122 lines
12 KiB
XML
122 lines
12 KiB
XML
|
<document id="68EDD25513F841EFD2CDDB9AAE640B47" ID-DOI="10.5281/zenodo.3374526" ID-GBIF-Dataset="2a40cf95-9ffb-4f3c-894b-cf53b8788a0a" ID-Zenodo-Dep="3374526" IM.metadata_requiresApprovalFor="plazi" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_requiresApprovalFor="plazi" IM.treatments_requiresApprovalFor="plazi" checkinTime="1566477704804" checkinUser="jeremy" docAuthor="Holtz, TR jr." docDate="2004" docId="012B87EDFF8ED81F4DBE25A9DC4DB6E3" docLanguage="en" docName="Holtz2004Tyrannosauroidea.pdf" docOrigin="The Dinosauria, University of California Press" docStyle="DocumentStyle{}" docTitle="Albertosaurus sarcophagus Osborn 1905" docType="treatment" docVersion="8" lastPageNumber="1" masterDocId="FD12FF95FF9AD80B4D3D2775DD29B37C" masterDocTitle="Tyrannosauroidea" masterLastPageNumber="136" masterPageNumber="111" pageNumber="1" updateTime="1698689907182" updateUser="ExternalLinkService">
|
||
|
|
<mods:mods id="31769DD69E7D5C3CC5F557635D170A98" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
|
|
<mods:titleInfo id="9F433540E234F9547DE44E248AC30633">
|
||
|
|
<mods:title id="DD64DA61B8F931CE5FC20D2B840726A6">Tyrannosauroidea</mods:title>
|
||
|
|
</mods:titleInfo>
|
||
|
|
<mods:name id="8D2DC7957B21B52CCBD68321E51ECA35" type="personal">
|
||
|
|
<mods:role id="DB2003BFCE95B7DCAA7BBD0C057F464E">
|
||
|
|
<mods:roleTerm id="79E11F9D0879169A27AADEC239DC1E12">Author</mods:roleTerm>
|
||
|
|
</mods:role>
|
||
|
|
<mods:namePart id="F01B7818F141D89BAEA20A6E820EBC76">Holtz, TR jr.</mods:namePart>
|
||
|
|
</mods:name>
|
||
|
|
<mods:typeOfResource id="9978AB71008A36431E79A14E91C940DD">text</mods:typeOfResource>
|
||
|
|
<mods:relatedItem id="D62C0970D3178E3EBF93C5C8643E352A" type="host">
|
||
|
|
<mods:originInfo id="2799E8F1B2C63F4AB9620D7AB3F214B5">
|
||
|
|
<mods:dateIssued id="641BF77585583111A6E4DFE8C17A639A">2004</mods:dateIssued>
|
||
|
|
<mods:publisher id="BF13C99D5C6AF2C96FAC14E4CCDCB648">University of California Press</mods:publisher>
|
||
|
|
</mods:originInfo>
|
||
|
|
<mods:name id="1D962DA82DF9BA93BA8A03E186CB715C" type="personal">
|
||
|
|
<mods:role id="13770358F23B3CC826175BB688053A17">
|
||
|
|
<mods:roleTerm id="EF97A6EF80C94698F2A4C89E8E6B07FD">Editor</mods:roleTerm>
|
||
|
|
</mods:role>
|
||
|
|
<mods:namePart id="71924F070C4B43C8F06DA467D8C58883">Weishampel DB</mods:namePart>
|
||
|
|
</mods:name>
|
||
|
|
<mods:name id="1E4C178C09275B675401AB1A7B270DC4" type="personal">
|
||
|
|
<mods:role id="ABF0D61AA0E59BC18A3DF4A3EF79919D">
|
||
|
|
<mods:roleTerm id="A9633E769150C1EA4457423376986EC8">Editor</mods:roleTerm>
|
||
|
|
</mods:role>
|
||
|
|
<mods:namePart id="58424FF87B7D85B1BBCB045C180651B4">Dodson P</mods:namePart>
|
||
|
|
</mods:name>
|
||
|
|
<mods:name id="ABD87F13343EF6D29D305D322F328B44" type="personal">
|
||
|
|
<mods:role id="82C4904D9CBA64A6203DC7F27EB35FB0">
|
||
|
|
<mods:roleTerm id="7C28A45BE71B7DC8928DFDE4B322242F">Editor</mods:roleTerm>
|
||
|
|
</mods:role>
|
||
|
|
<mods:namePart id="3C91873AF096D15001EE268E1850FE2A">Osmolska H</mods:namePart>
|
||
|
|
</mods:name>
|
||
|
|
<mods:titleInfo id="9208EE5F87479FA7C2365F660508DF17">
|
||
|
|
<mods:title id="A9EBF474078EF51942002D972AB4E84A">The Dinosauria</mods:title>
|
||
|
|
</mods:titleInfo>
|
||
|
|
<mods:part id="7684CB2FB9F95CA900C47A8C8777BA45">
|
||
|
|
