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351 lines
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<mods:title id="028C222758B5CD3B6AFAB9FE3A1E97F4">Endogean and Cavernicolous Coleoptera of the Balkans. XX. Notes on the genus Protamaurops J. Müller, 1944 (Coleoptera: Staphylinidae: Pselaphinae) with the description of a new species from Bulgaria</mods:title>
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<mods:namePart id="8EE50BDF3DEC0DDECB5BC7C3D1D3F2FA">Bekchiev, Rostislav</mods:namePart>
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<mods:affiliation id="EC949A5A962EB375EFC75558937591F1">National Museum of Natural History, Bulgarian Academy of Sciences, 1 Tsar Osvoboditel Blvd, 1000 Sofia, Bulgaria bekchiev @ nmnhs. com; https: // orcid. org / 0000 - 0001 - 6143 - 0184</mods:affiliation>
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<mods:namePart id="4E9D90F3CB114E06F51A0437C73C25C0">Hlaváč, Peter</mods:namePart>
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<mods:affiliation id="F4906659A86216DA23B48BC30077C3A7">Department of Entomology, National Museum, Natural History Museum, Cirkusová 1740, CZ- 193 00 Praha 9 - Horní Počernice, Czech Republic. peterclaviger @ gmail. com; https: // orcid. org / 0000 - 0001 - 5060 - 0811</mods:affiliation>
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<heading id="D08581AEA959DF6B2DCAFABDFD9E0A5C" bold="true" box="[151,605,1375,1402]" fontSize="11" level="1" pageId="1" pageNumber="368" reason="1">
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<emphasis id="B906EAD0A959DF6B2DCAFABDFD9E0A5C" bold="true" box="[151,605,1375,1402]" pageId="1" pageNumber="368">
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Genus
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<taxonomicName id="4C724D41A959DF6B2DB3FA82FD9E0A5C" ID-CoL="8XP63" authority="J. Muller, 1944" authorityName="J. Muller" authorityYear="1944" box="[238,605,1376,1402]" class="Insecta" family="Staphylinidae" genus="Protamaurops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="368" phylum="Arthropoda" rank="genus">
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<emphasis id="B906EAD0A959DF6B2DB3FA82FE580A5C" bold="true" box="[238,411,1376,1402]" italics="true" pageId="1" pageNumber="368">Protamaurops</emphasis>
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J.
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<bibRefCitation id="EFE34B33A959DF6B2C9DFA82FD9E0A5C" author="Muller, G." box="[448,605,1376,1402]" pageId="1" pageNumber="368" pagination="82 - 103" refId="ref6585" refString="Muller, G. (1944) Die Amaurops-Arten der Balkanhalbinsel. Mitteilungen der Munchner Entomologischen Gesellschaft, 34, 82 - 103." type="journal article" year="1944">Müller, 1944</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C3686549A959DF6B2DCAFA47FADB093E" pageId="1" pageNumber="368" type="reference_group">
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<treatmentCitation id="0AD310D3A959DF6B2DCAFA47FE320A9A" author="Muller, G." box="[151,497,1443,1468]" page="84" pageId="1" pageNumber="368" year="1944">
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<taxonomicName id="4C724D41A959DF6B2DCAFA47FE320A9A" ID-CoL="8XP63" authority="J. Muller, 1944: 84" authorityName="J. Muller" authorityPageNumber="84" authorityYear="1944" box="[151,497,1443,1468]" class="Insecta" family="Staphylinidae" genus="Protamaurops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="368" phylum="Arthropoda" rank="genus">
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<emphasis id="B906EAD0A959DF6B2DCAFA47FEEB0A9A" box="[151,296,1445,1468]" italics="true" pageId="1" pageNumber="368">Protamaurops</emphasis>
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J.
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<bibRefCitation id="EFE34B33A959DF6B2C18FA41FE320A9A" author="Muller, G." box="[325,497,1443,1468]" pageId="1" pageNumber="368" pagination="82 - 103" refId="ref6585" refString="Muller, G. (1944) Die Amaurops-Arten der Balkanhalbinsel. Mitteilungen der Munchner Entomologischen Gesellschaft, 34, 82 - 103." type="journal article" year="1944">Müller, 1944: 84</bibRefCitation>
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</taxonomicName>
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.
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<typeStatus id="54C98860A959DF6B2CA6FA46FDEE0A9A" box="[507,557,1444,1468]" pageId="1" pageNumber="368">Type</typeStatus>
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species:
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<taxonomicName id="4C724D41A959DF6B2FD7FA46FBF40A9A" ID-CoL="CK6N" authority="J. Muller, 1944" authorityName="J. Muller" authorityYear="1944" box="[650,1079,1443,1468]" class="Insecta" family="Staphylinidae" genus="Amaurops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="368" phylum="Arthropoda" rank="species" species="macrophthalma">
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<emphasis id="B906EAD0A959DF6B2FD7FA46FC550A9A" box="[650,918,1444,1468]" italics="true" pageId="1" pageNumber="368">Amaurops macrophthalma</emphasis>
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J.
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<bibRefCitation id="EFE34B33A959DF6B2EE9FA41FBF40A9A" author="Muller, G." box="[948,1079,1443,1468]" pageId="1" pageNumber="368" pagination="82 - 103" refId="ref6585" refString="Muller, G. (1944) Die Amaurops-Arten der Balkanhalbinsel. Mitteilungen der Munchner Entomologischen Gesellschaft, 34, 82 - 103." type="journal article" year="1944">Müller, 1944</bibRefCitation>
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<paragraph id="8BCD36C2A959DF6B2DCAFA21FBE10ADC" blockId="1.[151,1436,1443,1560]" pageId="1" pageNumber="368">
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<treatmentCitationGroup id="AB6211ECA959DF6B2DCAFA21FBE10ADC" pageId="1" pageNumber="368">
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<taxonomicName id="4C724D41A959DF6B2DCAFA21FE4E0AFD" ID-CoL="8XP63" authority="J. Muller" authorityName="J. Muller" authorityYear="1944" box="[151,397,1474,1499]" class="Insecta" family="Staphylinidae" genus="Protamaurops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="368" phylum="Arthropoda" rank="genus">
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<emphasis id="B906EAD0A959DF6B2DCAFA21FEEB0AFC" box="[151,296,1475,1498]" italics="true" pageId="1" pageNumber="368">Protamaurops</emphasis>
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J. Müller
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</taxonomicName>
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: J.
