330 lines
33 KiB
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330 lines
33 KiB
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<document ID-DOI="10.37520/aemnp.2022.012" ID-ISSN="1804-6487" ID-Zenodo-Dep="7161302" ID-ZooBank="D3830DDC-6589-4B0C-8DEE-015053395254" approvalRequired="11" approvalRequired_for_textStreams="10" approvalRequired_for_treatments="1" checkinTime="1665202473490" checkinUser="felipe" docAuthor="Švec, Zdeněk" docDate="2022" docId="8C011006FFCCFFF3FCB6F9FDFD7CE170" docLanguage="en" docName="ActaEntMusNatPra.62.1.155-164.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 62 (1)" docSource="http://dx.doi.org/10.37520/aemnp.2022.012" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.6:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="6" docTitle="Agathidium (Macroceble) cornigerum Švec 2022, sp. nov." docType="treatment" docVersion="2" lastPageNumber="160" masterDocId="7038687EFFCFFFF6FFDAFFC5FFE0E700" masterDocTitle="Leiodidae (Coleoptera) of the Hainan Island with new faunistic records from China and with notes on the unique body modifications in the genus Agathidium" masterLastPageNumber="164" masterPageNumber="155" pageNumber="158" updateTime="1665202904673" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Leiodidae (Coleoptera) of the Hainan Island with new faunistic records from China and with notes on the unique body modifications in the genus Agathidium</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Švec, Zdeněk</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Acta Entomologica Musei Nationalis Pragae</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="series">
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<mods:title>Acta. Ent. Mus. Natl. Pragae</mods:title>
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</mods:detail>
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<mods:detail type="pubDate">
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<mods:number>2022-08-28</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>62</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>155</mods:start>
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<mods:end>164</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.37520/aemnp.2022.012</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.37520/aemnp.2022.012</mods:identifier>
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<mods:identifier type="ISSN">1804-6487</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7161302</mods:identifier>
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<mods:identifier type="ZooBank">D3830DDC-6589-4B0C-8DEE-015053395254</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:8C011006FFCCFFF3FCB6F9FDFD7CE170" httpUri="http://treatment.plazi.org/id/8C011006FFCCFFF3FCB6F9FDFD7CE170" lastPageId="5" lastPageNumber="160" pageId="3" pageNumber="158">
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<subSubSection box="[876,1388,1592,1616]" pageId="3" pageNumber="158" type="nomenclature">
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<paragraph blockId="3.[876,1388,1592,1616]" box="[876,1388,1592,1616]" pageId="3" pageNumber="158">
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<heading box="[876,1388,1592,1616]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="158" reason="3">
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<emphasis bold="true" box="[876,1388,1592,1616]" pageId="3" pageNumber="158">
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<taxonomicName authority="Švec, 2022" authorityName="Švec" authorityYear="2022" box="[876,1296,1592,1616]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="158" phylum="Arthropoda" rank="species" species="cornigerum" status="sp. nov." subGenus="Macroceble">
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<emphasis bold="true" box="[876,1006,1592,1616]" italics="true" pageId="3" pageNumber="158">Agathidium</emphasis>
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(
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<emphasis bold="true" box="[1022,1151,1592,1616]" italics="true" pageId="3" pageNumber="158">Macroceble</emphasis>
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)