<mods:extent id="757B4E36BFE915BB8020524F3297536D" unit="page">
|
||
|
|
<mods:start id="439ADC272A6C9BABB95FB8F29C5B0A1F">111</mods:start>
|
||
|
|
<mods:end id="CD809D270C117A61F778929123AFDE89">136</mods:end>
|
||
|
|
</mods:extent>
|
||
|
|
</mods:part>
|
||
|
|
</mods:relatedItem>
|
||
|
|
<mods:classification id="783F28A3F3A0DF6AF52AC7A91525D8D0">book chapter</mods:classification>
|
||
|
|
<mods:identifier id="4FF44E1943011345C5DD85BA40346857" type="DOI">10.5281/zenodo.3374526</mods:identifier>
|
||
|
|
<mods:identifier id="25E59F1242B80AC65F6FBA9250CEC138" type="GBIF-Dataset">2a40cf95-9ffb-4f3c-894b-cf53b8788a0a</mods:identifier>
|
||
|
|
<mods:identifier id="8AA93AB0A7C78CD9922AAEA904A60543" type="Zenodo-Dep">3374526</mods:identifier>
|
||
|
|
</mods:mods>
|
||
|
|
<treatment id="012B87EDFF8ED81F4DBE25A9DC4DB6E3" ID-DOI="http://doi.org/10.5281/zenodo.3483196" ID-GBIF-Taxon="163642056" ID-Zenodo-Dep="3483196" LSID="urn:lsid:plazi:treatment:012B87EDFF8ED81F4DBE25A9DC4DB6E3" httpUri="http://treatment.plazi.org/id/012B87EDFF8ED81F4DBE25A9DC4DB6E3" lastPageNumber="1" pageId="20" pageNumber="1">
|
||
|
|
<subSubSection id="C1986570FF8ED81F4DBE25A9DCADB193" box="[131,388,732,751]" pageId="20" pageNumber="1" type="nomenclature">
|
||
|
|
<paragraph id="893D36FBFF8ED81F4DBE25A9DCADB193" blockId="20.[99,716,100,1926]" box="[131,388,732,751]" pageId="20" pageNumber="1">
|
||
|
|
<taxonomicName id="4E824D78FF8ED81F4DBE25A9DCADB193" authorityName="Osborn" authorityYear="1905" box="[131,388,732,751]" class="Reptilia" family="Tyrannosauridae" genus="Albertosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="20" pageNumber="1" phylum="Chordata" rank="species" species="sarcophagus">
|
||
|
|
<emphasis id="BBF6EAE9FF8ED81F4DBE25A9DCADB193" box="[131,388,732,751]" italics="true" pageId="20" pageNumber="1">Albertosaurus sarcophagus</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
</paragraph>
|
||
|
|
</subSubSection>
|
||
|
|
<subSubSection id="C1986570FF8ED81F4CB025AEDC4DB6E3" pageId="20" pageNumber="1" type="discussion">
|
||
|
|
<paragraph id="893D36FBFF8ED81F4CB025AEDDDAB756" blockId="20.[99,716,100,1926]" pageId="20" pageNumber="1">
|
||
|
|
is known only from the lower Maastrichtian Horseshoe Canyon Formation of Alberta (Osborn 1905); no material referred to this taxon from outside this unit has yet demonstrated derived features of the species. Lambe (1904a) and Osborn (1906) described the skull, but much of the postcranial except for the cervical and caudal regions was later described under the name
|
||
|
|
<taxonomicName id="4E824D78FF8ED81F4C5724FDDFE1B0E6" authority="Parks, 1928" authorityName="Parks" authorityYear="1928" box="[362,712,903,923]" class="Reptilia" family="Tyrannosauridae" genus="Albertosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="20" pageNumber="1" phylum="Chordata" rank="species" species="arctunguis">
|
||
|
|
<emphasis id="BBF6EAE9FF8ED81F4C5724FDDF7BB0E6" box="[362,594,904,923]" italics="true" pageId="20" pageNumber="1">Albertosaurus arctunguis</emphasis>
|
||
|
|
Parks, 1928
|
||
|
|
</taxonomicName>
|
||
|
|
. The juvenile tyrannosaurid from the Horseshoe Canyon Formation referred to
|
||
|
|
<taxonomicName id="4E824D78FF8ED81F4C2124B4DC80B0A8" baseAuthorityName="Russell" baseAuthorityYear="1970" box="[284,425,961,980]" class="Reptilia" family="Tyrannosauridae" genus="Daspletosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="20" pageNumber="1" phylum="Chordata" rank="genus">
