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<mods:title id="5AE858B11DA2BD7CD722EF9CE4AF0A59">Taxonomy of the freshwater crabs of Costa Rica, with a revision of the genus Ptychophallus Smalley, 1964 (Crustacea: Decapoda: Pseudothelphusidae)</mods:title>
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<treatment id="710D87BB291C1602D6BDFC2FFCC5F89F" ID-DOI="http://doi.org/10.5281/zenodo.5491516" ID-GBIF-Taxon="119590155" ID-Zenodo-Dep="5491516" LSID="urn:lsid:plazi:treatment:710D87BB291C1602D6BDFC2FFCC5F89F" httpUri="http://treatment.plazi.org/id/710D87BB291C1602D6BDFC2FFCC5F89F" lastPageNumber="311" pageId="10" pageNumber="311">
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<paragraph id="F91B36AD291C1602D6BDFC2FFD79FC76" blockId="10.[151,680,908,967]" box="[151,680,908,934]" pageId="10" pageNumber="311">
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<taxonomicName id="3EA44D2E291C1602D6BDFC2FFD79FC76" ID-CoL="6WN77" authority="Pretzmann, 1965" authorityName="Pretzmann" authorityYear="1965" box="[151,680,908,934]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="goldmanni">
<emphasis id="CBD0EABF291C1602D6BDFC2FFD79FC76" bold="true" box="[151,680,908,934]" pageId="10" pageNumber="311">
<emphasis id="CBD0EABF291C1602D6BDFC2FFE1CFC76" bold="true" box="[151,461,908,934]" italics="true" pageId="10" pageNumber="311">Ptychophallus goldmanni</emphasis>
<bibRefCitation id="9D354B5C291C1602D7FEFC2FFD79FC76" author="Pretzmann" box="[468,680,908,934]" pageId="10" pageNumber="311" refString="Pretzmann, G. (1965) Vorlaufiger Bericht uber die Familie Pseudothelphusidae. Sitzungsberichte der Mathematisch- Naturwissenschaftlichen Klasse der Osterreichischen Akademie der Wissenschaften, Wien, 1, 1 - 11." type="journal article" year="1965">Pretzmann, 1965</bibRefCitation>
</emphasis>
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</paragraph>
<paragraph id="F91B36AD291C1602D6BDFC0DFE89FC17" blockId="10.[151,680,908,967]" box="[151,344,942,967]" pageId="10" pageNumber="311">
(
<figureCitation id="619F2A28291C1602D6B5FC0DFEF6FC17" box="[159,295,942,967]" captionStart="FIGURES 10 21" captionStartId="7.[151,264,1746,1769]" captionTargetBox="[159,1427,197,1724]" captionTargetId="figure@7.[157,1429,193,1725]" captionTargetPageId="7" captionText="FIGURES 10 21. Ptychophallus colombianus (Rathbun 1896): 10, left first gonopod, INPA 1845, caudo-mesial view; 11, same data, latero-cephalic view of distal part; 12, outline of left first gonopod, INPA 1654, caudo-mesial view; 13, same data, INPA 1659; 14, same data, NHMW 4001. Ptychophallus costaricensis Villalobos, 1974: 15, right first gonopod, paratype, UCR-MZ 719, latero-cephalic view of distal part; 16, same data, caudo-mesial view. Ptychophallus goldmanni Pretzmann, 1965: 17, left first gonopod, holotype, USNM 54044, caudo-mesial view; 18, same data, cephalic-apical view of distal part; 19, same data, latero-cephalic view of distal part. Ptychophallus kuna Campos &amp; Lemaitre, 1999: 20, left first gonopod, holotype, USNM 184340, latero-cephalic view of distal part; 21, same data, caudo-mesial view (dr, disto-caudal ridge; lp, lateral process; me, mesial process; mp, marginal process). Scale bars = 1 mm." httpUri="https://zenodo.org/record/235129/files/figure.png" pageId="10" pageNumber="311">Figs. 1719</figureCitation>
