<documentid="AD788AEADFA1BFAAA971375D5744F0CA"ID-CLB-Dataset="279456"ID-DOI="10.11646/zootaxa.5374.4.1"ID-GBIF-Dataset="6f75d877-d9e0-49de-8ce6-b739b532136a"ID-ISSN="1175-5326"ID-Zenodo-Dep="10158722"ID-ZooBank="EEFF13BD-664A-4FD8-9FED-4E4B21A475E4"IM.bibliography_approvedBy="felipe"IM.illustrations_approvedBy="felipe"IM.materialsCitations_approvedBy="jonas"IM.metadata_approvedBy="felipe"IM.taxonomicNames_approvedBy="jonas"IM.treatments_approvedBy="jonas"checkinTime="1700469135046"checkinUser="plazi"docAuthor="Ahrens, Dirk, Lukic, Daniel, Pham, Phu, Li, Wei & Liu, Wangang"docDate="2023"docId="03B73169BE7A2747D9E4085EFA7DFE58"docLanguage="en"docName="zootaxa.5374.4.1.pdf"docOrigin="Zootaxa 5374 (4)"docSource="https://mapress.com/zt/article/download/zootaxa.5374.4.1/52301"docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article"docStyleId="647186512141C8FC8976D5BCC54AEB7D"docStyleName="Zootaxa.2013-.journal_article"docStyleVersion="9"docTitle="Tetraserica hubleyi Ahrens 2023, new species"docType="treatment"docVersion="7"lastPageNumber="461"masterDocId="FF8E4911BE73274DD9730A19FFFAFFD1"masterDocTitle="Tetraserica Ahrens, 2004 of continental Southeast Asia: new records, new species and an updated key to species (Coleoptera: Scarabaeidae: Sericinae: Sericini)"masterLastPageNumber="486"masterPageNumber="451"pageNumber="460"updateTime="1732584714017"updateUser="ExternalLinkService"zenodo-license-document="CLOSED">
<mods:titleid="8314AA3D15F22CA785FCC724E0BD740E">Tetraserica Ahrens, 2004 of continental Southeast Asia: new records, new species and an updated key to species (Coleoptera: Scarabaeidae: Sericinae: Sericini)</mods:title>
<mods:affiliationid="E7F89D8486D4E626EF9ECB487BE9D1AD">Museum A. Koenig, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Adenauerallee 127, 53113 Bonn, Germany & University of Salzburg, Salzburg, Austria</mods:affiliation>
<mods:affiliationid="20707953BA894C8EEA33BEA241F13835">Museum A. Koenig, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Adenauerallee 127, 53113 Bonn, Germany & Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam</mods:affiliation>
<mods:affiliationid="A3C48F6B60DCF084B60C85F5AB636942">Institute of Earth and Environment, Chinese Academy of Sciences, Yanxiang Road 97 #, Yanta District, Xi’an 710061, P. R. China. & liweiv @ stumail. nwu. edu. cn; https: // orcid. org / 0009 - 0001 - 0195 - 7889</mods:affiliation>
<mods:affiliationid="FA2C6C9E2197EED409082BEEBF60ACAE">Museum A. Koenig, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Adenauerallee 127, 53113 Bonn, Germany & Institute of Earth and Environment, Chinese Academy of Sciences, Yanxiang Road 97 #, Yanta District, Xi’an 710061, P. R. China. & liuwangang @ ieecas. cn; https: // orcid. org / 0000 - 0003 - 4788 - 7967</mods:affiliation>
<figureCitationid="13259CFABE7A2744D9E40872FEFBFD54"box="[151,257,619,645]"captionStart="FIGURE 3"captionStartId="8.[152,255,1523,1547]"captionTargetBox="[162,1427,192,1488]"captionTargetId="figure-23@8.[151,1436,181,1499]"captionTargetPageId="8"captionText="FIGURE 3. A–D: Tetraserica hornburgi Ahrens, new species (holotype); E–H: T. hubleyi Ahrens, new species (holotype). A, E: aedeagus, left side lateral view; B, F: parameres, dorsal view; C, G: aedeagus, right side lateral view; D, H: habitus; I: hind leg. Scale: 0.5 mm. Habitus not to scale."figureDoi="http://doi.org/10.5281/zenodo.10158728"httpUri="https://zenodo.org/record/10158728/files/figure.png"pageId="9"pageNumber="460">Fig. 3E–I</figureCitation>
<materialsCitationid="3B768A22BE7A2744D8A008AAFC92FCC4"ID-GBIF-Occurrence="4442319388"collectingDate="1999-05-21"collectingDateMax="1999-05-31"collectingDateMin="1999-05-21"collectionCode="ROMC"collectorName="B. Hubley & A. Leslie"country="Vietnam"elevation="120"latitude="11.456944"location="Crocodile Lake"longLatPrecision="21"longitude="107.351944"pageId="9"pageNumber="460"specimenCount="1"stateProvince="Dong Nai"typeStatus="holotype">
Labroclypeus subtrapezoidal, wider than long, widest at base, lateral margins moderately convex and convergent to strongly rounded anterior angles, anterior margin weakly emarginate medially, margins moderately reflexed; surface weakly convex, moderately shiny, finely and densely punctate, glabrous; frontoclypeal suture indistinctly incised, flat and weakly curved medially; ocular canthus short and triangular, impunctate, with a single terminal seta. Smooth area anterior to eye three times as wide as long. Frons dull, with sparse, fine punctures, with two single erect setae beside each eye. Antenna yellowish, with ten antennomeres; club composed of four antennomeres in male, straight, 1.1 times as long as remaining antennomeres combined. Eyes large, ratio of diameter/interocular width: 0.81.