|
||
|
|
<emphasis id="BBF6EAE9FF8ED81F4C2124B4DC80B0A8" box="[284,425,961,980]" italics="true" pageId="20" pageNumber="1">Daspletosaurus</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
(a taxon otherwise unknown from the Maastrichtian) by Russell (1970a) is almost certainly a juvenile
|
||
|
|
<emphasis id="BBF6EAE9FF8ED81F4D88248FDC10B771" box="[181,313,1018,1037]" italics="true" pageId="20" pageNumber="1">
|
||
|
|
<taxonomicName id="4E824D78FF8ED81F4D88248FDC1DB771" baseAuthorityName="Parks" baseAuthorityYear="1933" box="[181,308,1018,1037]" class="Reptilia" family="Tyrannosauridae" genus="Albertosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="20" pageNumber="1" phylum="Chordata" rank="genus">Albertosaurus</taxonomicName>
|
||
|
|
.
|
||
|
|
</emphasis>
|
||
|
|
Additional material is described by Currie (2000, 2003b).
|
||
|
|
</paragraph>
|
||
|
|
<paragraph id="893D36FBFF8ED81F4DBE2341DC4DB6E3" blockId="20.[99,716,100,1926]" pageId="20" pageNumber="1">
|
||
|
|
<taxonomicName id="4E824D78FF8ED81F4DBE2341DDD6B73B" baseAuthorityName="Parks" baseAuthorityYear="1933" box="[131,255,1076,1095]" class="Reptilia" family="Tyrannosauridae" genus="Albertosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="20" pageNumber="1" phylum="Chordata" rank="genus">
|
||
|
|
<emphasis id="BBF6EAE9FF8ED81F4DBE2341DDD6B73B" box="[131,255,1076,1095]" italics="true" pageId="20" pageNumber="1">Albertosaurus</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
is comparable to (perhaps slightly smaller than)
|
||
|
|
<taxonomicName id="4E824D78FF8ED81F4D5E2324DDFFB718" authorityName="Lambe" authorityYear="1914" box="[99,214,1105,1124]" class="Reptilia" family="Tyrannosauridae" genus="Gorgosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="20" pageNumber="1" phylum="Chordata" rank="genus">
|
||
|
|
<emphasis id="BBF6EAE9FF8ED81F4D5E2324DDFFB718" box="[99,214,1105,1124]" italics="true" pageId="20" pageNumber="1">Gorgosaurus</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
in size. However, it is similar to
|
||
|
|
<taxonomicName id="4E824D78FF8ED81F4F2F2325DFB6B71F" baseAuthorityName="Russell" baseAuthorityYear="1970" box="[530,671,1104,1123]" class="Reptilia" family="Tyrannosauridae" genus="Daspletosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="20" pageNumber="1" phylum="Chordata" rank="genus">
|
||
|
|
<emphasis id="BBF6EAE9FF8ED81F4F2F2325DFB6B71F" box="[530,671,1104,1123]" italics="true" pageId="20" pageNumber="1">Daspletosaurus</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
and the Two Medicine tyrannosaurine in having dorsally oriented lacrimal horns. It shares with
|
||
|
|
<taxonomicName id="4E824D78FF8ED81F4CB023FFDF3EB7E0" authorityName="Osborn" authorityYear="1905" box="[397,535,1162,1180]" class="Reptilia" family="Tyrannosauridae" genus="Tyrannosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="20" pageNumber="1" phylum="Chordata" rank="genus">
|
||
|
|
<emphasis id="BBF6EAE9FF8ED81F4CB023FFDF3EB7E0" box="[397,535,1162,1180]" italics="true" pageId="20" pageNumber="1">Tyrannosaurus</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
and
|
||
|
|
