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<mods:title id="B65C4AF4C20045C749B9A85454A9AC54">Revision of the genus Porcellionides Miers, 1877 (Isopoda: Porcellionidae) in the Ibero-Balearic region</mods:title>
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<mods:namePart id="8E7F7BA1B7128A87C0FE74F0182B0084">Cifuentes, Julio</mods:namePart>
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<mods:affiliation id="0B348F7D9FED2C9CEAD63BB9F975EC4F">Departamento de Biología (Zoología), Facultad de Ciencias, Universidad Autónoma de Madrid, 28049 Cantoblanco, Madrid, Spain.</mods:affiliation>
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<mods:namePart id="B3B11B1A66F6A9E834F3D70716E76C73">Da Silva, Luís P.</mods:namePart>
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<mods:affiliation id="535461C70681A3F0716AF3D233844B06">CIBIO, Centro de Investigação em Biodiversidade e Recursos Genéticos, InBIO Laboratório Associado, Campus de Vairão, Universidade do Porto, 4485 - 661 Vairão, Portugal. &amp; BIOPOLIS Program in Genomics, Biodiversity and Land Planning, CIBIO, Campus de Vairão, 4485 - 661 Vairão, Portugal.</mods:affiliation>
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<taxonomicName id="4C024D50FFA8FF89345EFD08FBEEFD5B" authority="(Vandel, 1953)" baseAuthorityName="Vandel" baseAuthorityYear="1953" box="[544,1043,671,698]" class="Malacostraca" family="Porcellionidae" genus="Porcellionides" kingdom="Animalia" order="Isopoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="lucasioides">
<emphasis id="B976EAC1FFA8FF89345EFD08FCA0FD5B" bold="true" box="[544,861,671,698]" italics="true" pageId="21" pageNumber="22">Porcellionides lucasioides</emphasis>
(
<bibRefCitation id="EF934B22FFA8FF893513FD08FBF6FD5B" author="Vandel A." box="[877,1035,671,698]" pageId="21" pageNumber="22" pagination="45 - 75" refId="ref25690" refString="Vandel A. 1953. Les isopodes terrestres des provinces d'Almeria et de Granada. Archivos del Instituto de Aclimatacion (Almeria) 1: 45 - 75." type="journal article" year="1953">Vandel, 1953</bibRefCitation>
)
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<subSubSection id="C3186558FFA8FF893446FD55FC01FD3C" box="[568,1020,706,733]" pageId="21" pageNumber="22" type="description">
<paragraph id="8BBD36D3FFA8FF893446FD55FC01FD3C" blockId="21.[544,1043,671,733]" box="[568,1020,706,733]" pageId="21" pageNumber="22">
<figureCitation id="13392A56FFA8FF893446FD55FD4CFD3C" box="[568,689,706,733]" captionStart="Fig" captionStartId="6.[189,232,1716,1742]" captionTargetBox="[213,1355,503,1668]" captionTargetId="figure-106@6.[189,1398,496,1681]" captionTargetPageId="6" captionText="Fig. 2. Relation between noduli laterales coordinates (following Vandel 1960) d/c (y-axis) and b/c (x-axis) multiplied by 100 for all studied species. A. Porcellionides cingendus (Kinahan, 1857). B. Porcellionides elegans(Pollo Zorita, 1982). C. Porcellionides fuscomarmoratus(Budde-Lund, 1885). D. Porcellionides glaber (Koch, 1856). E. Porcellionides hispanus (Vandel, 1953). F. Porcellionides lucasioides (Vandel, 1953). G. Porcellionides lusitanus (Vandel, 1946). H. Porcellionides molleri (Verhoeff, 1901). I. Porcellionides pruinosus (Brandt, 1833). J. Porcellionides sexfasciatus (BuddeLund, 1885). K. Porcellionides ibericus sp. nov. L. P. sexfasciatus and its previously considered subspecies: P. glaber (g), P. hispanus (h), P. lucasioides (l), P. lusitanus (lu), P. molleri (m) and P. sexfasciatus (s)." figureDoi="http://doi.org/10.5281/zenodo.12157016" httpUri="https://zenodo.org/record/12157016/files/figure.png" pageId="21" pageNumber="22">Figs 2F, L</figureCitation>
, 