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<treatmentCitation id="0AD310D3A959DF6B2CF1FA20FD950AFD" author="Muller, G." box="[428,598,1474,1499]" page="89" pageId="1" pageNumber="368" year="1944">
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<bibRefCitation id="EFE34B33A959DF6B2CF1FA20FD950AFD" author="Muller, G." box="[428,598,1474,1499]" pageId="1" pageNumber="368" pagination="82 - 103" refId="ref6585" refString="Muller, G. (1944) Die Amaurops-Arten der Balkanhalbinsel. Mitteilungen der Munchner Entomologischen Gesellschaft, 34, 82 - 103." type="journal article" year="1944">Müller, 1944: 89</bibRefCitation>
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</treatmentCitation>
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(key), 92 (key to species);
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<treatmentCitation id="0AD310D3A959DF6B2E34FA21FBF90AFD" author="Karaman, Z." box="[873,1082,1474,1499]" page="153" pageId="1" pageNumber="368" year="1961">
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<bibRefCitation id="EFE34B33A959DF6B2E34FA21FBF90AFD" author="Karaman, Z." box="[873,1082,1474,1499]" pageId="1" pageNumber="368" pagination="149 - 169" refId="ref6455" refString="Karaman, Z. (1961) Prilog Poznavanju Amauropsina Balkana. Beitrag zur Kenntnis der Amauropsinen des Balkans. Godisen Zbornik na Zemjodelsko-Sumarski Fakultet na Univerzitet, Skopje, 14, 149 - 169." type="journal article" year="1961">
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<collectingRegion id="49B6F820A959DF6B2E34FA21FC0B0AFC" box="[873,968,1475,1498]" country="Turkey" name="Karaman" pageId="1" pageNumber="368">Karaman</collectingRegion>
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, 1961: 153
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</bibRefCitation>
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</treatmentCitation>
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, 164 (diagnosis); Newton & Chan- dler, 1989: 32 (catalogue);
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<treatmentCitation id="0AD310D3A959DF6B2C84FA03FCF80ADC" author="Nonveiller, G. & Pavicevic, D." box="[473,827,1505,1530]" page="263" pageId="1" pageNumber="368" year="2008">
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<bibRefCitation id="EFE34B33A959DF6B2C84FA03FCF80ADC" author="Nonveiller, G. & Pavicevic, D." box="[473,827,1505,1530]" pageId="1" pageNumber="368" refId="ref6663" refString="Nonveiller, G. & Pavicevic, D. (2008) Une nouvelle espece et deux nouveaux sous-genres de Seracamaurops, avec un catalogue des Amauropini mediterraneens (Coleoptera, Staphylinidae, Pselaphinae) (Recherches sur les Coleopteres endoges et troglobiontes de Serbie et des regions Institute for Nature Conservation of Serbia, Belgrade, 564 pp." type="book" year="2008">Nonveiller & Pavićević, 2008: 263</bibRefCitation>
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</treatmentCitation>
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(key), 265 (catalogue)
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BCD36C2A959DF6B2DCAF9E2FADB093E" blockId="1.[151,1436,1443,1560]" box="[151,1304,1535,1560]" pageId="1" pageNumber="368">
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<treatmentCitationGroup id="AB6211ECA959DF6B2DCAF9E2FADB093E" box="[151,1304,1535,1560]" pageId="1" pageNumber="368">
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<treatmentCitation id="0AD310D3A959DF6B2DCAF9E2FE66093E" author="Lobl, I." box="[151,421,1535,1560]" page="1" pageId="1" pageNumber="368" year="1967">
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<taxonomicName id="4C724D41A959DF6B2DCAF9E2FE66093E" ID-CoL="8XPS3" authority="Lobl, 1967: 1" authorityName="Lobl" authorityPageNumber="1" authorityYear="1967" box="[151,421,1535,1560]" class="Insecta" family="Staphylinidae" genus="Schweigeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="368" phylum="Arthropoda" rank="genus">
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<emphasis id="B906EAD0A959DF6B2DCAF9E2FED7093E" box="[151,276,1536,1560]" italics="true" pageId="1" pageNumber="368">Schweigeria</emphasis>
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<bibRefCitation id="EFE34B33A959DF6B2C46FA1DFE66093E" author="Lobl, I." box="[283,421,1535,1560]" pageId="1" pageNumber="368" pagination="1 - 4" refId="ref6526" refString="Lobl, I. (1967) Ein neuer Pselaphide (Coleoptera) aus der Turkei. Annotationes Zoologicae et Botanicae, 40 1 - 4." type="journal article" year="1967">Löbl, 1967: 1</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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<typeStatus id="54C98860A959DF6B2CEDF9E2FE21093E" box="[432,482,1536,1560]" pageId="1" pageNumber="368">Type</typeStatus>
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species:
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<taxonomicName id="4C724D41A959DF6B2F62F9E2FC77093E" ID-CoL="4VGGJ" authority="Lobl, 1967" authorityName="Lobl" authorityYear="1967" box="[575,948,1535,1560]" class="Insecta" family="Staphylinidae" genus="Schweigeria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="368" phylum="Arthropoda" rank="species" species="uludagensis">
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<emphasis id="B906EAD0A959DF6B2F62F9E2FCFF093E" box="[575,828,1536,1560]" italics="true" pageId="1" pageNumber="368">Schweigeria uludagensis</emphasis>
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<bibRefCitation id="EFE34B33A959DF6B2E1FFA1DFC77093E" author="Lobl, I." box="[834,948,1535,1560]" pageId="1" pageNumber="368" pagination="1 - 4" refId="ref6526" refString="Lobl, I. (1967) Ein neuer Pselaphide (Coleoptera) aus der Turkei. Annotationes Zoologicae et Botanicae, 40 1 - 4." type="journal article" year="1967">Löbl, 1967</bibRefCitation>
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</taxonomicName>
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, synonymy in
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<treatmentCitation id="0AD310D3A959DF6B2915FA1DFADB093E" author="Besuchet, C." box="[1096,1304,1535,1560]" page="258" pageId="1" pageNumber="368" year="1986">
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<bibRefCitation id="EFE34B33A959DF6B2915FA1DFADB093E" author="Besuchet, C." box="[1096,1304,1535,1560]" pageId="1" pageNumber="368" pagination="257 - 264" refId="ref6155" refString="Besuchet, C. (1986) Synonymes et homonyme nouveaux de quelques genres de Pselaphides (Coleoptera). Revue Suisse de Zoologie, 93, 257 - 264. https: // doi. org / 10.5962 / bhl. part. 82019" type="journal article" year="1986">Besuchet, 1986: 258</bibRefCitation>
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</treatmentCitation>
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</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3686549A959DF682DCAF9A7FB2E088D" lastPageId="2" lastPageNumber="369" pageId="1" pageNumber="368" type="description">
|
|||
|
|
<paragraph id="8BCD36C2A959DF6B2DCAF9A7FC4F09A3" blockId="1.[151,1436,1605,2029]" pageId="1" pageNumber="368">
|
|||
|
|
<emphasis id="B906EAD0A959DF6B2DCAF9A7FEFE0946" bold="true" box="[151,317,1605,1632]" pageId="1" pageNumber="368">Redescription</emphasis>
|
|||
|
|
. Body (
|
|||
|
|
<figureCitation id="13492A47A959DF6B2CC3F9A4FE2C0946" box="[414,495,1606,1632]" captionStart="FIGURE 1" captionStartId="2.[151,250,1254,1279]" captionTargetBox="[161,1426,184,1215]" captionTargetId="figure@2.[151,1436,181,1229]" captionTargetPageId="2" captionText="FIGURE 1. Habitus and head of Protamaurops—a) P. macrophthalmus (male), scale—1 mm; b) head with the carina, scale— 0.5 mm; c) P. bulgaricus sp. n., head with the carinae (male, holotype), scale—0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3835419" httpUri="https://zenodo.org/record/3835419/files/figure.png" pageId="1" pageNumber="368">Fig. 1a</figureCitation>