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<emphasis bold="true" box="[1167,1296,1592,1616]" italics="true" pageId="3" pageNumber="158">cornigerum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1302,1388,1593,1616]" pageId="3" pageNumber="158" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[1056,1209,1623,1642]" pageId="3" pageNumber="158" type="description">
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<paragraph blockId="3.[828,1437,1623,2072]" box="[1056,1209,1623,1642]" pageId="3" pageNumber="158">
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(
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<figureCitation box="[1062,1139,1623,1642]" captionStart="Figs 1–8. 1, 2, 6, 7" captionStartId="3.[151,188,1016,1035]" captionTargetBox="[188,1406,191,997]" captionTargetId="figure-669@3.[158,1431,157,997]" captionTargetPageId="3" captionText="Figs 1–8. 1, 2, 6, 7 – aedeagus laterally and tip of aedeagus dorsally. 3, 5 – lateral view of body. 4, 8 – spermatheca. 1–4 – Agathidium (Agathidium) pilulum sp. nov.; 5–8 – Agathidium (Macroceble) cornigerum sp. nov. Scale bars in Figs 1, 2, 4, 6, 7, 8 = 0.1 mm; 3, 5 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7161304" httpUri="https://zenodo.org/record/7161304/files/figure.png" pageId="3" pageNumber="158">Figs 5–8</figureCitation>
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,
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<figureCitation box="[1148,1202,1623,1642]" captionStart="Figs 13–16. 13, 15, 16" captionStartId="5.[149,186,1096,1115]" captionTargetBox="[150,1435,189,1072]" captionTargetId="figure-486@5.[150,1435,189,1072]" captionTargetPageId="5" captionText="Figs 13–16. 13, 15, 16 – head dorsally; 14 – head laterally. 13 – Agathidium (Agathidium) pilulum sp. nov.; 14, 16 – Agathidium (Macroceble) cornigerum sp. nov.; 15 – Agathidium (Macroceble) apiforme sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7161308" httpUri="https://zenodo.org/record/7161308/files/figure.png" pageId="3" pageNumber="158">14, 16</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="158" type="materials_examined">
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<paragraph blockId="3.[828,1437,1623,2072]" pageId="3" pageNumber="158">
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<emphasis bold="true" box="[828,951,1657,1676]" pageId="3" pageNumber="158">
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<typeStatus box="[828,873,1657,1676]" pageId="3" pageNumber="158">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry box="[958,1011,1657,1676]" name="China" pageId="3" pageNumber="158">China</collectingCountry>
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,
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<collectingRegion box="[1022,1084,1657,1676]" country="China" name="Hainan" pageId="3" pageNumber="158">Hainan</collectingRegion>
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Island, Limushan Mts, 19°10.5–10.9′N, 109°44–45′E,
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<quantity box="[950,1049,1684,1703]" metricMagnitude="2" metricUnit="m" metricValue="7.75" metricValueMax="9.0" metricValueMin="6.5" pageId="3" pageNumber="158" unit="m" value="775.0" valueMax="900.0" valueMin="650.0">
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650–
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<specimenCount box="[993,1049,1684,1703]" pageId="3" pageNumber="158" type="generic">900 m</specimenCount>
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</quantity>
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.
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</paragraph>
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<paragraph blockId="3.[828,1437,1623,2072]" pageId="3" pageNumber="158">
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<emphasis bold="true" box="[828,960,1718,1737]" pageId="3" pageNumber="158">Type material.</emphasis>
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Hඈඅඈඍඒඉൾ: J, “
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<collectingCountry box="[1101,1157,1718,1737]" name="China" pageId="3" pageNumber="158">China</collectingCountry>