,
<figureCitation id="619F2A28291C1602D718FC0CFE9EFC17" box="[306,335,943,967]" captionStart="FIGURE 81" captionStartId="35.[151,250,1838,1860]" captionTargetBox="[154,1431,813,1813]" captionTargetId="figure@35.[151,1437,805,1817]" captionTargetPageId="35" captionText="FIGURE 81. Distribution map of Ptychophallus goldmanni Pretzmann, 1965, and P. kuna Campos &amp; Lemaitre, 1999." httpUri="https://zenodo.org/record/235139/files/figure.png" pageId="10" pageNumber="311">81</figureCitation>
)
</paragraph>
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<subSubSection id="B1BE6526291C1602D6BDFC55FC56FB76" pageId="10" pageNumber="311" type="reference_group">
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<treatmentCitationGroup id="D9B41183291C1602D6BDFC55FE5DFBFB" pageId="10" pageNumber="311">
<taxonomicName id="3EA44D2E291C1602D6BDFC55FCB9FBDC" ID-CoL="6WN77" authority="Pretzmann, 1965: 5" authorityName="Pretzmann" authorityPageNumber="5" authorityYear="1965" box="[151,872,1014,1036]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="goldmanni" subGenus="Microptychophallus">
<emphasis id="CBD0EABF291C1602D6BDFC55FEF9FBDC" box="[151,296,1014,1036]" italics="true" pageId="10" pageNumber="311">Ptychophallus</emphasis>
(
<emphasis id="CBD0EABF291C1602D717FC55FDD9FBDC" box="[317,520,1014,1036]" italics="true" pageId="10" pageNumber="311">Microptychophallus</emphasis>
)
<emphasis id="CBD0EABF291C1602D437FC55FD58FBDC" box="[541,649,1014,1036]" italics="true" pageId="10" pageNumber="311">goldmanni</emphasis>
<treatmentCitation id="780510BC291C1602D4BDFC54FCB9FBDC" author="Pretzmann" box="[663,872,1014,1036]" page="5" pageId="10" pageNumber="311" year="1965">
<bibRefCitation id="9D354B5C291C1602D4BDFC54FC9CFBDC" author="Pretzmann" box="[663,845,1014,1036]" pageId="10" pageNumber="311" refString="Pretzmann, G. (1965) Vorlaufiger Bericht uber die Familie Pseudothelphusidae. Sitzungsberichte der Mathematisch- Naturwissenschaftlichen Klasse der Osterreichischen Akademie der Wissenschaften, Wien, 1, 1 - 11." type="journal article" year="1965">Pretzmann, 1965</bibRefCitation>
: 5
</treatmentCitation>
</taxonomicName>
.—
<treatmentCitation id="780510BC291C1602D5ACFC55FBB0FBDC" author="Pretzmann" box="[902,1121,1014,1036]" page="21" pageId="10" pageNumber="311" year="1971">
<bibRefCitation id="9D354B5C291C1602D5ACFC55FBE8FBDC" author="Pretzmann" box="[902,1081,1014,1036]" pageId="10" pageNumber="311" refString="Pretzmann, G. (1971) Fortschritte in der Klassifizierung der Pseudothelphusidae. Anzeiger der Mathematisch Naturwissenschaftliche Klasse der Osterreichischen Akademie der Wissenschaften, Series 1, 179 (1 - 4), 12 - 24." type="journal article" year="1971">Pretzmann 1971</bibRefCitation>
: 21
</treatmentCitation>
.—
<treatmentCitation id="780510BC291C1602D254FC55FA8EFBDC" author="Pretzmann" box="[1150,1375,1014,1036]" page="90" pageId="10" pageNumber="311" year="1972">
<bibRefCitation id="9D354B5C291C1602D254FC55FAE3FBDC" author="Pretzmann" box="[1150,1330,1014,1036]" pageId="10" pageNumber="311" refString="Pretzmann, G. (1972) Die Pseudothelphusidae (Crustacea Brachyura). Zoologica, 42 (120), pt. 1, 1 - 182." type="journal article" year="1972">Pretzmann 1972</bibRefCitation>
: 90
</treatmentCitation>
, figs. 527529, 544546.