<figureCitation id="13392A56FFA8FF8934C3FD54FD20FD3C" box="[701,733,707,733]" captionStart="Fig" captionStartId="7.[189,232,1786,1812]" captionTargetBox="[231,1365,1102,1731]" captionTargetId="figure-323@7.[189,1398,1089,1751]" captionTargetPageId="7" captionText="Fig. 3. Scale-setae. A. Porcellionides cingendus (Kinahan, 1857), ♂ (JC104). B. Porcellionides elegans (Pollo Zorita, 1982), ♂ (MNCN 20.04/7306). C. Porcellionides fuscomarmoratus (BuddeLund, 1885), ♀ (MNCN 20.04/8129). D. Porcellionides glaber (Koch, 1856), ♀ (MNCN 20.04/11490). E. Porcellionides hispanus (Vandel, 1953), ♂ (MNCN 20.04/7163). F. Porcellionides lucasioides (Vandel, 1953), ♂ (JC352). G. Porcellionides lusitanus (Vandel, 1946), ♂ (LPS661). H. Porcellionides molleri (Verhoeff, 1901), ♀ (LPS135). I. Porcellionides pruinosus (Brandt, 1833), ♀ (JC318). J. Porcellionides sexfasciatus (Budde-Lund, 1885), ♂ (LPS455a). Scale bar = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.12157020" httpUri="https://zenodo.org/record/12157020/files/figure.png" pageId="21" pageNumber="22">3F</figureCitation>
, 
<figureCitation id="13392A56FFA8FF893497FD54FCF4FD3C" box="[745,777,707,733]" captionStart="Fig" captionStartId="22.[189,232,1963,1989]" captionTargetBox="[262,1324,265,1924]" captionTargetId="figure-16@22.[262,1324,265,1924]" captionTargetPageId="22" captionText="Fig. 14. Porcellionides lucasioides (Vandel, 1953), ♂ (JC648). A.Habitus. B. Noduli laterales coordinates. C. Exopod I. D. Exopod II. Scale bars: A = 1 mm; C–D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12157105" httpUri="https://zenodo.org/record/12157105/files/figure.png" pageId="21" pageNumber="22">14</figureCitation>
, 
<figureCitation id="13392A56FFA8FF89356BFD55FC93FD3C" box="[789,878,706,733]" captionStart="Fig" captionStartId="23.[189,232,1752,1778]" captionTargetBox="[328,1257,291,1695]" captionTargetId="figure-11@23.[286,1302,265,1714]" captionTargetPageId="23" captionText="Fig. 15. Pleopods. A–D. Porcellionides lucasioides (Vandel, 1953). A–B. ♂ (JC648). A. Exopod I. B. Exopod II. C–D. ♀ (JC216). C. exopod I. D. Exopod II. E–H. Porcellionides lusitanus (Vandel, 1946). E–F. ♂ (LPS661). E. Exopod I. F. Exopod II. G–H. ♀ (LPS644). G. Exopod I. H. Exopod II. I–L. Porcellionides molleri (Verhoeff, 1901), ♂ (JC237). I. Exopod I. J. Exopod II. K. Exopod I. L. Exopod II.M–P. Porcellionides pruinosus(Brandt, 1833). M–N. ♂ (JC383). M. Exopod I.N. Exopod II. O–P. ♀ (JC42). O. Exopod I. P. Exopod II. Q–T. Porcellionides sexfasciatus (Budde-Lund, 1885). Q–R. ♂ (LPS455a). Q. Exopod I. R. Exopod II. S–T. ♀ (JC75). S. Exopod I. T. Exopod II. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12157127" httpUri="https://zenodo.org/record/12157127/files/figure.png" pageId="21" pageNumber="22">15A–D</figureCitation>
, 
<figureCitation id="13392A56FFA8FF893504FD55FC67FD3D" box="[890,922,706,732]" captionStart="Fig" captionStartId="24.[189,232,1963,1989]" captionTargetBox="[228,1362,925,1915]" captionTargetId="figure-206@24.[189,1398,917,1925]" captionTargetPageId="24" captionText="Fig. 16. Localization of the Ibero-Balearic region in Europe and the distribution of Porcellionides lucasioides (Vandel, 1953)." figureDoi="http://doi.org/10.5281/zenodo.12157137" httpUri="https://zenodo.org/record/12157137/files/figure.png" pageId="21" pageNumber="22">16</figureCitation>
, 
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<treatmentCitation id="0AA310C2FFA8FF8936C3FC93FC9AFCFE" author="Vandel A." box="[189,871,772,799]" page="52" pageId="21" pageNumber="22" year="1953">
<taxonomicName id="4C024D50FFA8FF8936C3FC93FC9AFCFE" ID-CoL="5JNQY" authority="Vandel, 1953: 52" authorityName="Vandel" authorityPageNumber="52" authorityYear="1953" box="[189,871,772,799]" class="Malacostraca" family="Porcellionidae" genus="Metoponorthus" kingdom="Animalia" order="Isopoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="subSpecies" species="sexfasciatus" subSpecies="lucasioides">
<emphasis id="B976EAC1FFA8FF8936C3FC93FD6BFCFF" box="[189,662,772,798]" italics="true" pageId="21" pageNumber="22">Metoponorthus sexfasciatus lucasioides</emphasis>
<bibRefCitation id="EF934B22FFA8FF8934E3FC93FC9AFCFE" author="Vandel A." box="[669,871,772,799]" pageId="21" pageNumber="22" pagination="45 - 75" refId="ref25690" refString="Vandel A. 1953. Les isopodes terrestres des provinces d'Almeria et de Granada. Archivos del Instituto de Aclimatacion (Almeria) 1: 45 - 75." type="journal article" year="1953">Vandel, 1953: 52</bibRefCitation>
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, fig. 2.