<taxonomicName id="4E824D78FF8ED81F4F7423FFDF93B7E1" baseAuthorityName="Maleev" baseAuthorityYear="1974" box="[585,698,1162,1181]" class="Reptilia" family="Tyrannosauridae" genus="Tarbosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="20" pageNumber="1" phylum="Chordata" rank="genus">
|
||
|
|
<emphasis id="BBF6EAE9FF8ED81F4F7423FFDF93B7E1" box="[585,698,1162,1181]" italics="true" pageId="20" pageNumber="1">Tarbosaurus</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
a caudoventrally oriented occipital region, reduced basal tubera, two or more foramina on the lateral surface of the palatine, and (with
|
||
|
|
<taxonomicName id="4E824D78FF8ED81F4DA22395DC00B78E" authorityName="Osborn" authorityYear="1905" box="[159,297,1248,1266]" class="Reptilia" family="Tyrannosauridae" genus="Tyrannosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="20" pageNumber="1" phylum="Chordata" rank="genus">
|
||
|
|
<emphasis id="BBF6EAE9FF8ED81F4DA22395DC00B78E" box="[159,297,1248,1266]" italics="true" pageId="20" pageNumber="1">Tyrannosaurus</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
but not
|
||
|
|
<taxonomicName id="4E824D78FF8ED81F4CBD2395DCD8B78F" baseAuthorityName="Maleev" baseAuthorityYear="1974" box="[384,497,1248,1267]" class="Reptilia" family="Tyrannosauridae" genus="Tarbosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="20" pageNumber="1" phylum="Chordata" rank="genus">
|
||
|
|
<emphasis id="BBF6EAE9FF8ED81F4CBD2395DCD8B78F" box="[384,497,1248,1267]" italics="true" pageId="20" pageNumber="1">Tarbosaurus</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
) a caudally expanded scapula and large deltopectoral crest. In
|
||
|
|
<taxonomicName id="4E824D78FF8ED81F4CCE2389DF5BB673" baseAuthorityName="Parks" baseAuthorityYear="1933" box="[499,626,1276,1295]" class="Reptilia" family="Tyrannosauridae" genus="Albertosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="20" pageNumber="1" phylum="Chordata" rank="genus">
|
||
|
|
<emphasis id="BBF6EAE9FF8ED81F4CCE2389DF5BB673" box="[499,626,1276,1295]" italics="true" pageId="20" pageNumber="1">Albertosaurus</emphasis>
|
||
|
|
</taxonomicName>
|
||
|
|
each basisphenoid foramen lies within a distinct fossa. The rostral margin of the postorbital suborbital prong is jagged rather than smooth in
|
||
|
|
<emphasis id="BBF6EAE9FF8ED81F4DF12227DC7BB619" box="[204,338,1362,1381]" italics="true" pageId="20" pageNumber="1">
|
||
|
|
<taxonomicName id="4E824D78FF8ED81F4DF12227DC67B619" baseAuthorityName="Parks" baseAuthorityYear="1933" box="[204,334,1362,1381]" class="Reptilia" family="Tyrannosauridae" genus="Albertosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="20" pageNumber="1" phylum="Chordata" rank="genus">Albertosaurus</taxonomicName>
|
||
|
|
,
|
||
|
|
</emphasis>
|
||
|
|
suggesting the presence of a tendinous sheet beneath the orbit homologous to the fully ossified condition found in
|
||
|
|
<emphasis id="BBF6EAE9FF8ED81F4DD722F9DC4DB6E3" box="[234,356,1420,1439]" italics="true" pageId="20" pageNumber="1">
|
||
|
|
<taxonomicName id="4E824D78FF8ED81F4DD722F9DC49B6E3" authorityName="Lambe" authorityYear="1914" box="[234,352,1420,1439]" class="Reptilia" family="Tyrannosauridae" genus="Gorgosaurus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Dinosauria" pageId="20" pageNumber="1" phylum="Chordata" rank="genus">Gorgosaurus</taxonomicName>
|
||
|
|
.
|
||
|
|
</emphasis>
|
||
|
|
</paragraph>
|
||
|
|
</subSubSection>
|
||
|
|
</treatment>
|
||
|
|
</document>
|