</treatmentCitationGroup>
</paragraph>
<paragraph id="F91B36AD291C1602D6BDFB97FDEFFB57" blockId="10.[151,1436,1014,1190]" pageId="10" pageNumber="311">
<treatmentCitationGroup id="D9B41183291C1602D6BDFB97FDEFFB57" pageId="10" pageNumber="311">
<taxonomicName id="3EA44D2E291C1602D6BDFB97FE4BFB9A" ID-CoL="6WN77" box="[151,410,1076,1098]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="goldmanni">
<emphasis id="CBD0EABF291C1602D6BDFB97FE4BFB9A" box="[151,410,1076,1098]" italics="true" pageId="10" pageNumber="311">Ptychophallus goldmanni</emphasis>
</taxonomicName>
<treatmentCitation id="780510BC291C1602D79EFB97FDADFB9A" author="Rodriguez" box="[436,636,1076,1098]" page="86" pageId="10" pageNumber="311" year="1982">
<bibRefCitation id="9D354B5C291C1602D79EFB97FD8BFB9A" author="Rodriguez" box="[436,602,1076,1098]" pageId="10" pageNumber="311" refString="Rodriguez, G. (1982) Les crabes d'eau douce d'Amerique. Familie des Pseudothelphusidae. ORSTOM, Paris, 223 pp. [Faune Tropicale 22]" type="book" year="1982">Rodriguez 1982</bibRefCitation>
: 86
</treatmentCitation>
.—
<treatmentCitation id="780510BC291C1602D4B1FB97FCA3FB9A" author="Villalobos" box="[667,882,1076,1098]" page="220" pageId="10" pageNumber="311" year="1982">
<bibRefCitation id="9D354B5C291C1602D4B1FB97FC93FB9A" author="Villalobos" box="[667,834,1076,1098]" pageId="10" pageNumber="311" refString="Villalobos, A. (1982) Decapoda. In: Hurlbert, S. H. &amp; Villalobos, A. (Eds.), Aquatic Biota of Mexico, Central America and the West Indies. San Diego State University, San Diego, pp. 215 - 239." type="book chapter" year="1982">Villalobos 1982</bibRefCitation>
: 220
</treatmentCitation>
(in list).—
<treatmentCitation id="780510BC291C1602D5CAFB97FB6AFB9A" author="Rodriguez" box="[992,1211,1076,1098]" page="304" pageId="10" pageNumber="311" year="1994">
<bibRefCitation id="9D354B5C291C1602D5CAFB97FB58FB9A" author="Rodriguez" box="[992,1161,1076,1098]" pageId="10" pageNumber="311" refString="Rodriguez, G. (1994) A revision of the type material of some species of Hypolobocera and Ptychophallus (Crustacea: Decapoda: Pseudothelphusidae) in the National Museum of Natural History, Washington, D. C., with descriptions of a new species and a new subspecies. Proceedings of the Biological Society of Washington, 107 (2), 296 - 307." type="journal article" year="1994">Rodríguez 1994</bibRefCitation>
: 304
</treatmentCitation>
, fig. 3FH.—
<treatmentCitation id="780510BC291C1602D36CFB97FE7DFBB8" author="Campos" page="560" pageId="10" pageNumber="311" year="1999">
<bibRefCitation id="9D354B5C291C1602D36CFB97FEAAFBB8" author="Campos" pageId="10" pageNumber="311" refString="Campos, M. R. &amp; Lemaitre, R. (1999) Two new freshwater crabs of the genus Ptychophallus Smalley, 1964 (Crustacea: Decapoda: Brachyura: Pseudothelphusidae) from Panama, with notes on the distribution of the genus. Proceedings of the Biological Society of Washington, 112 (3), 553 - 561." type="journal article" year="1999">Campos &amp; Lemaitre 1999</bibRefCitation>
: 560
</treatmentCitation>
(map).—
<treatmentCitation id="780510BC291C1602D426FBF1FCA1FBB8" author="Rodriguez" box="[524,880,1106,1128]" page="424" pageId="10" pageNumber="311" year="2000">