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<emphasis id="B976EAC1FFA8FF8936C3FCD2FE50FCBE" bold="true" box="[189,429,837,863]" pageId="21" pageNumber="22">Material examined</emphasis>
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leg.; cited by 
<bibRefCitation id="EF934B22FFA8FF893378FCE3FEE1FC50" author="Cifuentes J." pageId="21" pageNumber="22" pagination="133" refId="ref23773" refString="Cifuentes J. 2021 a. Los isopodos terrestres de Andalucia, Espana (Crustacea: Isopoda, Oniscidea). Graellsia 77 (1): e 133. https: // doi. org / 10.3989 / graellsia. 2021. v 77.276" type="journal article" year="2021">Cifuentes (2021a)</bibRefCitation>
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<specimenCode id="DBA49EA8FFA8FF893759FC00FDF3FC50" box="[295,526,919,945]" pageId="21" pageNumber="22">MNCN 20.04/7164</specimenCode>
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. – 
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<specimenCount id="9D04FD5AFFA8FF8934CBFC00FD04FC53" box="[693,761,919,946]" count="2" pageId="21" pageNumber="22" type="male">2 ♂♂</specimenCount>
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<collectorName id="26F75305FFA8FF89320DFC0FFB01FC50" box="[1139,1276,919,946]" pageId="21" pageNumber="22">J. Cifuentes</collectorName>
leg.; cited by 
<bibRefCitation id="EF934B22FFA8FF89369EFC2DFE4BFC35" author="Cifuentes J." box="[224,438,954,980]" pageId="21" pageNumber="22" pagination="133" refId="ref23773" refString="Cifuentes J. 2021 a. Los isopodos terrestres de Andalucia, Espana (Crustacea: Isopoda, Oniscidea). Graellsia 77 (1): e 133. https: // doi. org / 10.3989 / graellsia. 2021. v 77.276" type="journal article" year="2021">Cifuentes (2021a)</bibRefCitation>
; JC352
</materialsCitation>
• 
<materialsCitation id="3B6A3C8EFFA8FF89345FFC2DFE76FC16" ID-GBIF-Occurrence="4895508337" collectingDate="1985-04-03" collectorName="J. Cifuentes" country="Spain" location="Venta del Charco" pageId="21" pageNumber="22" specimenCount="17" specimenCount-female="17" stateProvince="Murcia">
<specimenCount id="9D04FD5AFFA8FF89345FFC2DFD8FFC34" box="[545,626,954,981]" count="17" pageId="21" pageNumber="22" type="female">17 ♀♀</specimenCount>
; same collection data as for preceding; cited by 
<bibRefCitation id="EF934B22FFA8FF8932E3FC2DFA8FFC35" author="Cifuentes J." box="[1181,1394,954,980]" pageId="21" pageNumber="22" pagination="133" refId="ref23773" refString="Cifuentes J. 2021 a. Los isopodos terrestres de Andalucia, Espana (Crustacea: Isopoda, Oniscidea). Graellsia 77 (1): e 133. https: // doi. org / 10.3989 / graellsia. 2021. v 77.276" type="journal article" year="2021">Cifuentes (2021a)</bibRefCitation>
; JC216. – 
<emphasis id="B976EAC1FFA8FF893750FC4AFE76FC16" bold="true" box="[302,395,989,1015]" pageId="21" pageNumber="22">Murcia</emphasis>
</materialsCitation>
• 
<materialsCitation id="3B6A3C8EFFA8FF8937DBFC49FB19FC16" ID-GBIF-Occurrence="4895508478" box="[421,1252,989,1016]" collectingDate="2022-05-14" collectorName="I. Escarabajal Bernabe" country="Spain" location="Cabo Tinoso" pageId="21" pageNumber="22" specimenCount="4" specimenCount-female="4" stateProvince="Cordoba">
<specimenCount id="9D04FD5AFFA8FF8937DBFC49FE14FC19" box="[421,489,990,1016]" count="4" pageId="21" pageNumber="22" type="female">4 ♀♀</specimenCount>
; 
<location id="8EDD6008FFA8FF893786FC4AFD6CFC16" LSID="urn:lsid:plazi:treatment:03AB87C5FFA8FF89345EFD08FBD4F850:8EDD6008FFA8FF893786FC4AFD6CFC16" box="[504,657,989,1015]" country="Spain" name="Cabo Tinoso" pageId="21" pageNumber="22" stateProvince="Cordoba">Cabo Tiñoso</location>
; 
<date id="FFBC1013FFA8FF8934E0FC49FCC2FC19" box="[670,831,990,1016]" pageId="21" pageNumber="22" value="2022-05-14">
<collectingDate id="EFF8E9FBFFA8FF8934E0FC49FCC2FC19" box="[670,831,990,1016]" pageId="21" pageNumber="22" value="2022-05-14">14 May 2022</collectingDate>
</date>
; 
<collectorName id="26F75305FFA8FF893535FC49FBABFC16" box="[843,1110,989,1016]" pageId="21" pageNumber="22">I. Escarabajal Bernabé</collectorName>
leg.; JC649
</materialsCitation>
• 
