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<mods:title id="1028E6FFF8BA763F1317CAE2F81C5BBB">Taxonomic review of the genus Urostylis (Hemiptera: Heteroptera: Urostylididae) from the Korean Peninsula, with description of a new species</mods:title>
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<taxonomicName id="4C304D6BFFACFF82FF13C1CBF955FD0D" authority="Scott, 1874" authorityName="Scott" authorityYear="1874" box="[151,565,727,754]" class="Insecta" family="Urostylididae" genus="Urostylis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="annulicornis">
<emphasis id="B944EAFAFFACFF82FF13C1CBF955FD0D" bold="true" box="[151,565,727,754]" pageId="2" pageNumber="447">
<emphasis id="B944EAFAFFACFF82FF13C1CBFAC5FD0E" bold="true" box="[151,421,727,753]" italics="true" pageId="2" pageNumber="447">Urostylis annulicornis</emphasis>
<bibRefCitation id="EFA14B19FFACFF82FE28C1C4F955FD0D" box="[428,565,728,754]" pageId="2" pageNumber="447" refId="ref8525" refString="Scott, J. (1874) On a collection of Hemiptera Heteroptera from Japan. Descriptions of various new genera and species. Annals and Magazine of Natural History, Series 4, 14, 289 - 304, 360 - 365, 46 - 452." type="journal article">Scott, 1874</bibRefCitation>
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<paragraph id="8B8F36E8FFACFF82FF13C1E6F944FCEC" blockId="2.[151,565,727,787]" box="[151,548,762,787]" pageId="2" pageNumber="447">
(
<figureCitation id="130B2A6DFFACFF82FF1AC1E6FA7DFCEC" box="[158,285,762,787]" captionStart="FIGURE 1" captionStartId="3.[151,250,1355,1377]" captionTargetBox="[171,1408,185,1335]" captionTargetId="figure@3.[151,1436,173,1345]" captionTargetPageId="3" captionText="FIGURE 1. Dorsal habitus of Urostylis spp. A: U. koreana sp. nov.; B: U. annulicornis; C: U. lateralis; D: U. striicornis; E: U. trullata; F: U. westwoodii. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.1290375" httpUri="https://zenodo.org/record/1290375/files/figure.png" pageId="2" pageNumber="447">Figures 1B</figureCitation>
, 
<figureCitation id="130B2A6DFFACFF82FEAEC1E7FA0FFCED" box="[298,367,763,787]" captionStart="FIGURE 2" captionStartId="5.[151,250,1495,1517]" captionTargetBox="[217,1386,199,1462]" captionTargetId="figure@5.[175,1415,177,1479]" captionTargetPageId="5" captionText="FIGURE 2. Genital capsule of Urostylis spp in dorsal view and in lateral view. A, G: U. koreana sp. nov.; B, H: U. annulicornis; C, I: U. lateralis; D, J: U. striicornis; E, K: U. trullata; F, L: U. westwoodii; A–F: dorsal view; G–L: lateral view. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.1290377" httpUri="https://zenodo.org/record/1290377/files/figure.png" pageId="2" pageNumber="447">2B, H</figureCitation>
, 
<figureCitation id="130B2A6DFFACFF82FEFFC1E7FAFEFCEC" box="[379,414,763,787]" captionStart="FIGURE 3" captionStartId="7.[151,250,1018,1040]" captionTargetBox="[169,1403,323,980]" captionTargetId="figure@7.[151,1436,301,996]" captionTargetPageId="7" captionText="FIGURE 3. Genital process of Urostylis spp. in caudal view (process was illustrated in dark). A: U. koreana sp. nov.; B: U. annulicornis; C: U. lateralis; D: U. striicornis; E: U. trullata; F: U. westwoodii. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.1290379" httpUri="https://zenodo.org/record/1290379/files/figure.png" pageId="2" pageNumber="447">3B</figureCitation>
, 
<figureCitation id="130B2A6DFFACFF82FE2DC1E7FAACFCEC" box="[425,460,763,787]" captionStart="FIGURE 4" captionStartId="7.[151,250,1534,1556]" captionTargetBox="[161,1392,1134,1494]" captionTargetId="figure@7.[151,1436,1120,1512]" captionTargetPageId="7" captionText="FIGURE 4. Paramere of Urostylis spp. A: U. koreana sp. nov.; B: U. annulicornis; C: U. lateralis; D: U. striicornis; E: U. trullata; F: U. westwoodii. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.1290381" httpUri="https://zenodo.org/record/1290381/files/figure.png" pageId="2" pageNumber="447">4B</figureCitation>
, 
<figureCitation id="130B2A6DFFACFF82FE5CC1E6F97CFCEC" box="[472,540,762,787]" captionStart="FIGURE 5" captionStartId="9.[151,250,1209,1231]" captionTargetBox="[287,1294,189,1185]" captionTargetId="figure@9.[264,1323,163,1188]" captionTargetPageId="9" captionText="FIGURE 5. Female genitalia of Urostylis spp. A–E: female genital plates (with drawings); F–J: spermatheca; A, F: U. koreana sp. nov.; B, G: U. annulicornis; C, H: U. lateralis; D, I: U. striicornis; E, J: U. westwoodii. VII: seventh abdominal segment; gc8: gonocoxites 8; la8: laterotergites 8; la9: laterotergites 9." figureDoi="http://doi.org/10.5281/zenodo.1290383" httpUri="https://zenodo.org/record/1290383/files/figure.png" pageId="2" pageNumber="447">5B, G</figureCitation>
)
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<taxonomicName id="4C304D6BFFACFF82FF13C05EF949FCA7" authority="Scott, 1874: 361" authorityName="Scott" authorityPageNumber="361" authorityYear="1874" box="[151,553,834,856]" class="Insecta" family="Urostylididae" genus="Urostylis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="annulicornis">
<emphasis id="B944EAFAFFACFF82FF13C05EFA19FCA7" box="[151,377,834,856]" italics="true" pageId="2" pageNumber="447">Urostylis annulicornis</emphasis>
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: 361
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<taxonomicName id="4C304D6BFFACFF82FF13C07DF9BEFC88" authority="Reuter in Autran &amp; Reuter, 1888: 199" authorityName="Reuter in Autran &amp; Reuter" authorityPageNumber="199" authorityYear="1888" box="[151,734,865,887]" class="Insecta" family="Urostylididae" genus="Urostylis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="virescens">
<emphasis id="B944EAFAFFACFF82FF13C07DFA36FC88" box="[151,342,865,887]" italics="true" pageId="2" pageNumber="447">Urostylis virescens</emphasis>
Reuter in 
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: 199
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, 200 (syn. 