<bibRefCitation id="9D354B5C291C1602D426FBF1FCEFFBB8" author="Rodriguez" box="[524,830,1106,1128]" pageId="10" pageNumber="311" refString="Rodriguez, G. &amp; Hedstrom, I. (2000) The freswater crabs of the Barbilla National Park, Costa Rica (Crustacea: Brachyura: Pseudothelphusidae), with notes on the evolution of structures for spermatophore retention. Proceedings of the Biological Society of Washington, 113 (2), 420 - 425." type="journal article" year="2000">Rodríguez &amp; Hedström 2000</bibRefCitation>
: 424
</treatmentCitation>
, fig. 2E, J.—
<treatmentCitation id="780510BC291C1602D5D2FBF0FB13FBB8" author="Ng" box="[1016,1218,1106,1128]" page="176" pageId="10" pageNumber="311" year="2008">
<bibRefCitation id="9D354B5C291C1602D5D2FBF0FB40FBB8" author="Ng" box="[1016,1169,1106,1128]" pageId="10" pageNumber="311" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. The Raffles Bulletin of Zoology Supplement, 17, 1 - 286." type="journal article" year="2008">
Ng
<emphasis id="CBD0EABF291C1602D235FBF0FB85FBB8" box="[1055,1108,1106,1128]" italics="true" pageId="10" pageNumber="311">et al.</emphasis>
2008
</bibRefCitation>
: 176
</treatmentCitation>
(in list).—
<treatmentCitation id="780510BC291C1602D31BFBF1FE36FB57" author="Villalobos" page="297" pageId="10" pageNumber="311" year="2008">
<bibRefCitation id="9D354B5C291C1602D31BFBF1FE66FB57" author="Villalobos" pageId="10" pageNumber="311" refString="Villalobos Hiriart, J. L. &amp; Alvarez, F. (2008) Los cangrejos de la familia Pseudothelphusidae (Decapoda: Brachyura: Eubrachyura) de Mexico, con un apendice de las especies citadas para America hasta el 2006. In: Alvarez, F. &amp; Rodriguez- Almaraz, G. (Eds.), Crustaceos de Mexico: Estado actual de su Conocimiento. Universidad Autonoma de Nuevo Leon, Monterrey, Nuevo Leon, pp. 239 - 300. [Mexico]" type="book chapter" year="2008">Villalobos Hiriart &amp; Álvarez 2008</bibRefCitation>
: 297
</treatmentCitation>
(in list).
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</paragraph>
<paragraph id="F91B36AD291C1602D6BDFB33FC56FB76" blockId="10.[151,1436,1014,1190]" box="[151,903,1168,1190]" pageId="10" pageNumber="311">
? Not
<taxonomicName id="3EA44D2E291C1602D6FEFB33FE09FB76" ID-CoL="6WN77" box="[212,472,1168,1190]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="goldmanni">
<emphasis id="CBD0EABF291C1602D6FEFB33FE09FB76" box="[212,472,1168,1190]" italics="true" pageId="10" pageNumber="311">Ptychophallus goldmanni</emphasis>
</taxonomicName>
.—
<bibRefCitation id="9D354B5C291C1602D7DCFB33FD15FB76" author="Abele" box="[502,708,1168,1190]" pageId="10" pageNumber="311" refString="Abele, L. G. &amp; Kim, W. (1989) The decapod crustaceans of the Panama Canal. Smithsonian Contributions to Zoology, 482, 1 - 50. http: // dx. doi. org / 10.5479 / si. 00810282.482" type="journal article" year="1989">Abele &amp; Kim, 1989</bibRefCitation>
: 39 (see Remarks).
</paragraph>
</subSubSection>
<subSubSection id="B1BE6526291C1602D6BDFB71FE1AFADF" pageId="10" pageNumber="311" type="materials_examined">
<paragraph id="F91B36AD291C1602D6BDFB71FE1AFADF" blockId="10.[151,1437,1234,1871]" pageId="10" pageNumber="311">
<emphasis id="CBD0EABF291C1602D6BDFB71FE53FB3B" bold="true" box="[151,386,1234,1259]" pageId="10" pageNumber="311">Material examined</emphasis>
.