<paragraphid="8BA1807FBE7A2744D9B40E9EFCB7FAE0"blockId="9.[151,1437,690,2013]"pageId="9"pageNumber="460">Pronotum moderately wide and convex, widest at base, lateral margins evenly convex, strongly narrowed anteriorly towards sharp and slightly produced anterior angles, posterior angles convex. Anterior margin of pronotum slightly convex, with fine, complete marginal line. Surface finely and densely punctate, except minute setae glabrous, lateral and lateral anterior margins sparsely setose. Hypomeron not carinate. Scutellum triangular, finely and densely punctate, basal midline widely impunctate.</paragraph>
<paragraphid="8BA1807FBE7A2744D9B40F22FB99FA10"blockId="9.[151,1437,690,2013]"pageId="9"pageNumber="460">Elytra oblong, widest in posterior third, striae distinctly impressed, finely and moderately densely punctate, intervals weakly convex, with fine and dense punctures concentrated along striae, with very minute setae in punctures and a very few short setae on odd intervals; epipleural edge robust, ending at convex external apical angle of elytra, epipleura densely setose; apical border with a narrow fringe of microtrichomes (100x).</paragraph>
<paragraphid="8BA1807FBE7A2744D9B40FD2FB34F9A4"blockId="9.[151,1437,690,2013]"pageId="9"pageNumber="460">Ventral surface dull, finely and densely punctate, metasternum sparsely covered with fine, short, or very minute setae; metacoxa glabrous, with a few single setae laterally; abdominal sternites finely and densely punctuate, with a transverse row of coarse punctures, each bearing a robust seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.81. Pygidium weakly convex and dull, densely punctate, without smooth midline, almost glabrous, with a few longer setae along apical margin.</paragraph>
Legs wide; femora finely and sparsely punctate; metafemur wide and moderately shiny or dull, anterior margin acute; ventral posterior margin smooth and widened in apical half, dorsal posterior margin smooth, with a few short setae basally. Posterior margin at middle with a large, sharp tooth directed basally subequal to one third of maximum metafemur width (
<figureCitationid="13259CFABE7A2744D8180CF2FE46F8D4"box="[363,444,1771,1797]"captionStart="FIGURE 3"captionStartId="8.[152,255,1523,1547]"captionTargetBox="[162,1427,192,1488]"captionTargetId="figure-23@8.[151,1436,181,1499]"captionTargetPageId="8"captionText="FIGURE 3. A–D: Tetraserica hornburgi Ahrens, new species (holotype); E–H: T. hubleyi Ahrens, new species (holotype). A, E: aedeagus, left side lateral view; B, F: parameres, dorsal view; C, G: aedeagus, right side lateral view; D, H: habitus; I: hind leg. Scale: 0.5 mm. Habitus not to scale."figureDoi="http://doi.org/10.5281/zenodo.10158728"httpUri="https://zenodo.org/record/10158728/files/figure.png"pageId="9"pageNumber="460">Fig. 3I</figureCitation>
). Metatibia short and wide, widest at middle; ratio width/length: 1/3.0; basal group of dorsal spines of metatibia at first third, apical one at two thirds of metatibial length; ventral margin weakly convex and serrate, with four equidistant robust setae; apical margin bluntly and slightly concavely truncated towards ventral margin. Tarsomeres dorsally smooth, with fine, sparse setae ventrally, neither laterally nor dorsally carinate; metatarsomeres with a strongly serrated ridge ventrally and glabrous; first metatarsomere distinctly shorter than following two tarsomeres combined, one third of its length longer than dorsal tibial spine. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex.
<figureCitationid="13259CFABE792747D8370A8EFE43FF60"box="[324,441,151,177]"captionStart="FIGURE 3"captionStartId="8.[152,255,1523,1547]"captionTargetBox="[162,1427,192,1488]"captionTargetId="figure-23@8.[151,1436,181,1499]"captionTargetPageId="8"captionText="FIGURE 3. A–D: Tetraserica hornburgi Ahrens, new species (holotype); E–H: T. hubleyi Ahrens, new species (holotype). A, E: aedeagus, left side lateral view; B, F: parameres, dorsal view; C, G: aedeagus, right side lateral view; D, H: habitus; I: hind leg. Scale: 0.5 mm. Habitus not to scale."figureDoi="http://doi.org/10.5281/zenodo.10158728"httpUri="https://zenodo.org/record/10158728/files/figure.png"pageId="10"pageNumber="461">Fig. 3E–G</figureCitation>
. Habitus:
<figureCitationid="13259CFABE792747DB5E0A8EFD78FF60"box="[557,642,151,177]"captionStart="FIGURE 3"captionStartId="8.[152,255,1523,1547]"captionTargetBox="[162,1427,192,1488]"captionTargetId="figure-23@8.[151,1436,181,1499]"captionTargetPageId="8"captionText="FIGURE 3. A–D: Tetraserica hornburgi Ahrens, new species (holotype); E–H: T. hubleyi Ahrens, new species (holotype). A, E: aedeagus, left side lateral view; B, F: parameres, dorsal view; C, G: aedeagus, right side lateral view; D, H: habitus; I: hind leg. Scale: 0.5 mm. Habitus not to scale."figureDoi="http://doi.org/10.5281/zenodo.10158728"httpUri="https://zenodo.org/record/10158728/files/figure.png"pageId="10"pageNumber="461">Fig. 3H</figureCitation>
) and the shape of the parameres: the left paramere is distinctly shorter being subequal to the median apical phallobasal lamina and straight in dorsal view (instead of being curved externally and longer as in