<treatmentCitation id="0A9110F9FFACFF82FCC9C07DFF41FC88" author="" box="[845,1057,865,887]" page="363" pageId="2" pageNumber="447">
<bibRefCitation id="EFA14B19FFACFF82FCC9C07DF88DFC88" box="[845,1005,865,887]" pageId="2" pageNumber="447" refId="ref8146" refString="Kerzhner, I. M. (1964) Materials on the synonymy of shieldbugs (Heteroptera Pentatomoidea) in the fauna of the USSR and adjacent countires. Entomologicheskoe Obozrenie, 43, 363 - 367. [in Russian]" type="journal article">Kerzhner, 1964</bibRefCitation>
: 363
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).
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</paragraph>
<paragraph id="8B8F36E8FFACFF82FF13C09CF830FC69" blockId="2.[151,1068,834,948]" box="[151,848,896,918]" pageId="2" pageNumber="447">
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<taxonomicName id="4C304D6BFFACFF82FF13C09CF952FC69" authority="Jakovlev, 1889: 74" authorityName="Jakovlev" authorityPageNumber="74" authorityYear="1889" box="[151,562,896,918]" class="Insecta" family="Urostylididae" genus="Urostylis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="geniculatus">
<emphasis id="B944EAFAFFACFF82FF13C09CFA0EFC69" box="[151,366,896,918]" italics="true" pageId="2" pageNumber="447">Urostylis geniculatus</emphasis>
<treatmentCitation id="0A9110F9FFACFF82FEF0C09CF952FC69" author="" box="[372,562,896,918]" page="74" pageId="2" pageNumber="447">
<bibRefCitation id="EFA14B19FFACFF82FEF0C09CF96FFC69" box="[372,527,896,918]" pageId="2" pageNumber="447" refId="ref8071" refString="Jakovlev, V. E. (1889) Hemiptera Heteroptera sibirica. Horae Societatis Entomologicae Rossicae, 23, 72 - 82." type="journal article">Jakovlev, 1889</bibRefCitation>
: 74
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</taxonomicName>
(syn. 
<treatmentCitation id="0A9110F9FFACFF82FDF5C09CF824FC69" author="" box="[625,836,896,918]" page="363" pageId="2" pageNumber="447">
<bibRefCitation id="EFA14B19FFACFF82FDF5C09CF871FC69" box="[625,785,896,918]" pageId="2" pageNumber="447" refId="ref8146" refString="Kerzhner, I. M. (1964) Materials on the synonymy of shieldbugs (Heteroptera Pentatomoidea) in the fauna of the USSR and adjacent countires. Entomologicheskoe Obozrenie, 43, 363 - 367. [in Russian]" type="journal article">Kerzhner, 1964</bibRefCitation>
: 363
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).
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</paragraph>
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<taxonomicName id="4C304D6BFFACFF82FF13C082F976FC4B" authority="Nonnaizab, 1984: 342" authorityName="Nonnaizab" authorityPageNumber="342" authorityYear="1984" box="[151,534,926,948]" class="Insecta" family="Urostylididae" genus="Urostylis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="adiai">
<emphasis id="B944EAFAFFACFF82FF13C082FA4DFC4B" box="[151,301,926,948]" italics="true" pageId="2" pageNumber="447">Urostylis adiai</emphasis>
<treatmentCitation id="0A9110F9FFACFF82FEB7C082F976FC4B" author="" box="[307,534,926,948]" page="342" pageId="2" pageNumber="447">
<bibRefCitation id="EFA14B19FFACFF82FEB7C082FA84FC4B" box="[307,484,926,948]" pageId="2" pageNumber="447" refId="ref8261" refString="Nonnaizab (1984) New and little known species of Pentatomoidea from Inner Mongolia and Sinkiang, China (Hemiptera: Urostylidae). Acta Entomologica Sinica, 27, 342 - 344 [in Chinese, English summary]." type="journal article">Nonnaizab, 1984</bibRefCitation>
: 342
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, 344 (syn. 