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:
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<collectingRegion box="[1165,1227,1718,1737]" country="China" name="Hainan" pageId="3" pageNumber="158">Hainan</collectingRegion>
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isl. [MF 19] / Limushan Mts., mountains / above first admin. centre / 19°10.5-10.9
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<superScript attach="right" box="[1341,1345,1743,1754]" fontSize="5" pageId="3" pageNumber="158">′</superScript>
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N 109°44- 45′E /
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<quantity box="[882,977,1771,1790]" metricMagnitude="2" metricUnit="m" metricValue="7.75" metricValueMax="9.0" metricValueMin="6.5" pageId="3" pageNumber="158" unit="m" value="775.0" valueMax="900.0" valueMin="650.0">650-900 m</quantity>
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,
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<date box="[985,1060,1771,1790]" pageId="3" pageNumber="158" value="2011-05-06">6.v.2011</date>
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, Fikáček // sifting small accumula- / tions of moist leaf litter along / and on the trail in secondary / forest partly with
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<taxonomicName box="[828,899,1825,1844]" class="Polypodiopsida" family="Cyatheaceae" genus="Cyathea" kingdom="Plantae" order="Cyatheales" pageId="3" pageNumber="158" phylum="Tracheophyta" rank="genus">
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<emphasis box="[828,899,1825,1844]" italics="true" pageId="3" pageNumber="158">Cyathea</emphasis>
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</taxonomicName>
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and / bamboo” (
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<collectionCode box="[1040,1105,1825,1844]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="3" pageNumber="158">NMPC</collectionCode>
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). Pൺඋൺඍඒඉൾඌ: 2 JJ
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<specimenCount box="[1271,1318,1818,1844]" pageId="3" pageNumber="158" type="female">5 ♀♀</specimenCount>
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, same data as
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<typeStatus box="[828,903,1851,1870]" pageId="3" pageNumber="158">holotype</typeStatus>
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(
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<collectionCode box="[914,978,1851,1870]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="3" pageNumber="158">NMPC</collectionCode>
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,
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<collectionCode box="[986,1041,1851,1870]" pageId="3" pageNumber="158">ZSPC</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="4" lastPageNumber="159" pageId="3" pageNumber="158" type="description">
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<paragraph blockId="3.[828,1437,1623,2072]" pageId="3" pageNumber="158">
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<emphasis bold="true" box="[828,963,1887,1910]" pageId="3" pageNumber="158">Description.</emphasis>
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Length of
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<typeStatus box="[1073,1164,1887,1910]" pageId="3" pageNumber="158">holotype</typeStatus>
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<quantity box="[1169,1250,1887,1910]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="158" unit="mm" value="1.6">1.6 mm</quantity>
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, head
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<quantity box="[1312,1390,1887,1910]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="158" unit="mm" value="0.2">0.2 mm</quantity>
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(
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<quantity metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="158" unit="mm" value="0.5">0.5 mm</quantity>