<materialsCitation id="3B6A3C8EFFA8FF893280FC49FD71FBFA" ID-GBIF-Occurrence="4895508463" collectingDate="2022-05-14" collectionCode="JC" collectorName="I. Escarabajal Bernabe" country="Spain" location="Cabo Tinoso" pageId="21" pageNumber="22" specimenCode="JC648" specimenCount="1" specimenCount-male="1" stateProvince="Cordoba">
<specimenCount id="9D04FD5AFFA8FF893280FC49FAD0FC19" box="[1278,1325,990,1016]" count="1" pageId="21" pageNumber="22" type="male">1 ♂</specimenCount>
; same collection data as for preceding; 
<specimenCode id="DBA49EA8FFA8FF893443FB96FD71FBFA" box="[573,652,1025,1051]" pageId="21" pageNumber="22">JC648</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3186558FFA8FF8936C3FBD5FAE0F977" pageId="21" pageNumber="22" type="discussion">
<paragraph id="8BBD36D3FFA8FF8936C3FBD5FECDFBBD" blockId="21.[189,304,1090,1116]" box="[189,304,1090,1116]" pageId="21" pageNumber="22">
<heading id="D0F581BFFFA8FF8936C3FBD5FECDFBBD" bold="true" box="[189,304,1090,1116]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="B976EAC1FFA8FF8936C3FBD5FECDFBBD" bold="true" box="[189,304,1090,1116]" pageId="21" pageNumber="22">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD36D3FFA8FF8936C3FBE6FD9EFA29" blockId="21.[187,1399,1136,1480]" pageId="21" pageNumber="22">
<bibRefCitation id="EF934B22FFA8FF8936C3FBE6FE66FB6A" author="Vandel A." box="[189,411,1136,1163]" pageId="21" pageNumber="22" pagination="45 - 75" refId="ref25690" refString="Vandel A. 1953. Les isopodes terrestres des provinces d'Almeria et de Granada. Archivos del Instituto de Aclimatacion (Almeria) 1: 45 - 75." type="journal article" year="1953">Vandel (1953: 52)</bibRefCitation>
originally classified this taxon as a subspecies of 
<taxonomicName id="4C024D50FFA8FF89326BFBE6FB3AFB6A" authorityName="Budde-Lund" authorityYear="1885" box="[1045,1223,1137,1163]" class="Malacostraca" family="Porcellionidae" genus="Metoponorthus" kingdom="Animalia" order="Isopoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="sexfasciatus">
<emphasis id="B976EAC1FFA8FF89326BFBE6FB3AFB6A" box="[1045,1223,1137,1163]" italics="true" pageId="21" pageNumber="22">P. sexfasciatus</emphasis>
</taxonomicName>
. He indicated distinctive characters, including pronounced granulations (
<figureCitation id="13392A56FFA8FF89350EFB03FC20FB4F" box="[880,989,1172,1198]" captionStart="Fig" captionStartId="22.[189,232,1963,1989]" captionTargetBox="[262,1324,265,1924]" captionTargetId="figure-16@22.[262,1324,265,1924]" captionTargetPageId="22" captionText="Fig. 14. Porcellionides lucasioides (Vandel, 1953), ♂ (JC648). A.Habitus. B. Noduli laterales coordinates. C. Exopod I. D. Exopod II. Scale bars: A = 1 mm; C–D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12157105" httpUri="https://zenodo.org/record/12157105/files/figure.png" pageId="21" pageNumber="22">Fig. 14A</figureCitation>
), few pores in the glandular fields, cephalon with a small median lobe, and medium-sized lateral lobes. In males, the carpus of the pereopods 1 and 2 possess dense setae, and the exopod I has a widened inner lobe at the base, terminating in a fine, narrow point (
<figureCitation id="13392A56FFA8FF8937D8FB6AFDE9FAF9" box="[422,532,1277,1304]" captionStart="Fig" captionStartId="22.[189,232,1963,1989]" captionTargetBox="[262,1324,265,1924]" captionTargetId="figure-16@22.[262,1324,265,1924]" captionTargetPageId="22" captionText="Fig. 14. Porcellionides lucasioides (Vandel, 1953), ♂ (JC648). A.Habitus. B. Noduli laterales coordinates. C. Exopod I. D. Exopod II. Scale bars: A = 1 mm; C–D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12157105" httpUri="https://zenodo.org/record/12157105/files/figure.png" pageId="21" pageNumber="22">Figs 14C</figureCitation>
, 