<treatmentCitation id="0A9110F9FFACFF82FD01C083F844FC4B" author="" box="[645,804,926,948]" page="294" pageId="2" pageNumber="447">
<bibRefCitation id="EFA14B19FFACFF82FD01C083F990FC4B" box="[645,752,926,948]" pageId="2" pageNumber="447" refId="ref8302" refString="Ren, S. Z. (1997) Hemiptera: Urostylidae. In: Yang, X. K. (Ed.), Insects of the Three Gorge Reservoir area of Yangtze River. Chongqing Publishing House, Chongqing, pp. 291 - 300." type="book chapter">Ren, 1997</bibRefCitation>
: 294
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).
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</paragraph>
</subSubSection>
<subSubSection id="C32A6563FFACFF82FF13C0FDFBA8FB75" pageId="2" pageNumber="447" type="diagnosis">
<paragraph id="8B8F36E8FFACFF82FF13C0FDFBA8FB75" blockId="2.[151,1437,993,1990]" pageId="2" pageNumber="447">
<emphasis id="B944EAFAFFACFF82FF13C0FDFA41FC05" bold="true" box="[151,289,993,1018]" pageId="2" pageNumber="447">DIAGNOSIS:</emphasis>
Pronotum, scutellum and hemelytra with dark puncture; membrane with distinct dark markings (
<figureCitation id="130B2A6DFFACFF82FF1AC719FA73FBE1" box="[158,275,1029,1054]" captionStart="FIGURE 1" captionStartId="3.[151,250,1355,1377]" captionTargetBox="[171,1408,185,1335]" captionTargetId="figure@3.[151,1436,173,1345]" captionTargetPageId="3" captionText="FIGURE 1. Dorsal habitus of Urostylis spp. A: U. koreana sp. nov.; B: U. annulicornis; C: U. lateralis; D: U. striicornis; E: U. trullata; F: U. westwoodii. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.1290375" httpUri="https://zenodo.org/record/1290375/files/figure.png" pageId="2" pageNumber="447">Figure 1B</figureCitation>
); lateral margin of genital capsule rounded, apex relatively thick with densely long setae in dorsal view (
<figureCitation id="130B2A6DFFACFF82FF1AC735FA74FBBD" box="[158,276,1065,1090]" captionStart="FIGURE 2" captionStartId="5.[151,250,1495,1517]" captionTargetBox="[217,1386,199,1462]" captionTargetId="figure@5.[175,1415,177,1479]" captionTargetPageId="5" captionText="FIGURE 2. Genital capsule of Urostylis spp in dorsal view and in lateral view. A, G: U. koreana sp. nov.; B, H: U. annulicornis; C, I: U. lateralis; D, J: U. striicornis; E, K: U. trullata; F, L: U. westwoodii; A–F: dorsal view; G–L: lateral view. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.1290377" httpUri="https://zenodo.org/record/1290377/files/figure.png" pageId="2" pageNumber="447">Figure 2B</figureCitation>
); median process of genital capsule short, not reaching to dorsal wall of capsule in lateral view (
<figureCitation id="130B2A6DFFACFF82FAD5C735FBDCFB99" captionStart="FIGURE 2" captionStartId="5.[151,250,1495,1517]" captionTargetBox="[217,1386,199,1462]" captionTargetId="figure@5.[175,1415,177,1479]" captionTargetPageId="5" captionText="FIGURE 2. Genital capsule of Urostylis spp in dorsal view and in lateral view. A, G: U. koreana sp. nov.; B, H: U. annulicornis; C, I: U. lateralis; D, J: U. striicornis; E, K: U. trullata; F, L: U. westwoodii; A–F: dorsal view; G–L: lateral view. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.1290377" httpUri="https://zenodo.org/record/1290377/files/figure.png" pageId="2" pageNumber="447">Figure 2H</figureCitation>
), process rectanglar and broader apically (
<figureCitation id="130B2A6DFFACFF82FD1CC751F86DFB99" box="[664,781,1101,1126]" captionStart="FIGURE 3" captionStartId="7.[151,250,1018,1040]" captionTargetBox="[169,1403,323,980]" captionTargetId="figure@7.[151,1436,301,996]" captionTargetPageId="7" captionText="FIGURE 3. Genital process of Urostylis spp. in caudal view (process was illustrated in dark). A: U. koreana sp. nov.; B: U. annulicornis; C: U. lateralis; D: U. striicornis; E: U. trullata; F: U. westwoodii. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.1290379" httpUri="https://zenodo.org/record/1290379/files/figure.png" pageId="2" pageNumber="447">Figure 3B</figureCitation>
); paramere curved subapically, its apex bifurcated (
<figureCitation id="130B2A6DFFACFF82FAD5C751FBDBFB75" captionStart="FIGURE 4" captionStartId="7.[151,250,1534,1556]" captionTargetBox="[161,1392,1134,1494]" captionTargetId="figure@7.[151,1436,1120,1512]" captionTargetPageId="7" captionText="FIGURE 4. Paramere of Urostylis spp. A: U. koreana sp. nov.; B: U. annulicornis; C: U. lateralis; D: U. striicornis; E: U. trullata; F: U. westwoodii. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.1290381" httpUri="https://zenodo.org/record/1290381/files/figure.png" pageId="2" pageNumber="447">Figure 4B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32A6563FFACFF83FF43C789FA27F8C7" lastPageId="3" lastPageNumber="448" pageId="2" pageNumber="447" type="description">
<paragraph id="8B8F36E8FFACFF82FF43C789FFA3F911" blockId="2.[151,1437,993,1990]" pageId="2" pageNumber="447">
<emphasis id="B944EAFAFFACFF82FF43C789FA0FFB51" bold="true" box="[199,367,1173,1198]" pageId="2" pageNumber="447">DESCRIPTION:</emphasis>
<emphasis id="B944EAFAFFACFF82FEFBC789FAA2FB52" box="[383,450,1173,1197]" italics="true" pageId="2" pageNumber="447">Male:</emphasis>
Body oval and elongated, length 12.30–13.26. COLOR. mostly greenish. 