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including mandibular horn), pronotum
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<quantity box="[1284,1367,1919,1942]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="158" unit="mm" value="0.7">0.7 mm</quantity>
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, elytra
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<quantity box="[828,912,1951,1974]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="158" unit="mm" value="0.7">0.7 mm</quantity>
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, antenna
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<quantity box="[1014,1099,1951,1974]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="158" unit="mm" value="0.7">0.7 mm</quantity>
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, aedeagus
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<quantity box="[1216,1315,1951,1974]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" pageId="3" pageNumber="158" unit="mm" value="0.78">0.78 mm</quantity>
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. Maximum width of head
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<quantity box="[981,1065,1983,2006]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="158" unit="mm" value="0.9">0.9 mm</quantity>
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, pronotum
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<quantity box="[1186,1270,1983,2006]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="158" unit="mm" value="1.1">1.1 mm</quantity>
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, elytra
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<quantity box="[1348,1433,1983,2006]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="158" unit="mm" value="1.1">1.1 mm</quantity>
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. Body very shortly oval. Dorsum, antennomere 1 and legs light chestnut coloured, tarsi lighter; A2–A8
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.7161306" ID-Zenodo-Dep="7161306" httpUri="https://zenodo.org/record/7161306/files/figure.png" pageId="4" pageNumber="159" startId="4.[151,188,946,965]" targetBox="[242,1286,191,915]" targetPageId="4">
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<paragraph blockId="4.[151,1437,946,992]" pageId="4" pageNumber="159">
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Figs 9–12. 9, 10 – aedeagus laterally and tip of aedeagus dorsally. 11 – lateral view of body.12 – spermatheca. 9–11 –
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<taxonomicName authority="Švec, 2022" authorityName="Švec" authorityYear="2022" box="[1137,1437,946,965]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="159" phylum="Arthropoda" rank="species" species="apiforme" status="sp. nov." subGenus="Macroceble">
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<emphasis box="[1137,1237,946,965]" italics="true" pageId="4" pageNumber="159">Agathidium</emphasis>
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(
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<emphasis box="[1248,1349,946,965]" italics="true" pageId="4" pageNumber="159">Macroceble</emphasis>
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)
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<emphasis box="[1360,1437,946,965]" italics="true" pageId="4" pageNumber="159">apiforme</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[151,220,973,992]" pageId="4" pageNumber="159" rank="species">sp. nov.</taxonomicNameLabel>
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; 12 –
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<taxonomicName authority="Švec, 2022" authorityName="Švec" authorityYear="2022" box="[271,505,973,992]" class="Insecta" family="Leiodidae" genus="Dermatohomoeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="159" phylum="Arthropoda" rank="species" species="pennatus" status="sp. nov.">
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<emphasis box="[271,505,973,992]" italics="true" pageId="4" pageNumber="159">Dermatohomoeus pennatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[510,577,973,992]" pageId="4" pageNumber="159" rank="species">sp. nov.</taxonomicNameLabel>