<figureCitation id="13392A56FFA8FF89345DFB6AFDA4FAF9" box="[547,601,1277,1304]" captionStart="Fig" captionStartId="23.[189,232,1752,1778]" captionTargetBox="[328,1257,291,1695]" captionTargetId="figure-11@23.[286,1302,265,1714]" captionTargetPageId="23" captionText="Fig. 15. Pleopods. A–D. Porcellionides lucasioides (Vandel, 1953). A–B. ♂ (JC648). A. Exopod I. B. Exopod II. C–D. ♀ (JC216). C. exopod I. D. Exopod II. E–H. Porcellionides lusitanus (Vandel, 1946). E–F. ♂ (LPS661). E. Exopod I. F. Exopod II. G–H. ♀ (LPS644). G. Exopod I. H. Exopod II. I–L. Porcellionides molleri (Verhoeff, 1901), ♂ (JC237). I. Exopod I. J. Exopod II. K. Exopod I. L. Exopod II.M–P. Porcellionides pruinosus(Brandt, 1833). M–N. ♂ (JC383). M. Exopod I.N. Exopod II. O–P. ♀ (JC42). O. Exopod I. P. Exopod II. Q–T. Porcellionides sexfasciatus (Budde-Lund, 1885). Q–R. ♂ (LPS455a). Q. Exopod I. R. Exopod II. S–T. ♀ (JC75). S. Exopod I. T. Exopod II. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12157127" httpUri="https://zenodo.org/record/12157127/files/figure.png" pageId="21" pageNumber="22">15A</figureCitation>
). The b/c and d/c coordinates are relatively low (
<figureCitation id="13392A56FFA8FF8932D9FB6AFADBFAF9" box="[1191,1318,1277,1304]" captionStart="Fig" captionStartId="6.[189,232,1716,1742]" captionTargetBox="[213,1355,503,1668]" captionTargetId="figure-106@6.[189,1398,496,1681]" captionTargetPageId="6" captionText="Fig. 2. Relation between noduli laterales coordinates (following Vandel 1960) d/c (y-axis) and b/c (x-axis) multiplied by 100 for all studied species. A. Porcellionides cingendus (Kinahan, 1857). B. Porcellionides elegans(Pollo Zorita, 1982). C. Porcellionides fuscomarmoratus(Budde-Lund, 1885). D. Porcellionides glaber (Koch, 1856). E. Porcellionides hispanus (Vandel, 1953). F. Porcellionides lucasioides (Vandel, 1953). G. Porcellionides lusitanus (Vandel, 1946). H. Porcellionides molleri (Verhoeff, 1901). I. Porcellionides pruinosus (Brandt, 1833). J. Porcellionides sexfasciatus (BuddeLund, 1885). K. Porcellionides ibericus sp. nov. L. P. sexfasciatus and its previously considered subspecies: P. glaber (g), P. hispanus (h), P. lucasioides (l), P. lusitanus (lu), P. molleri (m) and P. sexfasciatus (s)." figureDoi="http://doi.org/10.5281/zenodo.12157016" httpUri="https://zenodo.org/record/12157016/files/figure.png" pageId="21" pageNumber="22">Figs 2F, L</figureCitation>
, 
<figureCitation id="13392A56FFA8FF89334AFB69FA94FAF9" box="[1332,1385,1278,1304]" captionStart="Fig" captionStartId="22.[189,232,1963,1989]" captionTargetBox="[262,1324,265,1924]" captionTargetId="figure-16@22.[262,1324,265,1924]" captionTargetPageId="22" captionText="Fig. 14. Porcellionides lucasioides (Vandel, 1953), ♂ (JC648). A.Habitus. B. Noduli laterales coordinates. C. Exopod I. D. Exopod II. Scale bars: A = 1 mm; C–D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12157105" httpUri="https://zenodo.org/record/12157105/files/figure.png" pageId="21" pageNumber="22">14B</figureCitation>
). Additionally, it is noteworthy that the scale-setae possess a broad tip (
<figureCitation id="13392A56FFA8FF893271FAB6FB96FADA" box="[1039,1131,1313,1339]" captionStart="Fig" captionStartId="7.[189,232,1786,1812]" captionTargetBox="[231,1365,1102,1731]" captionTargetId="figure-323@7.[189,1398,1089,1751]" captionTargetPageId="7" captionText="Fig. 3. Scale-setae. A. Porcellionides cingendus (Kinahan, 1857), ♂ (JC104). B. Porcellionides elegans (Pollo Zorita, 1982), ♂ (MNCN 20.04/7306). C. Porcellionides fuscomarmoratus (BuddeLund, 1885), ♀ (MNCN 20.04/8129). D. Porcellionides glaber (Koch, 1856), ♀ (MNCN 20.04/11490). E. Porcellionides hispanus (Vandel, 1953), ♂ (MNCN 20.04/7163). F. Porcellionides lucasioides (Vandel, 1953), ♂ (JC352). G. Porcellionides lusitanus (Vandel, 1946), ♂ (LPS661). H. Porcellionides molleri (Verhoeff, 1901), ♀ (LPS135). I. Porcellionides pruinosus (Brandt, 1833), ♀ (JC318). J. Porcellionides sexfasciatus (Budde-Lund, 1885), ♂ (LPS455a). Scale bar = 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.12157020" httpUri="https://zenodo.org/record/12157020/files/figure.png" pageId="21" pageNumber="22">Fig. 3F</figureCitation>
). The male exopod of the pleopod II is triangular (
<figureCitation id="13392A56FFA8FF893464FAD3FD73FABF" box="[538,654,1348,1374]" captionStart="Fig" captionStartId="22.[189,232,1963,1989]" captionTargetBox="[262,1324,265,1924]" captionTargetId="figure-16@22.[262,1324,265,1924]" captionTargetPageId="22" captionText="Fig. 14. Porcellionides lucasioides (Vandel, 1953), ♂ (JC648). A.Habitus. B. Noduli laterales coordinates. C. Exopod I. D. Exopod II. Scale bars: A = 1 mm; C–D = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12157105" httpUri="https://zenodo.org/record/12157105/files/figure.png" pageId="21" pageNumber="22">Figs 14D</figureCitation>