|
|||
|
|
) reddish-brown, covered with long, suberect golden setae; length
|
|||
|
|
<quantity id="4C8A9B27A959DF6B29B2F9A7FA5B0947" box="[1263,1432,1605,1633]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.65" metricValueMin="2.15" pageId="1" pageNumber="368" unit="mm" value="2.4" valueMax="2.65" valueMin="2.15">2.15–2.65 mm</quantity>
|
|||
|
|
; lateral side of head under ocular spines with long and erected setae.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BCD36C2A959DF6B2D9AF96CFA5B087A" blockId="1.[151,1436,1605,2029]" pageId="1" pageNumber="368">
|
|||
|
|
Head (
|
|||
|
|
<figureCitation id="13492A47A959DF6B2C4FF96CFEBC098F" box="[274,383,1678,1705]" captionStart="FIGURE 1" captionStartId="2.[151,250,1254,1279]" captionTargetBox="[161,1426,184,1215]" captionTargetId="figure@2.[151,1436,181,1229]" captionTargetPageId="2" captionText="FIGURE 1. Habitus and head of Protamaurops—a) P. macrophthalmus (male), scale—1 mm; b) head with the carina, scale— 0.5 mm; c) P. bulgaricus sp. n., head with the carinae (male, holotype), scale—0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3835419" httpUri="https://zenodo.org/record/3835419/files/figure.png" pageId="1" pageNumber="368">Fig. 1b, c</figureCitation>
|
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|
|
,
|
|||
|
|
<figureCitation id="13492A47A959DF6B2CD6F96CFE64098E" box="[395,423,1678,1704]" captionStart="FIGURE 2" captionStartId="3.[151,250,1767,1792]" captionTargetBox="[154,1420,194,1741]" captionTargetId="figure@3.[151,1436,181,1742]" captionTargetPageId="3" captionText="FIGURE 2. Morphology of Protamaurops (P. macedonicus)—a) Head (dorsal view); b) Pronotum (dorsal and ventral view); c) Meso-and metaventrite; d) Elytra; e) Abdomen.Abbreviations: vf—vertexal fovea; laf—lateral antebasal foveae; maf—me- dian antebasal fovea; blf—basolateral foveae; lpcf—lateral procoxal foveae; mmvf—median mesoventral foveae; lmsf—lateral mesoventral foveae; lmcf—lateral mesocoxal foveae; lmvf—lateral metaventral foveae; bef—basal foveae; ec—external ca- rina; mc—median carinae. Scale: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3835421" httpUri="https://zenodo.org/record/3835421/files/figure.png" pageId="1" pageNumber="368">2a</figureCitation>
|
|||
|
|
) wider than long, antennal tubercles prominent; frontal sulcus large, triangular, opened on frontal rostrum, which is triangular and extended anteriorly; with well-defined lateral carinae orginated from anten- nal tubercle and reaching occipital region; with more or less prominent hump, bearing one long (
|
|||
|
|
<figureCitation id="13492A47A959DF6B29B3F934FAFC09D7" box="[1262,1343,1750,1777]" captionStart="FIGURE 1" captionStartId="2.[151,250,1254,1279]" captionTargetBox="[161,1426,184,1215]" captionTargetId="figure@2.[151,1436,181,1229]" captionTargetPageId="2" captionText="FIGURE 1. Habitus and head of Protamaurops—a) P. macrophthalmus (male), scale—1 mm; b) head with the carina, scale— 0.5 mm; c) P. bulgaricus sp. n., head with the carinae (male, holotype), scale—0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3835419" httpUri="https://zenodo.org/record/3835419/files/figure.png" pageId="1" pageNumber="368">Fig. 1b</figureCitation>
|
|||
|
|
) or tree vertexal carinae of different length, one long, and two short (
|
|||
|
|
<figureCitation id="13492A47A959DF6B2E17F918FC580832" box="[842,923,1786,1812]" captionStart="FIGURE 1" captionStartId="2.[151,250,1254,1279]" captionTargetBox="[161,1426,184,1215]" captionTargetId="figure@2.[151,1436,181,1229]" captionTargetPageId="2" captionText="FIGURE 1. Habitus and head of Protamaurops—a) P. macrophthalmus (male), scale—1 mm; b) head with the carina, scale— 0.5 mm; c) P. bulgaricus sp. n., head with the carinae (male, holotype), scale—0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3835419" httpUri="https://zenodo.org/record/3835419/files/figure.png" pageId="1" pageNumber="368">Fig. 1c</figureCitation>
|
|||
|
|
), punctuation from sparse to dense; with two deep vertexal foveae. Eyes from well-developed to more or less reduced, always with distinct sharp lateral spine (
|
|||
|
|
<figureCitation id="13492A47A959DF6B2DC2F8A0FEE8087B" box="[159,299,1858,1885]" captionStart="FIGURE 1" captionStartId="2.[151,250,1254,1279]" captionTargetBox="[161,1426,184,1215]" captionTargetId="figure@2.[151,1436,181,1229]" captionTargetPageId="2" captionText="FIGURE 1. Habitus and head of Protamaurops—a) P. macrophthalmus (male), scale—1 mm; b) head with the carina, scale— 0.5 mm; c) P. bulgaricus sp. n., head with the carinae (male, holotype), scale—0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3835419" httpUri="https://zenodo.org/record/3835419/files/figure.png" pageId="1" pageNumber="368">Figs 1a, b, c</figureCitation>
|
|||
|
|
). Maxillary palpi with palpomeres I short, II lengthily pedunculate, III small, IV large and triangular.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BCD36C2A959DF6B2D9AF884FE7C08CB" blockId="1.[151,1436,1605,2029]" pageId="1" pageNumber="368">
|
|||
|
|
Antennae with eleven antennomeres; scape slightly longer than wide or as long as wide; pedicel longer than wide; antennomeres III and IV longer than wide or as long as wide; antennomere V–VII longer than wide, VIII and IX as long as wide, X wider than long and XI longer than wide, antennal club 3-segmented, lacking any modifications, long
|
|||
|
|
<quantity id="4C8A9B27A959DF6B2C49F830FE7908CB" box="[276,442,2002,2029]" metricMagnitude="-4" metricUnit="m" metricValue="3.4499999999999997" metricValueMax="3.7" metricValueMin="3.2" pageId="1" pageNumber="368" unit="mm" value="0.345" valueMax="0.37" valueMin="0.32">0.32–0.37 mm</quantity>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF0D664AA95ADF682DCAFB04FBD30A04" ID-DOI="http://doi.org/10.5281/zenodo.3835419" ID-Zenodo-Dep="3835419" httpUri="https://zenodo.org/record/3835419/files/figure.png" pageId="2" pageNumber="369" startId="2.[151,250,1254,1279]" targetBox="[161,1426,184,1215]" targetPageId="2">
|
|||
|
|
<paragraph id="8BCD36C2A95ADF682DCAFB04FBD30A04" blockId="2.[151,1436,1254,1315]" pageId="2" pageNumber="369">
|
|||
|
|
<emphasis id="B906EAD0A95ADF682DCAFB04FED00BD9" bold="true" box="[151,275,1254,1279]" pageId="2" pageNumber="369">FIGURE 1.</emphasis>
|
|||
|
|
Habitus and head of
|
|||
|
|
<taxonomicName id="4C724D41A95ADF682CB4FB0AFDB90BD9" authorityName="J. Muller" authorityYear="1944" box="[489,634,1256,1279]" class="Insecta" family="Staphylinidae" genus="Protamaurops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="369" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B906EAD0A95ADF682CB4FB0AFDB90BD9" box="[489,634,1256,1279]" italics="true" pageId="2" pageNumber="369">Protamaurops</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
—
|
|||
|
|
<emphasis id="B906EAD0A95ADF682FCEFB04FD6A0BD9" bold="true" box="[659,681,1254,1279]" pageId="2" pageNumber="369">a)</emphasis>
|
|||
|
|
<taxonomicName id="4C724D41A95ADF682FF3FB0AFCB00BD9" authorityName="J. Muller" authorityYear="1944" box="[686,883,1255,1279]" class="Insecta" family="Staphylinidae" genus="Protamaurops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="369" phylum="Arthropoda" rank="species" species="macrophthalmus">
|
|||
|
|
<emphasis id="B906EAD0A95ADF682FF3FB0AFD7E0BD9" box="[686,701,1256,1279]" italics="true" pageId="2" pageNumber="369">P</emphasis>
|
|||
|
|
.