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Scale bars in Figs 9, 10 = 0.2 mm; 11 = 0.5 mm; 12 = 0.1 mm.
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</paragraph>
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</caption>
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<paragraph blockId="4.[151,761,1090,2076]" pageId="4" pageNumber="159">yellowish, A9–A10 brown, A11 yellow-red. Ventral surface yellow-red. Dorsum punctured, without microsculpture. Sutural striae absent; femoral lines absent, metaventrite shortened, supraocular carina low all along its length.</paragraph>
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<paragraph blockId="4.[151,761,1090,2076]" pageId="4" pageNumber="159">
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Head very broad, maximum width of head just before posterior margin of eyes. Eyes, feebly flattened, somewhat drop-shaped (
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<figureCitation box="[296,375,1285,1308]" captionStart="Figs 13–16. 13, 15, 16" captionStartId="5.[149,186,1096,1115]" captionTargetBox="[150,1435,189,1072]" captionTargetId="figure-486@5.[150,1435,189,1072]" captionTargetPageId="5" captionText="Figs 13–16. 13, 15, 16 – head dorsally; 14 – head laterally. 13 – Agathidium (Agathidium) pilulum sp. nov.; 14, 16 – Agathidium (Macroceble) cornigerum sp. nov.; 15 – Agathidium (Macroceble) apiforme sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7161308" httpUri="https://zenodo.org/record/7161308/files/figure.png" pageId="4" pageNumber="159">Fig. 16</figureCitation>
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). Supraocular carina low with raised anterior edge of right carina forming triangular process. Subocular line not developed. Clypeus feebly roundly emarginate, clypeal line lacking. Antennomere 3 distinctly longer than A2 (A3/A2 = 1.7). Ratio of lengths of A2–A11 (A2 = 1.0): 1.0-1.7-0.9-0.9-0.7-0.9-0.9-1.4-1.1-2.6. Ratio of widths of A2–A11 (A2 = 1.0): 1.0-0.8-0.8-0.8-0.8-1.2- 1.4-2.0-2.2-2.4. Ratio of W/L of A2–A11: 0.7-0.3-0.7- 0.7-0.8-1.0-1.2-1.0-1.4-0.7. Surface of head smooth with very fine and very sparse punctures, separated by more than 10 times their own diameters. Left mandible bears long upwards oriented caudally bent horn (
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<figureCitation box="[609,733,1637,1660]" captionStart="Figs 13–16. 13, 15, 16" captionStartId="5.[149,186,1096,1115]" captionTargetBox="[150,1435,189,1072]" captionTargetId="figure-486@5.[150,1435,189,1072]" captionTargetPageId="5" captionText="Figs 13–16. 13, 15, 16 – head dorsally; 14 – head laterally. 13 – Agathidium (Agathidium) pilulum sp. nov.; 14, 16 – Agathidium (Macroceble) cornigerum sp. nov.; 15 – Agathidium (Macroceble) apiforme sp. nov." figureDoi="http://doi.org/10.5281/zenodo.7161308" httpUri="https://zenodo.org/record/7161308/files/figure.png" pageId="4" pageNumber="159">Figs 14, 16</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph blockId="4.[151,761,1090,2076]" pageId="4" pageNumber="159">
|
|||
|
|
Pronotum. Lateral outline broadly rounded in lateral view (
|
|||
|
|
<figureCitation box="[218,282,1701,1724]" captionStart="Figs 1–8. 1, 2, 6, 7" captionStartId="3.[151,188,1016,1035]" captionTargetBox="[188,1406,191,997]" captionTargetId="figure-669@3.[158,1431,157,997]" captionTargetPageId="3" captionText="Figs 1–8. 1, 2, 6, 7 – aedeagus laterally and tip of aedeagus dorsally. 3, 5 – lateral view of body. 4, 8 – spermatheca. 1–4 – Agathidium (Agathidium) pilulum sp. nov.; 5–8 – Agathidium (Macroceble) cornigerum sp. nov. Scale bars in Figs 1, 2, 4, 6, 7, 8 = 0.1 mm; 3, 5 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7161304" httpUri="https://zenodo.org/record/7161304/files/figure.png" pageId="4" pageNumber="159">Fig. 5</figureCitation>
|
|||
|
|
). Puncturation sparse and fine, rather similar to that on head with punctures separated by about 8–10 or even more times their own diameter.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph blockId="4.[151,761,1090,2076]" pageId="4" pageNumber="159">
|
|||
|
|
Elytra widest at lateral angles, then distinctly narrowed posteriorly. Elytral margins straight behind lateral angles in lateral view (