, 
<figureCitation id="13392A56FFA8FF8934E1FAD3FD2EFABE" box="[671,723,1348,1375]" captionStart="Fig" captionStartId="23.[189,232,1752,1778]" captionTargetBox="[328,1257,291,1695]" captionTargetId="figure-11@23.[286,1302,265,1714]" captionTargetPageId="23" captionText="Fig. 15. Pleopods. A–D. Porcellionides lucasioides (Vandel, 1953). A–B. ♂ (JC648). A. Exopod I. B. Exopod II. C–D. ♀ (JC216). C. exopod I. D. Exopod II. E–H. Porcellionides lusitanus (Vandel, 1946). E–F. ♂ (LPS661). E. Exopod I. F. Exopod II. G–H. ♀ (LPS644). G. Exopod I. H. Exopod II. I–L. Porcellionides molleri (Verhoeff, 1901), ♂ (JC237). I. Exopod I. J. Exopod II. K. Exopod I. L. Exopod II.M–P. Porcellionides pruinosus(Brandt, 1833). M–N. ♂ (JC383). M. Exopod I.N. Exopod II. O–P. ♀ (JC42). O. Exopod I. P. Exopod II. Q–T. Porcellionides sexfasciatus (Budde-Lund, 1885). Q–R. ♂ (LPS455a). Q. Exopod I. R. Exopod II. S–T. ♀ (JC75). S. Exopod I. T. Exopod II. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12157127" httpUri="https://zenodo.org/record/12157127/files/figure.png" pageId="21" pageNumber="22">15B</figureCitation>
). Females exhibit a minor incision in the tracheal field on exopods I and II (
<figureCitation id="13392A56FFA8FF8937BCFAF0FDA8FA63" box="[450,597,1383,1410]" captionStart="Fig" captionStartId="23.[189,232,1752,1778]" captionTargetBox="[328,1257,291,1695]" captionTargetId="figure-11@23.[286,1302,265,1714]" captionTargetPageId="23" captionText="Fig. 15. Pleopods. A–D. Porcellionides lucasioides (Vandel, 1953). A–B. ♂ (JC648). A. Exopod I. B. Exopod II. C–D. ♀ (JC216). C. exopod I. D. Exopod II. E–H. Porcellionides lusitanus (Vandel, 1946). E–F. ♂ (LPS661). E. Exopod I. F. Exopod II. G–H. ♀ (LPS644). G. Exopod I. H. Exopod II. I–L. Porcellionides molleri (Verhoeff, 1901), ♂ (JC237). I. Exopod I. J. Exopod II. K. Exopod I. L. Exopod II.M–P. Porcellionides pruinosus(Brandt, 1833). M–N. ♂ (JC383). M. Exopod I.N. Exopod II. O–P. ♀ (JC42). O. Exopod I. P. Exopod II. Q–T. Porcellionides sexfasciatus (Budde-Lund, 1885). Q–R. ♂ (LPS455a). Q. Exopod I. R. Exopod II. S–T. ♀ (JC75). S. Exopod I. T. Exopod II. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.12157127" httpUri="https://zenodo.org/record/12157127/files/figure.png" pageId="21" pageNumber="22">Fig. 15C–D</figureCitation>
). Considering the aforementioned, we consider that 
<taxonomicName id="4C024D50FFA8FF8932AEFAFFFE96FA44" authority="(Vandel, 1953)" baseAuthorityName="Vandel" baseAuthorityYear="1953" class="Malacostraca" family="Porcellionidae" genus="Porcellionides" kingdom="Animalia" order="Isopoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="lucasioides">
<emphasis id="B976EAC1FFA8FF8932AEFAFFFA8BFA60" box="[1232,1398,1383,1409]" italics="true" pageId="21" pageNumber="22">P. lucasioides</emphasis>
(
<bibRefCitation id="EF934B22FFA8FF8936BBFA1DFE9EFA44" author="Vandel A." box="[197,355,1418,1445]" pageId="21" pageNumber="22" pagination="45 - 75" refId="ref25690" refString="Vandel A. 1953. Les isopodes terrestres des provinces d'Almeria et de Granada. Archivos del Instituto de Aclimatacion (Almeria) 1: 45 - 75." type="journal article" year="1953">Vandel, 1953</bibRefCitation>
)
</taxonomicName>
possesses sufficient distinct characters to be considered a valid species (
<tableCitation id="C6800368FFA8FF8932BEFA1DFAE5FA45" box="[1216,1304,1418,1444]" captionStart="Table 1" captionStartId="5.[185,255,713,741]" captionTargetPageId="5" captionText="Table 1. Summary of comparative characters for species within the genus Porcellionides Miers, 1877 found in the Ibero-Balearic region." httpUri="http://table.plazi.org/id/DF7D665BFFB8FF9936C7FD5EF8F9FD04" pageId="21" pageNumber="22" tableUuid="DF7D665BFFB8FF9936C7FD5EF8F9FD04">Table 1</tableCitation>
), rather than a subspecies of 
<taxonomicName id="4C024D50FFA8FF8937CEFA39FDA1FA29" authorityName="Budde-Lund" authorityYear="1885" box="[432,604,1454,1480]" class="Malacostraca" family="Porcellionidae" genus="Metoponorthus" kingdom="Animalia" order="Isopoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="sexfasciatus">
