "https://binary.pensoft.net/fig/896000" pageId= "0" pageNumber= "165" > 5-8</figureCitation>
). Length of legs I-IV (without coxae): 340 (328-348); 270 (276-286); 325 (317-328); 375 (367-384), respectively. Chaetotaxy on legs I-IV (solenidia in parentheses and not included in setal counts): coxa 2-2-2-1; trochanter 1-1-2-1; femora 6-5-3-2; genua 5(+κ)-5(+κ)-4-4; tibiae 5(+φρ)-5(+φρ)-4(+φρ)-4(+φρ); tarsi 21(1ω) -20(1ω)-15-14.
</paragraph>
<caption id= "C7EE2A971E65B24A9509C9B65AAA4AAC" doi= "10.3897/zookeys.1176.106224.figures5-8" httpUri= "https://binary.pensoft.net/fig/896000" pageId= "0" pageNumber= "165" start= "Figures 5– startId= "F5" >
<paragraph id= "85C107D04589A42CE80D43411D6FEDA6" pageId= "0" pageNumber= "165" >
<emphasis id= "BC056304AD26DCFB15A3C38450C1BD0E" bold= "true" pageId= "0" pageNumber= "165" > Figures 5-8.</emphasis>
2024-11-25 00:49:40 +01:00
<taxonomicName id= "89F74B7C4FA2012F0D0D98EEFD346F8C" family= "Raphignathidae" genus= "Raphignathus" kingdom= "Animalia" lsidName= "Raphignathus (Diraphignathus) neohecmatanaensis" order= "Prostigmata" pageId= "0" pageNumber= "165" rank= "species" species= "neohecmatanaensis" subGenus= "Diraphignathus" > Raphignathus (Diraphignathus) neohecmatanaensis</taxonomicName>
2024-06-21 12:36:04 +02:00
sp. nov. (female), Legs I-IV. Scale bar: 20
<normalizedToken id= "58D35E83F99ECEA27947F9DAA39CFD3D" originalValue= "µm" > µm</normalizedToken>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id= "A4D9A2266785B173E2BE5E1223902448" pageId= "0" pageNumber= "165" type= "male and immature stages" >
<paragraph id= "957428EDE88C6A7CA56C3F6538C7AC79" pageId= "0" pageNumber= "165" > Male and immature stages.</paragraph>
<paragraph id= "05EECDE8A5E31E8C26BD3A93032A760B" pageId= "0" pageNumber= "165" > Unknown.</paragraph>
</subSubSection>
<subSubSection id= "8CF52E08A3678DC18A34701A3449C41F" pageId= "0" pageNumber= "165" type= "type material" >
<paragraph id= "7BE510CDE2101453F96EACFC636C4338" pageId= "0" pageNumber= "165" > Type materials.</paragraph>
<paragraph id= "5440C05D5EDD4CE263A912D80D649F03" pageId= "0" pageNumber= "165" >
<emphasis id= "C53C2579D97F5A652457A9E80A3989C6" bold= "true" italics= "true" pageId= "0" pageNumber= "165" > Holotype</emphasis>
female and three paratype females, Faifa, Jazan,
<geoCoordinate id= "26822D3D5C45BA90E0577579CBE404D8" degrees= "24" direction= "north" minutes= "30.412" orientation= "latitude" precision= "1" value= "24.506866" > 24°30.412'N</geoCoordinate>
,
<geoCoordinate id= "05C0502F8D3DA4120C1999652B045DFC" degrees= "39" direction= "east" minutes= "36.578" orientation= "longitude" precision= "1" value= "39.609634" > 39°36.578'E</geoCoordinate>
, 8 Oct., 2020, collected from
<taxonomicName id= "3CF8582E92EF550F1C7648E5B68A02D6" class= "Dicotyledoneae" family= "Rhamnaceae" genus= "Ziziphus" higherTaxonomySource= "GBIF" kingdom= "Plantae" lsidName= "Ziziphus spina-christi" order= "Rhamnales" pageId= "0" pageNumber= "165" phylum= "Angiospermae" rank= "species" species= "spina-christi" >
<emphasis id= "0BA2FA7006917362924F1B1F5ABC354B" italics= "true" pageId= "0" pageNumber= "165" > Ziziphus spina-christi</emphasis>
</taxonomicName>
Mill. (
<taxonomicName id= "0DD9B63BE6CA09470CEC37F7B4F6EC25" class= "Dicotyledoneae" family= "Rhamnaceae" higherTaxonomySource= "GBIF" kingdom= "Plantae" lsidName= "" order= "Rhamnales" pageId= "0" pageNumber= "165" phylum= "Angiospermae" rank= "family" > Rhamnaceae</taxonomicName>
) by Eid M. Khan, Jawwad H. Mirza & Hafiz S. Mushtaq.