<emphasis id="B944EAFAFFACFF82FAD7C789FEFCFB51" bold="true" box="[1363,1436,1173,1198]" pageId="2" pageNumber="447">Head:</emphasis>
generally greenish, sometimes reddish; antennae almost brown, first segment with dark line laterally, third segment dark brown, 1/2 apical part of fourth and fifth segment dark brown; rostrum green, apex of fourth rostral segment dark brown. 
<emphasis id="B944EAFAFFACFF82FEADC61DFAEAFAE5" bold="true" box="[297,394,1281,1306]" pageId="2" pageNumber="447">Thorax:</emphasis>
pronotum entirely green except for pale anterolateral margin, with densely dark punctures, anterior part of pronotum with dark line laterally; scutellum entirely greenish, with dark punctures; hemelytra mostly greenish with entirely dark punctures; membrane transparent tinged with brownish, with dark V-shaped markings; legs almost green, sometimes reddish; base of tibia with dark band, apical part brown; third tarsal segment brown. 
<emphasis id="B944EAFAFFACFF82FED4C68DFAADFA55" bold="true" box="[336,461,1425,1450]" pageId="2" pageNumber="447">Abdomen:</emphasis>
entirely green. SURFACE AND VESTITURE: Body slightly glossy, with distinct dark punctures, covered with silvery short pubescences; antenna with densely short pubescences; legs with setae. STRUCTURE. 
<emphasis id="B944EAFAFFACFF82FECEC6C5FAF3FA0D" bold="true" box="[330,403,1497,1522]" pageId="2" pageNumber="447">Head:</emphasis>
width of head longer than length, mandibular plate projected as long as clypeus dorsally; length of antennae longer than length of body, first segment longer than 2 times length of head; rostrum not reaching to midcoxae. 
<emphasis id="B944EAFAFFACFF82FE24C53DF962F9C5" bold="true" box="[416,514,1569,1594]" pageId="2" pageNumber="447">Thorax:</emphasis>
anterior and posterior margin of pronotum rounded; mesal length of scutellum longer than width; lateral margin of hemelytra rounded; legs generally long; femur distinctly thick; first tarsal segment as long as third segment. 
<emphasis id="B944EAFAFFACFF82FDA4C575F9FDF97D" bold="true" box="[544,669,1641,1666]" pageId="2" pageNumber="447">Abdomen:</emphasis>
margin rounded, not reaching to apex of membrane. GENITALIA: lateral margin of genital capsule rounded, apex relatively thick with densely long setae in dorsal view (
<figureCitation id="130B2A6DFFACFF82FA9CC591FEEEF959" box="[1304,1422,1677,1702]" captionStart="FIGURE 2" captionStartId="5.[151,250,1495,1517]" captionTargetBox="[217,1386,199,1462]" captionTargetId="figure@5.[175,1415,177,1479]" captionTargetPageId="5" captionText="FIGURE 2. Genital capsule of Urostylis spp in dorsal view and in lateral view. A, G: U. koreana sp. nov.; B, H: U. annulicornis; C, I: U. lateralis; D, J: U. striicornis; E, K: U. trullata; F, L: U. westwoodii; A–F: dorsal view; G–L: lateral view. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.1290377" httpUri="https://zenodo.org/record/1290377/files/figure.png" pageId="2" pageNumber="447">Figure 2B</figureCitation>
); process of genital capsule short, not reaching to height of capsule in lateral view (
<figureCitation id="130B2A6DFFACFF82FBB1C5ADFFCEF935" box="[1077,1198,1713,1738]" captionStart="FIGURE 2" captionStartId="5.[151,250,1495,1517]" captionTargetBox="[217,1386,199,1462]" captionTargetId="figure@5.[175,1415,177,1479]" captionTargetPageId="5" captionText="FIGURE 2. Genital capsule of Urostylis spp in dorsal view and in lateral view. A, G: U. koreana sp. nov.; B, H: U. annulicornis; C, I: U. lateralis; D, J: U. striicornis; E, K: U. trullata; F, L: U. westwoodii; A–F: dorsal view; G–L: lateral view. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.1290377" httpUri="https://zenodo.org/record/1290377/files/figure.png" pageId="2" pageNumber="447">Figure 2H</figureCitation>
), process rectangular and broader apically (
<figureCitation id="130B2A6DFFACFF82FE0EC5C9FA9FF911" box="[394,511,1749,1774]" captionStart="FIGURE 3" captionStartId="7.[151,250,1018,1040]" captionTargetBox="[169,1403,323,980]" captionTargetId="figure@7.[151,1436,301,996]" captionTargetPageId="7" captionText="FIGURE 3. Genital process of Urostylis spp. in caudal view (process was illustrated in dark). A: U. koreana sp. nov.; B: U. annulicornis; C: U. lateralis; D: U. striicornis; E: U. trullata; F: U. westwoodii. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.1290379" httpUri="https://zenodo.org/record/1290379/files/figure.png" pageId="2" pageNumber="447">Figure 3B</figureCitation>
); paramere curved subapically, its apex bifurcated (
<figureCitation id="130B2A6DFFACFF82FBC4C5C9FFD6F911" box="[1088,1206,1749,1774]" captionStart="FIGURE 4" captionStartId="7.[151,250,1534,1556]" captionTargetBox="[161,1392,1134,1494]" captionTargetId="figure@7.[151,1436,1120,1512]" captionTargetPageId="7" captionText="FIGURE 4. Paramere of Urostylis spp. A: U. koreana sp. nov.; B: U. annulicornis; C: U. lateralis; D: U. striicornis; E: U. trullata; F: U. westwoodii. Scale bar: 1mm." figureDoi="http://doi.org/10.5281/zenodo.1290381" httpUri="https://zenodo.org/record/1290381/files/figure.png" pageId="2" pageNumber="447">Figure 4B</figureCitation>
).