<emphasis id="B976EAC1FFA8FF8937CEFA39FDA1FA29" box="[432,604,1454,1480]" italics="true" pageId="21" pageNumber="22">P. sexfasciatus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BBD36D3FFA8FF8936C3FA78FAE0F977" blockId="21.[187,1399,1519,1686]" pageId="21" pageNumber="22">
In some specimens of both immature and adult 
<taxonomicName id="4C024D50FFA8FF8934A7FA67FC87F9E8" baseAuthorityName="Vandel" baseAuthorityYear="1953" box="[729,890,1519,1545]" class="Malacostraca" family="Porcellionidae" genus="Porcellionides" kingdom="Animalia" order="Isopoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="lucasioides">
<emphasis id="B976EAC1FFA8FF8934A7FA67FC87F9E8" box="[729,890,1519,1545]" italics="true" pageId="21" pageNumber="22">P. lucasioides</emphasis>
</taxonomicName>
, the inner tip of the exopod I is occasionally underdeveloped, which increases the potential for misidentification with 
<taxonomicName id="4C024D50FFA8FF893208F984FB1FF9CD" baseAuthorityName="C.Koch" baseAuthorityYear="1856" box="[1142,1250,1554,1580]" class="Malacostraca" family="Porcellionidae" genus="Porcellionides" kingdom="Animalia" order="Isopoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="glabe">
<emphasis id="B976EAC1FFA8FF893208F984FB1FF9CD" box="[1142,1250,1554,1580]" italics="true" pageId="21" pageNumber="22">P. glabe</emphasis>
</taxonomicName>
r. However, distinguishing between these two species is feasible, as 
<taxonomicName id="4C024D50FFA8FF893534F9A1FC11F9AE" baseAuthorityName="Vandel" baseAuthorityYear="1953" box="[842,1004,1589,1615]" class="Malacostraca" family="Porcellionidae" genus="Porcellionides" kingdom="Animalia" order="Isopoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="lucasioides">
<emphasis id="B976EAC1FFA8FF893534F9A1FC11F9AE" box="[842,1004,1589,1615]" italics="true" pageId="21" pageNumber="22">P. lucasioides</emphasis>
</taxonomicName>
has pronounced granulations and medium-sized lateral lobes on the cephalon, while 
<taxonomicName id="4C024D50FFA8FF893566F9CEFC8AF993" baseAuthorityName="C.Koch" baseAuthorityYear="1856" box="[792,887,1624,1650]" class="Malacostraca" family="Porcellionidae" genus="Porcellionides" kingdom="Animalia" order="Isopoda" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="glabe">
<emphasis id="B976EAC1FFA8FF893566F9CEFC8AF993" box="[792,887,1624,1650]" italics="true" pageId="21" pageNumber="22">P. glabe</emphasis>
</taxonomicName>
r either presents a smooth body or, in some specimens, may have sparse and very fine granulations on the cephalon, and lacks lateral lobes.
</paragraph>
</subSubSection>
<subSubSection id="C3186558FFA8FF8936C3F92AFBD4F850" pageId="21" pageNumber="22" type="distribution">
<paragraph id="8BBD36D3FFA8FF8936C3F92AFEA5F936" blockId="21.[189,344,1725,1751]" box="[189,344,1725,1751]" pageId="21" pageNumber="22">
<heading id="D0F581BFFFA8FF8936C3F92AFEA5F936" bold="true" box="[189,344,1725,1751]" fontSize="11" level="3" pageId="21" pageNumber="22" reason="3">
<emphasis id="B976EAC1FFA8FF8936C3F92AFEA5F936" bold="true" box="[189,344,1725,1751]" pageId="21" pageNumber="22">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD36D3FFA8FF8936C3F97BFE97F88E" blockId="21.[189,1399,1771,1903]" pageId="21" pageNumber="22">
Spanish providences of 
<emphasis id="B976EAC1FFA8FF893798F97CFDB3F8E7" bold="true" box="[486,590,1771,1798]" pageId="21" pageNumber="22">Almería</emphasis>
: Fondón, Laujar de Andarax (
<bibRefCitation id="EF934B22FFA8FF8935B4F97BFB9AF8E7" author="Vandel A." box="[970,1127,1771,1798]" pageId="21" pageNumber="22" pagination="45 - 75" refId="ref25690" refString="Vandel A. 1953. Les isopodes terrestres des provinces d'Almeria et de Granada. Archivos del Instituto de Aclimatacion (Almeria) 1: 45 - 75." type="journal article" year="1953">Vandel 1953</bibRefCitation>
) and Níjar (
<bibRefCitation id="EF934B22FFA8FF89337AF97BFEF7F8C8" author="Cifuentes J." pageId="21" pageNumber="22" pagination="133" refId="ref23773" refString="Cifuentes J. 2021 a. Los isopodos terrestres de Andalucia, Espana (Crustacea: Isopoda, Oniscidea). Graellsia 77 (1): e 133. https: // doi. org / 10.3989 / graellsia. 2021. v 77.276" type="journal article" year="2021">Cifuentes 2021a</bibRefCitation>