<collectingCountry id="81B3763D291C1602D7B0FB70FDC4FB3B" box="[410,533,1235,1259]" name="Panama" pageId="10" pageNumber="311">PANAMA</collectingCountry>
. Darién Province, Pacific drainage: male,
<typeStatus id="261F880F291C1602D20EFB71FB58FB3B" box="[1060,1161,1234,1259]" pageId="10" pageNumber="311" type="holotype">holotype</typeStatus>
,
<collectionCode id="9FB5AE68291C1602D2B7FB70FB3FFB3B" LSID="urn:lsid:biocol.org:col:34871" box="[1181,1262,1235,1259]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="311">USNM</collectionCode>
54044, Cana,
<date id="8D1A106D291C1602D6BDFB54FED6FADF" box="[151,263,1271,1295]" pageId="10" pageNumber="311" value="1912-05-24">24.v.1912</date>
, E.A. Goldmann.
</paragraph>
</subSubSection>
<subSubSection id="B1BE6526291C1602D6EDFAB9FC3DF9FF" pageId="10" pageNumber="311" type="description">
<paragraph id="F91B36AD291C1602D6EDFAB9FC3DF9FF" blockId="10.[151,1437,1234,1871]" pageId="10" pageNumber="311">
<emphasis id="CBD0EABF291C1602D6EDFAB9FE3CFAE3" bold="true" box="[199,493,1306,1331]" pageId="10" pageNumber="311">Description of gonopod</emphasis>
. Straight in mesiocaudal view, apex bent approximately 90° in laterocephalic direction. Marginal suture on mesial side, straight; marginal process rounded, not produced beyond caudal border of apex. Mesial process subtriangular, slightly downturned, its distal border rounded. Lateral process very poorly developed, with small lobe distally, not wider than lateral end of apex; lateral process with irregular border, nearly straight proximally, with few scattered, minute setae. Distocaudal ridge short, wide, subtriagular, weakly developed. Cephalic surface ending as wide, flat flange distally, forming distinct transversal sulcus between this flange and bent apex. Apex oblong, narrow; caudal border slightly convex, with small notch near lateral end. Field of apical spines moderately developed, directed towards laterocephalic side.
</paragraph>
</subSubSection>
<subSubSection id="B1BE6526291C1602D6EDF999FE84F96F" pageId="10" pageNumber="311" type="materials_examined">
<paragraph id="F91B36AD291C1602D6EDF999FE84F96F" blockId="10.[151,1437,1234,1871]" pageId="10" pageNumber="311">
<emphasis id="CBD0EABF291C1602D6EDF999FDD5F983" bold="true" box="[199,516,1594,1619]" pageId="10" pageNumber="311">
<typeStatus id="261F880F291C1602D6EDF999FEE5F983" box="[199,308,1594,1619]" pageId="10" pageNumber="311" type="holotype">Holotype</typeStatus>
and
<typeStatus id="261F880F291C1602D759F999FE77F983" box="[371,422,1594,1619]" pageId="10" pageNumber="311">type</typeStatus>
locality
</emphasis>
. Male, cw 26.4, cl 17.3,
<collectionCode id="9FB5AE68291C1602D50CF998FCA6F983" LSID="urn:lsid:biocol.org:col:34871" box="[806,887,1595,1619]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="311">USNM</collectionCode>
54044 (specimen not in good condition, mostly dismembered, abdomen detached and carapace partially detached from the body).
<collectingCountry id="81B3763D291C1602D21CF9FCFB43F9A6" box="[1078,1170,1631,1654]" name="Panama" pageId="10" pageNumber="311">Panama</collectingCountry>
, Darién Province, Cana (Pacific drainage); according to
<bibRefCitation id="9D354B5C291C1602D42AF921FD19F94B" author="Rodriguez" box="[512,712,1666,1691]" pageId="10" pageNumber="311" refString="Rodriguez, G. (1994) A revision of the type material of some species of Hypolobocera and Ptychophallus (Crustacea: Decapoda: Pseudothelphusidae) in the National Museum of Natural History, Washington, D. C., with descriptions of a new species and a new subspecies. Proceedings of the Biological Society of Washington, 107 (2), 296 - 307." type="journal article" year="1994">Rodríguez (1994)</bibRefCitation>
, Cana refers to Mount Cana, altitude
<quantity id="3E5C9B48291C1602D247F921FB6FF94B" box="[1133,1214,1666,1691]" metricMagnitude="3" metricUnit="m" metricValue="1.6149999999999998" pageId="10" pageNumber="311" unit="m" value="1615.0">1615m</quantity>
,
<geoCoordinate id="9C90506A291C1602D2E3F920FAFBF94B" box="[1225,1322,1667,1691]" direction="north" orientation="latitude" pageId="10" pageNumber="311" precision="925" value="7.8">07°48N</geoCoordinate>
<geoCoordinate id="9C90506A291C1602D31BF920FA49F94B" box="[1329,1432,1667,1691]" direction="west" orientation="longitude" pageId="10" pageNumber="311" precision="925" value="-77.53333">77°32W</geoCoordinate>
, Darién Province.