</paragraph>
<paragraph id="8B8F36E8FFACFF82FF43C5E5F98EF839" blockId="2.[151,1437,993,1990]" pageId="2" pageNumber="447">
<emphasis id="B944EAFAFFACFF82FF43C5E5FA7BF8EE" box="[199,283,1785,1809]" italics="true" pageId="2" pageNumber="447">Female</emphasis>
: Body oval and elongated, length 13.28–13.34. COLOR: as in male. SURFACE AND VESTITURE: as in male. STRUCTURE: as in male except for genital segment. GENITALIA: lateral seventh abdominal segment flat, central seventh abdominal segment concave and convex at center; gonocoxites 8 large and divergent, extending the same length with laterotergites 9; laterotergites 8 small and compact, shorter than gonocoxite 8 and laterotergite 9; laterotergites 9 slightly visible, touching at base but divergent at apex (
<figureCitation id="130B2A6DFFACFF82FB04C495FF98F85D" box="[1152,1272,1929,1954]" captionStart="FIGURE 5" captionStartId="9.[151,250,1209,1231]" captionTargetBox="[287,1294,189,1185]" captionTargetId="figure@9.[264,1323,163,1188]" captionTargetPageId="9" captionText="FIGURE 5. Female genitalia of Urostylis spp. A–E: female genital plates (with drawings); F–J: spermatheca; A, F: U. koreana sp. nov.; B, G: U. annulicornis; C, H: U. lateralis; D, I: U. striicornis; E, J: U. westwoodii. VII: seventh abdominal segment; gc8: gonocoxites 8; la8: laterotergites 8; la9: laterotergites 9." figureDoi="http://doi.org/10.5281/zenodo.1290383" httpUri="https://zenodo.org/record/1290383/files/figure.png" pageId="2" pageNumber="447">Figure 5B</figureCitation>
); spermatheca curved and robust, sclerotized area short (
<figureCitation id="130B2A6DFFACFF82FDEEC4B1F982F839" box="[618,738,1965,1990]" captionStart="FIGURE 5" captionStartId="9.[151,250,1209,1231]" captionTargetBox="[287,1294,189,1185]" captionTargetId="figure@9.[264,1323,163,1188]" captionTargetPageId="9" captionText="FIGURE 5. Female genitalia of Urostylis spp. A–E: female genital plates (with drawings); F–J: spermatheca; A, F: U. koreana sp. nov.; B, G: U. annulicornis; C, H: U. lateralis; D, I: U. striicornis; E, J: U. westwoodii. VII: seventh abdominal segment; gc8: gonocoxites 8; la8: laterotergites 8; la9: laterotergites 9." figureDoi="http://doi.org/10.5281/zenodo.1290383" httpUri="https://zenodo.org/record/1290383/files/figure.png" pageId="2" pageNumber="447">Figure 5G</figureCitation>
).