</paragraph>
</subSubSection>
<subSubSection id= "6DD3E53261C5EADC1C8FEC22476E063E" pageId= "0" pageNumber= "165" type= "etymology" >
<paragraph id= "BC20FEE951DD1852DD91FEE60CE85E76" pageId= "0" pageNumber= "165" > Etymology.</paragraph>
<paragraph id= "63141A627A90AA539A87920D8050E49A" pageId= "0" pageNumber= "165" >
The specific epithet is in reference to the similarity of the new species to
2024-11-25 00:49:40 +01:00
<taxonomicName id= "24CE9AB489FCCD60AF5CF5EE8E353D06" class= "Arachnida" family= "Raphignathidae" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "R. (D.) hecmatanaensis" order= "Prostigmata" pageId= "0" pageNumber= "165" phylum= "Arthropoda" rank= "species" species= "hecmatanaensis" subGenus= "D." > R. (D.) hecmatanaensis</taxonomicName>
2024-06-21 12:36:04 +02:00
;
<emphasis id= "4A7685E1AD8CC8DC7343909A1922AC64" italics= "true" pageId= "0" pageNumber= "165" > neo</emphasis>
= new.
</paragraph>
</subSubSection>
<subSubSection id= "8D66110CB16D78D1F225D32A7800C020" pageId= "0" pageNumber= "165" type= "remarks" >
<paragraph id= "C1E48608C433D44219785AAB2C42EE2F" pageId= "0" pageNumber= "165" > Remarks.</paragraph>
<paragraph id= "BD8447BEE5BCEA3D29791693B003C24B" pageId= "0" pageNumber= "165" >
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<taxonomicName id= "33ACBC9DEC7AB1A7A032951E485BE9A4" family= "Raphignathidae" genus= "Raphignathus" kingdom= "Animalia" lsidName= "Raphignathus (D.) neohecmatanaensis" order= "Prostigmata" pageId= "0" pageNumber= "165" rank= "species" species= "neohecmatanaensis" subGenus= "D." > Raphignathus (D.) neohecmatanaensis</taxonomicName>
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sp. nov., belongs to the subgenus
<taxonomicName id= "C904A0B0A9CB1E5CEDA3D9408D198008" genus= "Diraphignathus" lsidName= "Diraphignathus" pageId= "0" pageNumber= "165" rank= "genus" > Diraphignathus</taxonomicName>
subgen. nov. The new species resembles to
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<taxonomicName id= "FDD73418DCFB696BA7C3C03B9A654286" class= "Arachnida" family= "Raphignathidae" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "R. (D.) hecmatanaensis" order= "Prostigmata" pageId= "0" pageNumber= "165" phylum= "Arthropoda" rank= "species" species= "hecmatanaensis" subGenus= "D." > R. (D.) hecmatanaensis</taxonomicName>
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Khanjani & Ueckermann in having two pairs of setae (
<emphasis id= "DBDB7D8499F68B3C91D55FD3E3BA0BBF" italics= "true" pageId= "0" pageNumber= "165" > d1</emphasis>
and
<emphasis id= "F28D44F2A9B77CBBEAA38A8CEBA06AAC" italics= "true" pageId= "0" pageNumber= "165" > e1</emphasis>
) on the interscutal membrane, two setae on the palp femora, and two small plates present posterolateral to the median prodorsal shield. However, the new species differs from
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<taxonomicName id= "F31CD9ACE0F30C6E88232424136C1ADE" class= "Arachnida" family= "Raphignathidae" higherTaxonomySource= "CoL" kingdom= "Animalia" lsidName= "R. (D.) hecmatanaensis" order= "Prostigmata" pageId= "0" pageNumber= "165" phylum= "Arthropoda" rank= "species" species= "hecmatanaensis" subGenus= "D." > R. (D.) hecmatanaensis</taxonomicName>
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in the absence of an endopodal shield (vs present), femur IV with two setae (vs three), and leg tarsus I with one solenidion (vs two solenidia).
</paragraph>
</subSubSection>
</treatment>
</document>