|
|||
|
|
<emphasis id="B906EAD0A95ADF682F97FB05FCB00BD9" box="[714,883,1255,1279]" italics="true" pageId="2" pageNumber="369">macrophthalmus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(male), scale—1 mm;
|
|||
|
|
<emphasis id="B906EAD0A95ADF682906FB04FBB10BD9" bold="true" box="[1115,1138,1254,1279]" pageId="2" pageNumber="369">b)</emphasis>
|
|||
|
|
head with the carina, scale— 0.5 mm;
|
|||
|
|
<emphasis id="B906EAD0A95ADF682DAFFAE8FEC50A05" bold="true" box="[242,262,1290,1315]" pageId="2" pageNumber="369">c)</emphasis>
|
|||
|
|
<taxonomicName id="4C724D41A95ADF682C51FAEEFE520A05" authority="Bekchiev & Hlaváč, 2020" authorityName="Bekchiev & Hlaváč" authorityYear="2020" box="[268,401,1291,1315]" class="Insecta" family="Staphylinidae" genus="Protamaurops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="369" phylum="Arthropoda" rank="species" species="bulgaricus" status="sp. nov.">
|
|||
|
|
<emphasis id="B906EAD0A95ADF682C51FAEEFE520A05" box="[268,401,1291,1315]" italics="true" pageId="2" pageNumber="369">P. bulgaricus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B906EAD0A95ADF682CC5FAE9FE120A05" bold="true" box="[408,465,1291,1315]" pageId="2" pageNumber="369">
|
|||
|
|
<taxonomicNameLabel id="A23557ABA95ADF682CC5FAE9FE120A05" box="[408,465,1291,1315]" pageId="2" pageNumber="369" rank="species">sp. n.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
, head with the carinae (male, holotype), scale—0.5 mm.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BCD36C2A95ADF682D9AFAB0FDF30AF1" blockId="2.[151,1437,1361,1964]" pageId="2" pageNumber="369">
|
|||
|
|
Pronotum (
|
|||
|
|
<figureCitation id="13492A47A95ADF682C19FAB3FE570A4A" box="[324,404,1361,1388]" captionStart="FIGURE 2" captionStartId="3.[151,250,1767,1792]" captionTargetBox="[154,1420,194,1741]" captionTargetId="figure@3.[151,1436,181,1742]" captionTargetPageId="3" captionText="FIGURE 2. Morphology of Protamaurops (P. macedonicus)—a) Head (dorsal view); b) Pronotum (dorsal and ventral view); c) Meso-and metaventrite; d) Elytra; e) Abdomen.Abbreviations: vf—vertexal fovea; laf—lateral antebasal foveae; maf—me- dian antebasal fovea; blf—basolateral foveae; lpcf—lateral procoxal foveae; mmvf—median mesoventral foveae; lmsf—lateral mesoventral foveae; lmcf—lateral mesocoxal foveae; lmvf—lateral metaventral foveae; bef—basal foveae; ec—external ca- rina; mc—median carinae. Scale: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3835421" httpUri="https://zenodo.org/record/3835421/files/figure.png" pageId="2" pageNumber="369">Fig. 2b</figureCitation>
|
|||
|
|
) longer than wide, widest in the middle; dorsal keels well-definied, forming acute tooth close to base; area between keels strongly impressed; lateral antebasal foveae deep, connected by weak antebasal sulcus; median antebasal fovea large, connected with thin vertical carina to posterior margin of pronotum; basolateral fo- veae small, not connected by sulcus.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BCD36C2A95ADF682D9AFA00FBF20962" blockId="2.[151,1437,1361,1964]" pageId="2" pageNumber="369">
|
|||
|
|
Prosternum with two lateral procoxal foveae (
|
|||
|
|
<figureCitation id="13492A47A95ADF682F90FA03FCE20ADD" box="[717,801,1505,1531]" captionStart="FIGURE 2" captionStartId="3.[151,250,1767,1792]" captionTargetBox="[154,1420,194,1741]" captionTargetId="figure@3.[151,1436,181,1742]" captionTargetPageId="3" captionText="FIGURE 2. Morphology of Protamaurops (P. macedonicus)—a) Head (dorsal view); b) Pronotum (dorsal and ventral view); c) Meso-and metaventrite; d) Elytra; e) Abdomen.Abbreviations: vf—vertexal fovea; laf—lateral antebasal foveae; maf—me- dian antebasal fovea; blf—basolateral foveae; lpcf—lateral procoxal foveae; mmvf—median mesoventral foveae; lmsf—lateral mesoventral foveae; lmcf—lateral mesocoxal foveae; lmvf—lateral metaventral foveae; bef—basal foveae; ec—external ca- rina; mc—median carinae. Scale: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3835421" httpUri="https://zenodo.org/record/3835421/files/figure.png" pageId="2" pageNumber="369">Fig. 2b</figureCitation>
|
|||
|
|
,
|
|||
|
|
<emphasis id="B906EAD0A95ADF682E71FA03FC950ADD" box="[812,854,1505,1531]" italics="true" pageId="2" pageNumber="369">lpcf</emphasis>
|
|||
|
|
). Mesoventrite with one, large median mesoventral foveae, with deep, bifurcated, lateral mesoventral foveae and two deep lateral mesocoxal foveae (
|
|||
|
|
<figureCitation id="13492A47A95ADF6829AAF9E7FA890939" box="[1271,1354,1541,1567]" captionStart="FIGURE 2" captionStartId="3.[151,250,1767,1792]" captionTargetBox="[154,1420,194,1741]" captionTargetId="figure@3.[151,1436,181,1742]" captionTargetPageId="3" captionText="FIGURE 2. Morphology of Protamaurops (P. macedonicus)—a) Head (dorsal view); b) Pronotum (dorsal and ventral view); c) Meso-and metaventrite; d) Elytra; e) Abdomen.Abbreviations: vf—vertexal fovea; laf—lateral antebasal foveae; maf—me- dian antebasal fovea; blf—basolateral foveae; lpcf—lateral procoxal foveae; mmvf—median mesoventral foveae; lmsf—lateral mesoventral foveae; lmcf—lateral mesocoxal foveae; lmvf—lateral metaventral foveae; bef—basal foveae; ec—external ca- rina; mc—median carinae. Scale: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3835421" httpUri="https://zenodo.org/record/3835421/files/figure.png" pageId="2" pageNumber="369">Fig. 2c</figureCitation>