</paragraph>
<caption id="DF4F6660FFADFF83FF13C657F92FFA80" ID-DOI="http://doi.org/10.5281/zenodo.1290375" httpUri="https://zenodo.org/record/1290375/files/figure.png" pageId="3" pageNumber="448" startId="3.[151,250,1355,1377]" targetBox="[171,1408,185,1335]" targetPageId="3">
<paragraph id="8B8F36E8FFADFF83FF13C657F92FFA80" blockId="3.[151,1436,1355,1407]" pageId="3" pageNumber="448">
<emphasis id="B944EAFAFFADFF83FF13C657FA72FA9F" bold="true" box="[151,274,1355,1377]" pageId="3" pageNumber="448">FIGURE 1.</emphasis>
Dorsal habitus of 
<taxonomicName id="4C304D6BFFADFF83FE4FC657F947FA9E" authorityName="Westwood" authorityYear="1837" box="[459,551,1355,1377]" class="Insecta" family="Urostylididae" genus="Urostylis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="448" phylum="Arthropoda" rank="genus">
<emphasis id="B944EAFAFFADFF83FE4FC657F947FA9E" box="[459,551,1355,1377]" italics="true" pageId="3" pageNumber="448">Urostylis</emphasis>
</taxonomicName>
spp. A: 
<taxonomicName id="4C304D6BFFADFF83FDFEC650F98AFA9E" authority="Kim &amp; Roca-Cusachs &amp; Jung, 2018" authorityName="Kim &amp; Roca-Cusachs &amp; Jung" authorityYear="2018" box="[634,746,1355,1377]" class="Insecta" family="Urostylididae" genus="Urostylis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="448" phylum="Arthropoda" rank="species" species="koreana" status="sp. nov.">
<emphasis id="B944EAFAFFADFF83FDFEC650F98AFA9E" box="[634,746,1355,1377]" italics="true" pageId="3" pageNumber="448">U. koreana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2775781FFADFF83FD74C657F820FA9F" box="[752,832,1355,1376]" pageId="3" pageNumber="448" rank="species">
<emphasis id="B944EAFAFFADFF83FD74C657F820FA9F" bold="true" box="[752,832,1355,1376]" pageId="3" pageNumber="448">sp. nov.</emphasis>
</taxonomicNameLabel>
; B: 
<taxonomicName id="4C304D6BFFADFF83FCEFC650FF6AFA9E" authorityName="Scott" authorityYear="1874" box="[875,1034,1355,1377]" class="Insecta" family="Urostylididae" genus="Urostylis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="448" phylum="Arthropoda" rank="species" species="annulicornis">
<emphasis id="B944EAFAFFADFF83FCEFC650FF6AFA9E" box="[875,1034,1355,1377]" italics="true" pageId="3" pageNumber="448">U. annulicornis</emphasis>
</taxonomicName>
; C: 
<taxonomicName id="4C304D6BFFADFF83FBB0C650FFC6FA9E" authorityName="Walker" authorityYear="1867" box="[1076,1190,1355,1377]" class="Insecta" family="Urostylididae" genus="Urostylis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="448" phylum="Arthropoda" rank="species" species="lateralis">
<emphasis id="B944EAFAFFADFF83FBB0C650FFC6FA9E" box="[1076,1190,1355,1377]" italics="true" pageId="3" pageNumber="448">U. lateralis</emphasis>
</taxonomicName>
; D: 
<emphasis id="B944EAFAFFADFF83FB57C650FE02FA9E" box="[1235,1378,1356,1377]" italics="true" pageId="3" pageNumber="448">
<taxonomicName id="4C304D6BFFADFF83FB57C650FE3BFA9E" authorityName="Scott" authorityYear="1874" box="[1235,1371,1356,1377]" class="Insecta" family="Urostylididae" genus="Urostylis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="448" phylum="Arthropoda" rank="species" species="striicornis">U. striicornis</taxonomicName>
;
</emphasis>
E: 
<taxonomicName id="4C304D6BFFADFF83FA07C650FB83FA80" authorityName="Kerzhner" authorityYear="1966" class="Insecta" family="Urostylididae" genus="Urostylis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="448" phylum="Arthropoda" rank="species" species="trullata">
<emphasis id="B944EAFAFFADFF83FA07C650FB83FA80" italics="true" pageId="3" pageNumber="448">U. trullata</emphasis>
</taxonomicName>
; F: 
<taxonomicName id="4C304D6BFFADFF83FE88C676FAFDFA80" authorityName="Scott" authorityYear="1874" box="[268,413,1385,1407]" class="Insecta" family="Urostylididae" genus="Urostylis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="448" phylum="Arthropoda" rank="species" species="westwoodii">
<emphasis id="B944EAFAFFADFF83FE88C676FAFDFA80" box="[268,413,1385,1407]" italics="true" pageId="3" pageNumber="448">U. westwoodii</emphasis>
</taxonomicName>
. Scale bar: 1mm.
</paragraph>
</caption>
<paragraph id="8B8F36E8FFADFF83FF43C6B1FA27F8C7" blockId="3.[151,1437,1452,2028]" pageId="3" pageNumber="448">
MEASUREMENTS (IN MM): Male (n=4)/Female (n=2) Body length, head-apex of membrane: 12.30–13.26/ 13.28–13.34; head length: 1.05–1.19/1.09–1.21; head width, including compound eyes: 1.92–2.03/1.94–2.04; 1 
<taxonomicName id="4C304D6BFFADFF83FA16C6D3FB96F9ED" pageId="3" pageNumber="448" rank="subSpecies" subSpecies="antennal">st antennal</taxonomicName>
segment length: 3.45–3.54/2.75–2.79; 2nd antennal segment length: 4.16–4.32/3.27–3.70; 3rd antennal segment length: 1.65–1.79/1.47–1.54; 4th antennal segment length: 3.08–3.22/2.64–2.72; 5th antennal segment length: missing–1.93/2.18–2.38; total antennal length: 12.33 (1st to 4th)–14.80/12.39–13.07; mesal pronotal length: 2.30–2.44/2.15–2.30; basal pronotal maximal width (straight): 4.70–4.92/4.80–4.90; anterior scutelluar width: 2.82–2.97/3.14–3.20; mesal scutellar length: 3.47–3.59/3.60–4.14; costal margin length (straight): 7.92–8.21/7.70– 7.95; maximal width across hemelytron: 2.54–2.71/2.55–2.75; foreleg (femur: tibia: tarsus): 3.63–3.85:3.62– 3.81:1.57–1.63/3.86–3.90:3.31–3.32:1.70–1.90; midleg (femur: tibia: tarsus): 4.64–4.74:4.73–4.87:1.80–1.89/ 4.05–4.29:4.21–4.46:1.69–1.96; hindleg (femur: tibia: tarsus): 5.90–6.01:5.59–5.72:2.03–2.23/4.88–5.13:4.56– 4.99:1.89–2.07.