</paragraph>
</subSubSection>
<subSubSection id="B1BE6526291C1602D6EDF969FED1F8D7" pageId="10" pageNumber="311" type="distribution">
<paragraph id="F91B36AD291C1602D6EDF969FED1F8D7" blockId="10.[151,1437,1234,1871]" pageId="10" pageNumber="311">
<emphasis id="CBD0EABF291C1602D6EDF969FE88F933" bold="true" box="[199,345,1738,1763]" pageId="10" pageNumber="311">Distribution</emphasis>
. The species is so far known only from the
<typeStatus id="261F880F291C1602D56FF968FCA4F933" box="[837,885,1739,1763]" pageId="10" pageNumber="311">type</typeStatus>
locality, on the Pacific drainage, eastern
<collectingCountry id="81B3763D291C1602D369F968FA4DF932" box="[1347,1436,1739,1762]" name="Panama" pageId="10" pageNumber="311">Panama</collectingCountry>
(
<figureCitation id="619F2A28291C1602D6B5F94DFF22F8D7" box="[159,243,1774,1799]" captionStart="FIGURE 81" captionStartId="35.[151,250,1838,1860]" captionTargetBox="[154,1431,813,1813]" captionTargetId="figure@35.[151,1437,805,1817]" captionTargetPageId="35" captionText="FIGURE 81. Distribution map of Ptychophallus goldmanni Pretzmann, 1965, and P. kuna Campos &amp; Lemaitre, 1999." httpUri="https://zenodo.org/record/235139/files/figure.png" pageId="10" pageNumber="311">Fig. 81</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B1BE6526291C1602D6EDF8B1FCC5F89F" pageId="10" pageNumber="311" type="discussion">
<paragraph id="F91B36AD291C1602D6EDF8B1FCC5F89F" blockId="10.[151,1437,1234,1871]" pageId="10" pageNumber="311">
<emphasis id="CBD0EABF291C1602D6EDF8B1FEE5F8FB" bold="true" box="[199,308,1810,1835]" pageId="10" pageNumber="311">Remarks</emphasis>
.
<bibRefCitation id="9D354B5C291C1602D769F8B1FDE3F8FB" author="Abele" box="[323,562,1810,1835]" pageId="10" pageNumber="311" refString="Abele, L. G. &amp; Kim, W. (1989) The decapod crustaceans of the Panama Canal. Smithsonian Contributions to Zoology, 482, 1 - 50. http: // dx. doi. org / 10.5479 / si. 00810282.482" type="journal article" year="1989">Abele &amp; Kim (1989)</bibRefCitation>
recorded two males from the former
<collectingCountry id="81B3763D291C1602D5F5F8B0FBE9F8FA" box="[991,1080,1811,1834]" name="Panama" pageId="10" pageNumber="311">Panama</collectingCountry>
Canal Zone, but they probably belong to
<taxonomicName id="3EA44D2E291C1602D72DF894FE44F89F" box="[263,405,1846,1871]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="cocleensis">
<emphasis id="CBD0EABF291C1602D72DF894FE44F89F" box="[263,405,1846,1871]" italics="true" pageId="10" pageNumber="311">P. cocleensis</emphasis>
</taxonomicName>
(see Remarks under that species).
</paragraph>
</subSubSection>
</treatment>
</document>