|
|||
|
|
,
|
|||
|
|
<emphasis id="B906EAD0A95ADF682804F9E7FEC60965" italics="true" pageId="2" pageNumber="369">mmvf, lmvf, lmcf</emphasis>
|
|||
|
|
). Metaventrite with two small lateral metaventral foveae (
|
|||
|
|
<figureCitation id="13492A47A95ADF682ECEF9CBFC250965" box="[915,998,1577,1603]" captionStart="FIGURE 2" captionStartId="3.[151,250,1767,1792]" captionTargetBox="[154,1420,194,1741]" captionTargetId="figure@3.[151,1436,181,1742]" captionTargetPageId="3" captionText="FIGURE 2. Morphology of Protamaurops (P. macedonicus)—a) Head (dorsal view); b) Pronotum (dorsal and ventral view); c) Meso-and metaventrite; d) Elytra; e) Abdomen.Abbreviations: vf—vertexal fovea; laf—lateral antebasal foveae; maf—me- dian antebasal fovea; blf—basolateral foveae; lpcf—lateral procoxal foveae; mmvf—median mesoventral foveae; lmsf—lateral mesoventral foveae; lmcf—lateral mesocoxal foveae; lmvf—lateral metaventral foveae; bef—basal foveae; ec—external ca- rina; mc—median carinae. Scale: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3835421" httpUri="https://zenodo.org/record/3835421/files/figure.png" pageId="2" pageNumber="369">Fig. 2b</figureCitation>
|
|||
|
|
,
|
|||
|
|
<emphasis id="B906EAD0A95ADF682EACF9CBFBE20965" box="[1009,1057,1577,1603]" italics="true" pageId="2" pageNumber="369">lmvf</emphasis>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BCD36C2A95ADF682D9AF9AFFD6009AD" blockId="2.[151,1437,1361,1964]" pageId="2" pageNumber="369">
|
|||
|
|
Elytra (
|
|||
|
|
<figureCitation id="13492A47A95ADF682C41F9AFFEAD094E" box="[284,366,1613,1640]" captionStart="FIGURE 2" captionStartId="3.[151,250,1767,1792]" captionTargetBox="[154,1420,194,1741]" captionTargetId="figure@3.[151,1436,181,1742]" captionTargetPageId="3" captionText="FIGURE 2. Morphology of Protamaurops (P. macedonicus)—a) Head (dorsal view); b) Pronotum (dorsal and ventral view); c) Meso-and metaventrite; d) Elytra; e) Abdomen.Abbreviations: vf—vertexal fovea; laf—lateral antebasal foveae; maf—me- dian antebasal fovea; blf—basolateral foveae; lpcf—lateral procoxal foveae; mmvf—median mesoventral foveae; lmsf—lateral mesoventral foveae; lmcf—lateral mesocoxal foveae; lmvf—lateral metaventral foveae; bef—basal foveae; ec—external ca- rina; mc—median carinae. Scale: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3835421" httpUri="https://zenodo.org/record/3835421/files/figure.png" pageId="2" pageNumber="369">Fig. 2d</figureCitation>
|
|||
|
|
) wider than long; each elytron with two large basal foveae, inner one bifurcate at base; sutural and discal striae short, humeri well-developed.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BCD36C2A95ADF682D9AF977FDA7083D" blockId="2.[151,1437,1361,1964]" pageId="2" pageNumber="369">
|
|||
|
|
Abdomen (
|
|||
|
|
<figureCitation id="13492A47A95ADF682C18F977FE560989" box="[325,405,1685,1711]" captionStart="FIGURE 2" captionStartId="3.[151,250,1767,1792]" captionTargetBox="[154,1420,194,1741]" captionTargetId="figure@3.[151,1436,181,1742]" captionTargetPageId="3" captionText="FIGURE 2. Morphology of Protamaurops (P. macedonicus)—a) Head (dorsal view); b) Pronotum (dorsal and ventral view); c) Meso-and metaventrite; d) Elytra; e) Abdomen.Abbreviations: vf—vertexal fovea; laf—lateral antebasal foveae; maf—me- dian antebasal fovea; blf—basolateral foveae; lpcf—lateral procoxal foveae; mmvf—median mesoventral foveae; lmsf—lateral mesoventral foveae; lmcf—lateral mesocoxal foveae; lmvf—lateral metaventral foveae; bef—basal foveae; ec—external ca- rina; mc—median carinae. Scale: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3835421" httpUri="https://zenodo.org/record/3835421/files/figure.png" pageId="2" pageNumber="369">Fig. 2e</figureCitation>
|
|||
|
|
)
|
|||
|
|
<quantity id="4C8A9B27A95ADF682CF8F976FD8B0996" box="[421,584,1684,1712]" metricMagnitude="-4" metricUnit="m" metricValue="6.8999999999999995" metricValueMax="8.4" metricValueMin="5.4" pageId="2" pageNumber="369" unit="mm" value="0.69" valueMax="0.84" valueMin="0.54">0.54–0.84 mm</quantity>
|
|||
|
|
long; first visible tergite (IV) large, wider than long; external carinae paral- lel to margin, orignate from exterior part of basolateral fovea; median carinae triangular, area between them with shallow depression; visible tergites 2–4 (V–VII) smallest, wider than long, lacking foveae or carinae; first visible sternite (IV) with two basolateral foveae.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BCD36C2A95ADF682D9AF8C4FCDF0819" blockId="2.[151,1437,1361,1964]" box="[199,796,1829,1856]" pageId="2" pageNumber="369">Legs long and slender. Hindwings present or lacking.</paragraph>
|
|||
|
|
<paragraph id="8BCD36C2A95ADF682D9AF8ABFE4908AE" blockId="2.[151,1437,1361,1964]" pageId="2" pageNumber="369">
|
|||
|
|
Aedegus (
|
|||
|
|
<figureCitation id="13492A47A95ADF682C67F8ABFEBD0845" box="[314,382,1865,1891]" captionStart="FIGURE 3" captionStartId="4.[151,250,1945,1970]" captionTargetBox="[287,1299,190,1910]" captionTargetId="figure@4.[273,1314,181,1919]" captionTargetPageId="4" captionText="FIGURE 3. Aedeagi of Protamamaurops, ventral view (Scale: 0.1 mm)—a, b) P. macedonicus—a. Bos Dag Mts., b—Belasitsa Mts; c) P. macrophthalmus; d) P. winneguthi; e) P. bulgaricus sp. n. Scale: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.3835423" httpUri="https://zenodo.org/record/3835423/files/figure.png" pageId="2" pageNumber="369">Fig. 3</figureCitation>