|
|||
|
|
<figureCitation box="[329,397,1861,1884]" captionStart="Figs 1–8. 1, 2, 6, 7" captionStartId="3.[151,188,1016,1035]" captionTargetBox="[188,1406,191,997]" captionTargetId="figure-669@3.[158,1431,157,997]" captionTargetPageId="3" captionText="Figs 1–8. 1, 2, 6, 7 – aedeagus laterally and tip of aedeagus dorsally. 3, 5 – lateral view of body. 4, 8 – spermatheca. 1–4 – Agathidium (Agathidium) pilulum sp. nov.; 5–8 – Agathidium (Macroceble) cornigerum sp. nov. Scale bars in Figs 1, 2, 4, 6, 7, 8 = 0.1 mm; 3, 5 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7161304" httpUri="https://zenodo.org/record/7161304/files/figure.png" pageId="4" pageNumber="159">Fig. 5</figureCitation>
|
|||
|
|
). Lateral elytral channel narrow. Puncturation similar to that on head.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph blockId="4.[151,761,1090,2076]" pageId="4" pageNumber="159">
|
|||
|
|
Legs. Tarsi slim, all tibiae slim, metafemora without specific characters. Tarsal formula: 5-5-
|
|||
|
|
<quantity box="[573,616,1957,1980]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="4" pageNumber="159" unit="in" value="4.0">4 in</quantity>
|
|||
|
|
male.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph blockId="4.[151,761,1090,2076]" pageId="4" pageNumber="159">Mesoventrite. Posterior part subconcave. Anterior part raised without longitudinal carina. Lateral lines not developed.</paragraph>
|
|||
|
|
<paragraph blockId="4.[828,1437,1089,2072]" pageId="4" pageNumber="159">Metaventrite. Distinctly shortened – MTW/MTLM = 3; MTW/MTLC = 6. Femoral lines missing.</paragraph>
|
|||
|
|
<paragraph blockId="4.[828,1437,1089,2072]" box="[858,1151,1153,1176]" pageId="4" pageNumber="159">Membranous wings absent.</paragraph>
|
|||
|
|
<paragraph blockId="4.[828,1437,1089,2072]" pageId="4" pageNumber="159">
|
|||
|
|
Genitalia. Basal part of median lobe simple (
|
|||
|
|
<typeStatus box="[1347,1394,1185,1208]" pageId="4" pageNumber="159">type</typeStatus>
|
|||
|
|
A). Apical part of median lobe narrow, elongate in long process rounded at its top in dorsal view. Operculum small, oval. Parameres flattened, triangularly widened at apex with flat oval apendices (
|
|||
|
|
<figureCitation box="[1000,1096,1313,1336]" captionStart="Figs 1–8. 1, 2, 6, 7" captionStartId="3.[151,188,1016,1035]" captionTargetBox="[188,1406,191,997]" captionTargetId="figure-669@3.[158,1431,157,997]" captionTargetPageId="3" captionText="Figs 1–8. 1, 2, 6, 7 – aedeagus laterally and tip of aedeagus dorsally. 3, 5 – lateral view of body. 4, 8 – spermatheca. 1–4 – Agathidium (Agathidium) pilulum sp. nov.; 5–8 – Agathidium (Macroceble) cornigerum sp. nov. Scale bars in Figs 1, 2, 4, 6, 7, 8 = 0.1 mm; 3, 5 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7161304" httpUri="https://zenodo.org/record/7161304/files/figure.png" pageId="4" pageNumber="159">Figs 6, 7</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph blockId="4.[828,1437,1089,2072]" pageId="4" pageNumber="159">
|
|||
|
|
<emphasis bold="true" box="[828,1214,1345,1368]" pageId="4" pageNumber="159">Variation and sexual dimorphism.</emphasis>
|
|||
|
|
Length of body
|
|||
|
|
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.9" metricValueMin="1.6" pageId="4" pageNumber="159" unit="mm" value="1.75" valueMax="1.9" valueMin="1.6">1.6– 1.9 mm</quantity>
|
|||
|
|
in the type series. The ratio A3/A2 varies between 1.2–1.7. One of the male
|
|||
|
|
<typeStatus box="[1103,1205,1409,1432]" pageId="4" pageNumber="159">paratypes</typeStatus>
|
|||
|
|
possesses simple left mandible without any horn, the other male
|
|||
|
|
<typeStatus box="[1271,1362,1441,1464]" pageId="4" pageNumber="159">paratype</typeStatus>
|
|||
|
|
is with large mandibular horn as in the
|
|||
|
|
<typeStatus box="[1172,1269,1473,1496]" pageId="4" pageNumber="159">holotype</typeStatus>
|
|||
|
|
. Tarsal formula 4-4-
|
|||
|
|
<quantity box="[871,913,1505,1528]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="4" pageNumber="159" unit="in" value="4.0">4 in</quantity>
|
|||
|
|
female. Spermatheca slim, cylindrical, S-shaped (
|
|||
|
|