); 
<emphasis id="B976EAC1FFA8FF89375EF898FE9EF8CA" bold="true" box="[288,355,1807,1835]" pageId="21" pageNumber="22">Ávila</emphasis>
: Piedralaves (
<bibRefCitation id="EF934B22FFA8FF893470F898FD0BF8C8" author="Pollo Zorita A." box="[526,758,1807,1833]" pageId="21" pageNumber="22" pagination="191 - 196" refId="ref25055" refString="Pollo Zorita A. 1986 b. Oniscidos en suelos de bosques quemados de la zona del alto Tietar (Avila) (Isopoda, Crustacea). Graellsia 41: 191 - 196." type="journal article" year="1986">Pollo Zorita 1986b</bibRefCitation>
); 
<emphasis id="B976EAC1FFA8FF893571F899FC80F8C8" bold="true" box="[783,893,1806,1833]" pageId="21" pageNumber="22">Córdoba</emphasis>
: Morrón Grande (
<bibRefCitation id="EF934B22FFA8FF893227F898FB13F8C8" author="Garcia L." box="[1113,1262,1807,1833]" pageId="21" pageNumber="22" pagination="27 - 32" refId="ref24419" refString="Garcia L. 2019. Nuevos registros de Isopodos terrestres (Crustacea: Oniscidea) en Espana meridional (Andalucia y Murcia). Revista Sociedad Gaditana Historia Natural 13: 27 - 32." type="journal article" year="2019">Garcia 2019</bibRefCitation>
) and Venta del Charco (
<bibRefCitation id="EF934B22FFA8FF89372EF8A5FDE5F8AD" author="Cifuentes J." box="[336,536,1842,1868]" pageId="21" pageNumber="22" pagination="133" refId="ref23773" refString="Cifuentes J. 2021 a. Los isopodos terrestres de Andalucia, Espana (Crustacea: Isopoda, Oniscidea). Graellsia 77 (1): e 133. https: // doi. org / 10.3989 / graellsia. 2021. v 77.276" type="journal article" year="2021">Cifuentes 2021a</bibRefCitation>
); 
<emphasis id="B976EAC1FFA8FF893453F8A5FD76F8AD" bold="true" box="[557,651,1842,1868]" pageId="21" pageNumber="22">Cuenca</emphasis>
: Ruinas de Segóbriga (
<bibRefCitation id="EF934B22FFA8FF8935DFF8A5FB79F8AD" author="Pollo Zorita A." box="[929,1156,1842,1868]" pageId="21" pageNumber="22" pagination="173 - 189" refId="ref25027" refString="Pollo Zorita A. 1986 a. Oniscidos de la cuenca alta del rio Tajo (Isopoda, Crustacea). Graellsia 41: 173 - 189." type="journal article" year="1986">Pollo Zorita 1986a</bibRefCitation>
, 
<bibRefCitation id="EF934B22FFA8FF8932EFF8A5FB2DF8AC" author="Pollo Zorita A." box="[1169,1232,1842,1869]" pageId="21" pageNumber="22" refId="ref25090" refString="Pollo Zorita A. 2015. Estudio Taxonomico y Ecologico de los Isopodos Terrestres de la Cuenca Alta del Rio Tajo. PhD thesis, Universidad Complutense de Madrid, Spain." type="book" year="2015">2015</bibRefCitation>
); 
<emphasis id="B976EAC1FFA8FF893299F8A6FADCF8AD" bold="true" box="[1255,1313,1841,1868]" pageId="21" pageNumber="22">Jaén</emphasis>
: Peñas (
<bibRefCitation id="EF934B22FFA8FF8936B8F8C2FEA0F88E" author="Garcia L." box="[198,349,1877,1903]" pageId="21" pageNumber="22" pagination="27 - 32" refId="ref24419" refString="Garcia L. 2019. Nuevos registros de Isopodos terrestres (Crustacea: Oniscidea) en Espana meridional (Andalucia y Murcia). Revista Sociedad Gaditana Historia Natural 13: 27 - 32." type="journal article" year="2019">Garcia 2019</bibRefCitation>
).
</paragraph>
<paragraph id="8BBD36D3FFA8FF8936C3F800FBD4F850" blockId="21.[189,1065,1943,1969]" box="[189,1065,1943,1969]" pageId="21" pageNumber="22">
It is recorded for the first time in the Spanish province of 
<collectingRegion id="49C6F831FFA8FF89351EF800FC48F850" box="[864,949,1943,1969]" country="Spain" name="Murcia" pageId="21" pageNumber="22">Murcia</collectingRegion>
(
<figureCitation id="13392A56FFA8FF8935BAF800FBE1F850" box="[964,1052,1943,1969]" captionStart="Fig" captionStartId="24.[189,232,1963,1989]" captionTargetBox="[228,1362,925,1915]" captionTargetId="figure-206@24.[189,1398,917,1925]" captionTargetPageId="24" captionText="Fig. 16. Localization of the Ibero-Balearic region in Europe and the distribution of Porcellionides lucasioides (Vandel, 1953)." figureDoi="http://doi.org/10.5281/zenodo.12157137" httpUri="https://zenodo.org/record/12157137/files/figure.png" pageId="21" pageNumber="22">Fig. 16</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>