</paragraph>
</subSubSection>
<subSubSection id="C32A6563FFADFF83FF43C45FFFEDF813" pageId="3" pageNumber="448" type="materials_examined">
<paragraph id="8B8F36E8FFADFF83FF43C45FFFEDF813" blockId="3.[151,1437,1452,2028]" pageId="3" pageNumber="448">
<emphasis id="B944EAFAFFADFF83FF43C45FFA85F8A3" bold="true" box="[199,485,1859,1884]" pageId="3" pageNumber="448">MATERIALS EXAMINED:</emphasis>
[
<collectionCode id="ED21AE2DFFADFF83FE7CC45FF959F8A4" box="[504,569,1859,1883]" country="Korea" httpUri="http://grbio.org/cool/kyjm-bn0n" name="Chonbuk National University" pageId="3" pageNumber="448">CNU</collectionCode>
] 
<specimenCount id="9D36FD61FFADFF83FDCFC45FF9E5F8A4" box="[587,645,1859,1883]" pageId="3" pageNumber="448" type="male">2♂♂</specimenCount>
, Donam-ri, Geumnam-myeon, 
<collectingRegion id="49F4F80AFFADFF83FC77C45FFF24F8A4" box="[1011,1092,1859,1883]" country="South Korea" name="Sejong" pageId="3" pageNumber="448">Sejong</collectingRegion>
, 
<collectingCountry id="F3277678FFADFF83FBD0C458FFFBF8A4" box="[1108,1179,1860,1883]" name="South Korea" pageId="3" pageNumber="448">Korea</collectingCountry>
, 
<date id="FF8E1028FFADFF83FB2FC45FFE78F8A4" box="[1195,1304,1859,1883]" pageId="3" pageNumber="448" value="2015-06-03">3.vi.2015</date>
, H.U. Lee; [
<collectionCode id="ED21AE2DFFADFF83FF19C474FBBEF87F" box="[157,222,1896,1920]" country="Korea" httpUri="http://grbio.org/cool/kyjm-bn0n" name="Chonbuk National University" pageId="3" pageNumber="448">CNU</collectionCode>
] 
<specimenCount id="9D36FD61FFADFF83FF74C474FA74F87F" box="[240,276,1896,1920]" pageId="3" pageNumber="448" type="male">1♂</specimenCount>
, 
<collectingRegion id="49F4F80AFFADFF83FEA1C47BFAC0F880" box="[293,416,1895,1919]" country="South Korea" name="Chungnam" pageId="3" pageNumber="448">Chungnam</collectingRegion>
National Univ., Gung-dong, Yuseong-gu, 
<collectingRegion id="49F4F80AFFADFF83FC0BC47BF88FF880" box="[911,1007,1895,1919]" country="South Korea" name="Daejon" pageId="3" pageNumber="448">Daejeon</collectingRegion>
, 
<collectingCountry id="F3277678FFADFF83FC7AC474FF25F880" box="[1022,1093,1896,1919]" name="South Korea" pageId="3" pageNumber="448">Korea</collectingCountry>
, 
<date id="FF8E1028FFADFF83FBD1C47BFFA1F880" box="[1109,1217,1895,1919]" pageId="3" pageNumber="448" value="2015-06-01">1.vi.2015</date>
, Y.S. Kim; [
<collectionCode id="ED21AE2DFFADFF83FAD1C474FEF6F87F" box="[1365,1430,1896,1920]" country="Korea" httpUri="http://grbio.org/cool/kyjm-bn0n" name="Chonbuk National University" pageId="3" pageNumber="448">CNU</collectionCode>
] 
<specimenCount id="9D36FD61FFADFF83FF13C497FBB0F85C" box="[151,208,1931,1955]" pageId="3" pageNumber="448" type="female">2♀♀</specimenCount>
, 
<collectingRegion id="49F4F80AFFADFF83FF5AC497FA39F85C" box="[222,345,1931,1955]" country="South Korea" name="Chungnam" pageId="3" pageNumber="448">Chungnam</collectingRegion>
National Univ., Gung-dong, Yuseong-gu, 
<collectingRegion id="49F4F80AFFADFF83FCBFC497F8FBF85C" box="[827,923,1931,1955]" country="South Korea" name="Daejon" pageId="3" pageNumber="448">Daejeon</collectingRegion>
, 
<collectingCountry id="F3277678FFADFF83FC23C490F88EF85C" box="[935,1006,1932,1955]" name="South Korea" pageId="3" pageNumber="448">Korea</collectingCountry>
, 
<date id="FF8E1028FFADFF83FC7FC497FF0CF85C" box="[1019,1132,1931,1955]" pageId="3" pageNumber="448" value="2015-05-12">12.v.2015</date>
, S.A. Kwon; [
<collectionCode id="ED21AE2DFFADFF83FA8AC497FE2FF85C" box="[1294,1359,1931,1955]" country="Korea" httpUri="http://grbio.org/cool/kyjm-bn0n" name="Chonbuk National University" pageId="3" pageNumber="448">CNU</collectionCode>
] 
<specimenCount id="9D36FD61FFADFF83FAD9C497FEF7F85C" box="[1373,1431,1931,1955]" pageId="3" pageNumber="448" type="female">2♀♀</specimenCount>
, Jichon-ri, Yanggang-myeon, Yeongdong-gun, 