|
|||
|
|
) moderately sclerotized, symmetrical, basal capsule oval, apical lobe with large and slightly laterally curved apex.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BCD36C2A95ADF682D9AF872FB2E088D" blockId="2.[151,1437,1361,1964]" box="[199,1261,1936,1964]" pageId="2" pageNumber="369">
|
|||
|
|
<emphasis id="B906EAD0A95ADF682D9AF872FE73088D" bold="true" box="[199,432,1936,1963]" pageId="2" pageNumber="369">Sexual dimorphism</emphasis>
|
|||
|
|
. Males with metatrochanters modified, with one spine, in females simple.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<caption id="DF0D664AA95BDF692DCAF905FD8708B6" ID-DOI="http://doi.org/10.5281/zenodo.3835421" ID-Zenodo-Dep="3835421" httpUri="https://zenodo.org/record/3835421/files/figure.png" pageId="3" pageNumber="370" startId="3.[151,250,1767,1792]" targetBox="[154,1420,194,1741]" targetPageId="3">
|
|||
|
|
<paragraph id="8BCD36C2A95BDF692DCAF905FD8708B6" blockId="3.[151,1437,1767,1936]" pageId="3" pageNumber="370">
|
|||
|
|
<emphasis id="B906EAD0A95BDF692DCAF905FED70826" bold="true" box="[151,276,1767,1792]" pageId="3" pageNumber="370">FIGURE 2.</emphasis>
|
|||
|
|
Morphology of
|
|||
|
|
<taxonomicName id="4C724D41A95BDF692CE0F90BFD8D0826" authorityName="J. Muller" authorityYear="1944" box="[445,590,1769,1792]" class="Insecta" family="Staphylinidae" genus="Protamaurops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="370" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B906EAD0A95BDF692CE0F90BFD8D0826" box="[445,590,1769,1792]" italics="true" pageId="3" pageNumber="370">Protamaurops</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<taxonomicName id="4C724D41A95BDF692F00F90BFD380826" authorityName="J. Muller" authorityYear="1934" box="[605,763,1768,1792]" class="Insecta" family="Staphylinidae" genus="Protamaurops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="370" phylum="Arthropoda" rank="species" species="macedonicus">
|
|||
|
|
<emphasis id="B906EAD0A95BDF692F00F90BFD380826" box="[605,763,1768,1792]" italics="true" pageId="3" pageNumber="370">P. macedonicus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
)—
|
|||
|
|
<emphasis id="B906EAD0A95BDF692E40F905FCF00826" bold="true" box="[797,819,1767,1792]" pageId="3" pageNumber="370">a)</emphasis>
|
|||
|
|
Head (dorsal view);
|
|||
|
|
<emphasis id="B906EAD0A95BDF692951F905FBE00826" bold="true" box="[1036,1059,1767,1792]" pageId="3" pageNumber="370">b)</emphasis>
|
|||
|
|
Pronotum (dorsal and ventral view);
|
|||
|
|
<emphasis id="B906EAD0A95BDF692DCAF8E9FF680802" bold="true" box="[151,171,1803,1828]" pageId="3" pageNumber="370">c)</emphasis>
|
|||
|
|
Meso-and metaventrite;
|
|||
|
|
<emphasis id="B906EAD0A95BDF692CF5F8E9FE7C0802" bold="true" box="[424,447,1803,1828]" pageId="3" pageNumber="370">d)</emphasis>
|
|||
|
|
Elytra; e) Abdomen.Abbreviations: vf—vertexal fovea; laf—lateral antebasal foveae; maf—me- dian antebasal fovea; blf—basolateral foveae; lpcf—lateral procoxal foveae; mmvf—median mesoventral foveae; lmsf—lateral mesoventral foveae; lmcf—lateral mesocoxal foveae; lmvf—lateral metaventral foveae; bef—basal foveae; ec—external ca- rina; mc—median carinae. Scale: 0.1 mm.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="DF0D664AA95CDF6E2DCAF87BFC2908F3" ID-DOI="http://doi.org/10.5281/zenodo.3835423" ID-Zenodo-Dep="3835423" httpUri="https://zenodo.org/record/3835423/files/figure.png" pageId="4" pageNumber="371" startId="4.[151,250,1945,1970]" targetBox="[287,1299,190,1910]" targetPageId="4">
|
|||
|
|
<paragraph id="8BCD36C2A95CDF6E2DCAF87BFC2908F3" blockId="4.[151,1436,1945,2006]" pageId="4" pageNumber="371">
|
|||
|
|
<emphasis id="B906EAD0A95CDF6E2DCAF87BFED10894" bold="true" box="[151,274,1945,1970]" pageId="4" pageNumber="371">FIGURE 3.</emphasis>
|
|||
|
|
Aedeagi of
|
|||
|
|
<emphasis id="B906EAD0A95CDF6E2CD1F878FDF80897" box="[396,571,1946,1969]" italics="true" pageId="4" pageNumber="371">Protamamaurops</emphasis>
|
|||
|
|
, ventral view (Scale: 0.1 mm)—
|
|||
|
|
<emphasis id="B906EAD0A95CDF6E2ED8F878FC770894" bold="true" box="[901,948,1945,1970]" pageId="4" pageNumber="371">a, b)</emphasis>
|
|||
|
|
<emphasis id="B906EAD0A95CDF6E2EE7F878FBAE0894" box="[954,1133,1946,1970]" italics="true" pageId="4" pageNumber="371">P. macedonicus—</emphasis>
|
|||
|
|
a. Bos Dag Mts., b—Belasitsa Mts;
|
|||
|
|
<emphasis id="B906EAD0A95CDF6E2D8FF85FFF2508F0" bold="true" box="[210,230,1981,2006]" pageId="4" pageNumber="371">c)</emphasis>
|
|||
|
|
<taxonomicName id="4C724D41A95CDF6E2DB1F85CFE7208F0" authorityName="J. Muller" authorityYear="1944" box="[236,433,1982,2006]" class="Insecta" family="Staphylinidae" genus="Protamaurops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="371" phylum="Arthropoda" rank="species" species="macrophthalmus">
|
|||
|
|
<emphasis id="B906EAD0A95CDF6E2DB1F85CFF3808F3" box="[236,251,1982,2005]" italics="true" pageId="4" pageNumber="371">P</emphasis>
|
|||
|
|
.