<collectingRegion id="49F4F80AFFADFF83FD34C4B3F8FEF838" box="[688,926,1967,1991]" country="South Korea" name="Chungbuk" pageId="3" pageNumber="448">Chungcheongbuk-do</collectingRegion>
, 
<collectingCountry id="F3277678FFADFF83FC34C4ACF897F838" box="[944,1015,1968,1991]" name="South Korea" pageId="3" pageNumber="448">Korea</collectingCountry>
, 
<date id="FF8E1028FFADFF83FB8CC4B3FFE2F838" box="[1032,1154,1967,1991]" pageId="3" pageNumber="448" value="2014-06-24">24.vi.2014</date>
, H.U. Lee; [
<collectionCode id="ED21AE2DFFADFF83FA9AC4ACFE3FF837" box="[1310,1375,1968,1992]" country="Korea" httpUri="http://grbio.org/cool/kyjm-bn0n" name="Chonbuk National University" pageId="3" pageNumber="448">CNU</collectionCode>
] 
<specimenCount id="9D36FD61FFADFF83FAF6C4ACFEF7F837" box="[1394,1431,1968,1992]" pageId="3" pageNumber="448" type="male">1♂</specimenCount>
, Wondang-ri, Jangnam-myeon, Yeoncheon-gun, 
<collectingRegion id="49F4F80AFFADFF83FD2BC4CFF825F813" box="[687,837,2003,2028]" country="South Korea" name="Gyeonggi" pageId="3" pageNumber="448">Gyeonggi-do</collectingRegion>
, 
<collectingCountry id="F3277678FFADFF83FCD5C4C8F8F8F814" box="[849,920,2004,2027]" name="South Korea" pageId="3" pageNumber="448">Korea</collectingCountry>
, 
<date id="FF8E1028FFADFF83FC27C4CFFF78F813" box="[931,1048,2003,2028]" pageId="3" pageNumber="448" value="2014-07-03">3.vii.2014</date>
, J.G. Kim.
</paragraph>
</subSubSection>
<subSubSection id="C32A6563FFAAFF84FF43C38BFF2CFF2B" pageId="4" pageNumber="449" type="biology_ecology">
<paragraph id="8B8F36E8FFAAFF84FF43C38BF8BDFF50" blockId="4.[199,1100,151,213]" box="[199,989,151,176]" pageId="4" pageNumber="449">
<emphasis id="B944EAFAFFAAFF84FF43C38BFA7FFF4F" bold="true" box="[199,287,151,176]" pageId="4" pageNumber="449">HOSTS:</emphasis>
<taxonomicName id="4C304D6BFFAAFF84FEA2C384F9E6FF4F" box="[294,646,152,176]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="449" phylum="Tracheophyta" rank="species" species="mongolica">
<emphasis id="B944EAFAFFAAFF84FEA2C384F961FF4F" box="[294,513,152,176]" italics="true" pageId="4" pageNumber="449">Quercus mongolica</emphasis>
(Fagaceae)
</taxonomicName>
, 
<taxonomicName id="4C304D6BFFAAFF84FD15C384F82AFF4F" box="[657,842,152,176]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="449" phylum="Tracheophyta" rank="species" species="dentata">
<emphasis id="B944EAFAFFAAFF84FD15C384F82AFF4F" box="[657,842,152,176]" italics="true" pageId="4" pageNumber="449">Quercus dentata</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C304D6BFFAAFF84FCDCC384F8D6FF50" box="[856,950,152,175]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Fagales" pageId="4" pageNumber="449" phylum="Tracheophyta" rank="genus">
<emphasis id="B944EAFAFFAAFF84FCDCC384F8D6FF50" box="[856,950,152,175]" italics="true" pageId="4" pageNumber="449">Quercus</emphasis>
</taxonomicName>
sp.
</paragraph>
<paragraph id="8B8F36E8FFAAFF84FF43C3A0FF2CFF2B" blockId="4.[199,1100,151,213]" box="[199,1100,188,213]" pageId="4" pageNumber="449">
<emphasis id="B944EAFAFFAAFF84FF43C3A0FAE6FF2A" bold="true" box="[199,390,188,213]" pageId="4" pageNumber="449">DISTRIBUTIONS:</emphasis>
<collectingCountry id="F3277678FFAAFF84FE09C3A0FAB0FF2A" box="[397,464,188,213]" name="China" pageId="4" pageNumber="449">China</collectingCountry>
(Northern East, North), 
<collectingCountry id="F3277678FFAAFF84FD67C3A1F846FF2B" box="[739,806,189,212]" name="Japan" pageId="4" pageNumber="449">Japan</collectingCountry>
, 
<collectingCountry id="F3277678FFAAFF84FCB5C3A1F818FF2B" box="[817,888,189,212]" name="South Korea" pageId="4" pageNumber="449">Korea</collectingCountry>
, 
<collectingCountry id="F3277678FFAAFF84FC00C3A0F8AFFF2A" box="[900,975,188,213]" name="Russia" pageId="4" pageNumber="449">Russia</collectingCountry>
(Far Eest).
</paragraph>
</subSubSection>
</treatment>
</document>