|
|||
|
|
<emphasis id="B906EAD0A95CDF6E2C55F85CFE7208F0" box="[264,433,1982,2006]" italics="true" pageId="4" pageNumber="371">macrophthalmus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
;
|
|||
|
|
<emphasis id="B906EAD0A95CDF6E2CE2F85FFE1508F0" bold="true" box="[447,470,1981,2006]" pageId="4" pageNumber="371">d)</emphasis>
|
|||
|
|
<emphasis id="B906EAD0A95CDF6E2C81F85CFDAD08F0" box="[476,622,1982,2006]" italics="true" pageId="4" pageNumber="371">
|
|||
|
|
<taxonomicName id="4C724D41A95CDF6E2C81F85CFDAB08F0" authorityName="Szekessy" authorityYear="1943" box="[476,616,1982,2006]" class="Insecta" family="Staphylinidae" genus="Protamaurops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="371" phylum="Arthropoda" rank="species" species="winneguthi">P. winneguthi</taxonomicName>
|
|||
|
|
;
|
|||
|
|
</emphasis>
|
|||
|
|
<emphasis id="B906EAD0A95CDF6E2F29F85FFD4B08F0" bold="true" box="[628,648,1981,2006]" pageId="4" pageNumber="371">e)</emphasis>
|
|||
|
|
<taxonomicName id="4C724D41A95CDF6E2FD3F85CFCD008F0" authority="Bekchiev & Hlaváč, 2020" authorityName="Bekchiev & Hlaváč" authorityYear="2020" box="[654,787,1982,2006]" class="Insecta" family="Staphylinidae" genus="Protamaurops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="371" phylum="Arthropoda" rank="species" species="bulgaricus" status="sp. nov.">
|
|||
|
|
<emphasis id="B906EAD0A95CDF6E2FD3F85CFCD008F0" box="[654,787,1982,2006]" italics="true" pageId="4" pageNumber="371">P. bulgaricus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B906EAD0A95CDF6E2E47F85CFC9008F0" bold="true" box="[794,851,1982,2006]" pageId="4" pageNumber="371">
|
|||
|
|
<taxonomicNameLabel id="A23557ABA95CDF6E2E47F85CFC9008F0" box="[794,851,1982,2006]" pageId="4" pageNumber="371" rank="species">sp. n.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
Scale: 0.1 mm
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<subSubSection id="C3686549A95DDF6F2D9AFF7AFEAD0E39" pageId="5" pageNumber="372" type="biology_ecology">
|
|||
|
|
<paragraph id="8BCD36C2A95DDF6F2D9AFF7AFEAD0E39" blockId="5.[151,1437,152,683]" pageId="5" pageNumber="372">
|
|||
|
|
<emphasis id="B906EAD0A95DDF6F2D9AFF7AFEE20F95" bold="true" box="[199,289,152,179]" pageId="5" pageNumber="372">Biology</emphasis>
|
|||
|
|
.
|
|||
|
|
<taxonomicName id="4C724D41A95DDF6F2C73FF78FE0D0F95" authorityName="J. Muller" authorityYear="1944" box="[302,462,154,179]" class="Insecta" family="Staphylinidae" genus="Protamaurops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="372" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B906EAD0A95DDF6F2C73FF78FE0D0F95" box="[302,462,154,179]" italics="true" pageId="5" pageNumber="372">Protamaurops</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is most primitive and less specialised
|
|||
|
|
<taxonomicName id="4C724D41A95DDF6F2E2EFF7BFC390F95" authorityName="Jeannel" authorityYear="1948" box="[883,1018,153,179]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="372" phylum="Arthropoda" rank="tribe" tribe="Amauropini">Amauropini</taxonomicName>
|
|||
|
|
and from all other genera of the tribe, it is most similar to
|
|||
|
|
<taxonomicName id="4C724D41A95DDF6F2C2AFF5FFD440FF1" authority="Reitter, 1882" authorityName="Reitter" authorityYear="1882" box="[375,647,189,215]" class="Insecta" family="Staphylinidae" genus="Batrisodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="372" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B906EAD0A95DDF6F2C2AFF5FFE2C0FF1" box="[375,495,189,215]" italics="true" pageId="5" pageNumber="372">Batrisodes</emphasis>
|
|||
|
|
Reitter, 1882
|
|||
|
|
</taxonomicName>
|
|||
|
|
and it is only genus with rellatively well developed eyes and presense of wings. They can usually be collected by sifting leaf-litter, hand-collecting under stones or in ant nests in the mountains (
|
|||
|
|
<figureCitation id="13492A47A95DDF6F2C46FEE7FEA20E39" box="[283,353,261,287]" captionStart="FIGURE 4" captionStartId="6.[151,250,1730,1755]" captionTargetBox="[151,1436,741,1705]" captionTargetId="figure@6.[151,1436,741,1705]" captionTargetPageId="6" captionText="FIGURE 4. Habitat of Protamaurops macedonicus (Belasitsa Mts., Bulgaria)." figureDoi="http://doi.org/10.5281/zenodo.3835425" httpUri="https://zenodo.org/record/3835425/files/figure.png" pageId="5" pageNumber="372">Fig. 4</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3686549A95DDF6F2D9AFECAFC650E65" box="[199,934,296,323]" pageId="5" pageNumber="372" type="distribution">
|
|||
|
|
<paragraph id="8BCD36C2A95DDF6F2D9AFECAFC650E65" blockId="5.[151,1437,152,683]" box="[199,934,296,323]" pageId="5" pageNumber="372">
|
|||
|
|
<emphasis id="B906EAD0A95DDF6F2D9AFECAFE9A0E65" bold="true" box="[199,345,296,323]" pageId="5" pageNumber="372">Distribution</emphasis>
|
|||
|
|
. The Balkan Penisnula and
|
|||
|
|
<collectingCountry id="F3657652A95DDF6F2FCEFECBFD200E65" box="[659,739,297,323]" name="Turkey" pageId="5" pageNumber="372">Turkey</collectingCountry>
|
|||
|
|
(
|
|||
|
|
<tableCitation id="C6F00379A95DDF6F2FACFECBFC840E65" box="[753,839,297,323]" captionStart="TABLE 1" captionStartId="10.[151,239,1084,1108]" captionText="TABLE 1. List of species of genus Protamaurops." pageId="5" pageNumber="372">Table 1</tableCitation>
|
|||
|
|
;
|
|||
|
|
<figureCitation id="13492A47A95DDF6F2E09FECBFC5B0E65" box="[852,920,296,323]" captionStart="FIGURE 5" captionStartId="9.[151,250,1874,1899]" captionTargetBox="[151,1427,1145,1848]" captionTargetId="figure@9.[151,1436,1145,1848]" captionTargetPageId="9" captionText="FIGURE 5. Distribution of Protamaurops." figureDoi="http://doi.org/10.5281/zenodo.3835427" httpUri="https://zenodo.org/record/3835427/files/figure.png" pageId="5" pageNumber="372">Fig. 5</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3686549A95DDF6F2D9AFEAEFE3D0D8D" pageId="5" pageNumber="372" type="diagnosis">
|
|||
|
|
<paragraph id="8BCD36C2A95DDF6F2D9AFEAEFE3D0D8D" blockId="5.[151,1437,152,683]" pageId="5" pageNumber="372">
|
|||
|
|
<emphasis id="B906EAD0A95DDF6F2D9AFEAEFE810E41" bold="true" box="[199,322,332,359]" pageId="5" pageNumber="372">Diagnosis.</emphasis>
|
|||
|
|
As noted by
|
|||
|
|
<bibRefCitation id="EFE34B33A95DDF6F2C8DFEAFFDAD0E41" author="Muller, G." box="[464,622,333,359]" pageId="5" pageNumber="372" pagination="82 - 103" refId="ref6585" refString="Muller, G. (1944) Die Amaurops-Arten der Balkanhalbinsel. Mitteilungen der Munchner Entomologischen Gesellschaft, 34, 82 - 103." type="journal article" year="1944">Müller (1944)</bibRefCitation>
|
|||
|
|
,
|
|||
|
|
<bibRefCitation id="EFE34B33A95DDF6F2F25FEAFFCF10E4E" author="Karaman, Z." box="[632,818,333,360]" pageId="5" pageNumber="372" pagination="149 - 169" refId="ref6455" refString="Karaman, Z. (1961) Prilog Poznavanju Amauropsina Balkana. Beitrag zur Kenntnis der Amauropsinen des Balkans. Godisen Zbornik na Zemjodelsko-Sumarski Fakultet na Univerzitet, Skopje, 14, 149 - 169." type="journal article" year="1961">
|
|||
|
|
<collectingRegion id="49B6F820A95DDF6F2F25FEAFFD1C0E41" box="[632,735,333,359]" country="Turkey" name="Karaman" pageId="5" pageNumber="372">Karaman</collectingRegion>
|
|||
|
|
(1961)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
,
|
|||
|
|
<bibRefCitation id="EFE34B33A95DDF6F2E66FEAFFC370E4E" author="Besuchet, C." box="[827,1012,333,360]" pageId="5" pageNumber="372" pagination="257 - 264" refId="ref6155" refString="Besuchet, C. (1986) Synonymes et homonyme nouveaux de quelques genres de Pselaphides (Coleoptera). Revue Suisse de Zoologie, 93, 257 - 264. https: // doi. org / 10.5962 / bhl. part. 82019" type="journal article" year="1986">Besuchet (1986)</bibRefCitation>
|
|||
|
|
, and later by
|
|||
|
|
<bibRefCitation id="EFE34B33A95DDF6F29D9FEAFFF270EAD" author="Nonveiller, G. & Pavicevic, D." pageId="5" pageNumber="372" refId="ref6663" refString="Nonveiller, G. & Pavicevic, D. (2008) Une nouvelle espece et deux nouveaux sous-genres de Seracamaurops, avec un catalogue des Amauropini mediterraneens (Coleoptera, Staphylinidae, Pselaphinae) (Recherches sur les Coleopteres endoges et troglobiontes de Serbie et des regions Institute for Nature Conservation of Serbia, Belgrade, 564 pp." type="book" year="2008">Nonveiller and Pavićević (2008)</bibRefCitation>
|
|||
|
|
, this is a primitive genus, sharing some common characters with the tribe
|
|||
|
|
<taxonomicName id="4C724D41A95DDF6F2977FE93FAE40EAD" authority="Reitter, 1882" authorityName="Reitter" authorityYear="1882" box="[1066,1319,369,395]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="372" phylum="Arthropoda" rank="tribe" tribe="Batrisini">Batrisini Reitter, 1882</taxonomicName>
|
|||
|
|
. It is most closely related to the genus
|
|||
|
|
<taxonomicName id="4C724D41A95DDF6F2C8AFE74FCDB0E89" authority="Jeannel, 1948" authorityName="Jeannel" authorityYear="1948" box="[471,792,405,431]" class="Insecta" family="Staphylinidae" genus="Paramaurops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="372" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B906EAD0A95DDF6F2C8AFE74FDB30E89" box="[471,624,406,431]" italics="true" pageId="5" pageNumber="372">Paramaurops</emphasis>
|
|||
|
|
<bibRefCitation id="EFE34B33A95DDF6F2F24FE77FCDB0E89" author="Jeannel, R." box="[633,792,405,431]" pageId="5" pageNumber="372" pagination="1 - 19" refId="ref6340" refString="Jeannel, R. (1948) Revision des Amaurops et genres voisins (Pselaphidae). Revue Francaise d'Entomologie, 15 1 - 19." type="journal article" year="1948">Jeannel, 1948</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, sharing with this genus the presence of eyes and ocular spines, paralell-sided lateral carinae on the first visible tergite and well-defined medial longitudinal carinae.
|
|||
|
|
<taxonomicName id="4C724D41A95DDF6F281AFE58FF270ED1" authorityName="Jeannel" authorityYear="1948" class="Insecta" family="Staphylinidae" genus="Paramaurops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="372" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B906EAD0A95DDF6F281AFE58FF270ED1" italics="true" pageId="5" pageNumber="372">Paramaurops</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
differes from
|
|||
|
|
<taxonomicName id="4C724D41A95DDF6F2CDFFE3CFDE10ED1" authorityName="J. Muller" authorityYear="1944" box="[386,546,478,503]" class="Insecta" family="Staphylinidae" genus="Protamaurops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="372" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B906EAD0A95DDF6F2CDFFE3CFDE10ED1" box="[386,546,478,503]" italics="true" pageId="5" pageNumber="372">Protamaurops</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
by 1) elytra with completely atrophied basal foveae, 2) head with small and thin median carina, and 3) pronotum with small and thin lateral carinae. The genus
|
|||
|
|
<taxonomicName id="4C724D41A95DDF6F295FFDE0FB610D3D" authorityName="J. Muller" authorityYear="1944" box="[1026,1186,514,539]" class="Insecta" family="Staphylinidae" genus="Protamaurops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="372" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B906EAD0A95DDF6F295FFDE0FB610D3D" box="[1026,1186,514,539]" italics="true" pageId="5" pageNumber="372">Protamaurops</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
differes from all other members of the tribe
|
|||
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<taxonomicName id="4C724D41A95DDF6F2CDFFDC7FDCA0D19" authorityName="Jeannel" authorityYear="1948" box="[386,521,549,575]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="372" phylum="Arthropoda" rank="tribe" tribe="Amauropini">Amauropini</taxonomicName>
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by the following combination of characters: 1) eyes moderately to well devoloped above well-defined ocular spines; 2) carinae on the head and pronotum well-developed; 3) presence of two large basal fovea on the elytra; 4) humeri prominent; 5) external stria of the first abdominal tergite parallel to the tergal margin, median carinae present.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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