<figureCitation box="[836,901,1537,1560]" captionStart="Figs 1–8. 1, 2, 6, 7" captionStartId="3.[151,188,1016,1035]" captionTargetBox="[188,1406,191,997]" captionTargetId="figure-669@3.[158,1431,157,997]" captionTargetPageId="3" captionText="Figs 1–8. 1, 2, 6, 7 – aedeagus laterally and tip of aedeagus dorsally. 3, 5 – lateral view of body. 4, 8 – spermatheca. 1–4 – Agathidium (Agathidium) pilulum sp. nov.; 5–8 – Agathidium (Macroceble) cornigerum sp. nov. Scale bars in Figs 1, 2, 4, 6, 7, 8 = 0.1 mm; 3, 5 = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7161304" httpUri="https://zenodo.org/record/7161304/files/figure.png" pageId="4" pageNumber="159">Fig. 8</figureCitation>
|
|||
|
|
). Length of spermatheca
|
|||
|
|
<quantity box="[1172,1270,1537,1560]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="159" unit="mm" value="0.15">0.15 mm</quantity>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection lastPageId="5" lastPageNumber="160" pageId="4" pageNumber="159" type="diagnosis">
|
|||
|
|
<paragraph blockId="4.[828,1437,1089,2072]" lastBlockId="5.[150,759,1209,1648]" lastPageId="5" lastPageNumber="160" pageId="4" pageNumber="159">
|
|||
|
|
<emphasis bold="true" box="[828,1078,1569,1592]" pageId="4" pageNumber="159">Differential diagnosis.</emphasis>
|
|||
|
|
<taxonomicName authority="Švec, 2022" authorityName="Švec" authorityYear="2022" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="159" phylum="Arthropoda" rank="species" species="cornigerum" status="sp. nov." subGenus="Macroceble">
|
|||
|
|
<emphasis box="[1086,1210,1569,1592]" italics="true" pageId="4" pageNumber="159">Agathidium</emphasis>
|
|||
|
|
(
|
|||
|
|
<emphasis box="[1227,1353,1569,1592]" italics="true" pageId="4" pageNumber="159">Macroceble</emphasis>
|
|||
|
|
)
|
|||
|
|
<emphasis italics="true" pageId="4" pageNumber="159">cornigerum</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel box="[903,987,1601,1624]" pageId="4" pageNumber="159" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
possesses greatly shortened metaventrite; femoral lines on metaventrite missing. Therefore, the new species is attributed to the subgenus
|
|||
|
|
<taxonomicName authority="Angelini, 1993" authorityName="Angelini" authorityYear="1993" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="159" phylum="Arthropoda" rank="subGenus" subGenus="Macroceble">
|
|||
|
|
<emphasis box="[1206,1332,1665,1688]" italics="true" pageId="4" pageNumber="159">Macroceble</emphasis>
|
|||
|
|
<bibRefCitation author="ANGELINI F." pageId="4" pageNumber="159" pagination="29 - 44" refId="ref8148" refString="ANGELINI F. 1993: Studi sugli Agathidium. Designazione di un nuovo genere, un nuovo sottogenere e gruppi di specie. Bollettino della Societa Entomologica Italiana 125: 29 - 44." type="journal article" year="1993">Angelini, 1993</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. It is habitually similar to
|
|||
|
|
<taxonomicName authority="Svec, 2017" authorityName="Svec" authorityYear="2017" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="159" phylum="Arthropoda" rank="species" species="pokornyi" subGenus="Macroceble">
|
|||
|
|
<emphasis box="[1162,1178,1697,1720]" italics="true" pageId="4" pageNumber="159">A</emphasis>
|
|||
|
|
. (
|
|||
|
|
<emphasis box="[1200,1326,1697,1720]" italics="true" pageId="4" pageNumber="159">Macroceble</emphasis>
|
|||
|
|
)
|
|||
|
|
<emphasis box="[1341,1436,1697,1720]" italics="true" pageId="4" pageNumber="159">pokornyi</emphasis>
|
|||
|
|
<bibRefCitation author="SVEC Z." box="[828,947,1729,1752]" pageId="4" pageNumber="159" pagination="485 - 497" refId="ref8959" refString="SVEC Z. 2017: Contribution to the knowledge of Chinese species of the genus Agathidium Panzer, 1797 (Coleoptera: Leiodidae: Leiodinae) - part III. Studies and Reports, Taxonomical Series 13: 485 - 497." type="book chapter" year="2017">Švec, 2017</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, a mountain species from
|
|||
|
|
<collectingRegion box="[1213,1297,1729,1752]" country="China" name="Yunnan" pageId="4" pageNumber="159">Yunnan</collectingRegion>
|
|||
|
|
,
|
|||
|
|
<collectingCountry box="[1306,1369,1729,1752]" name="China" pageId="4" pageNumber="159">China</collectingCountry>
|
|||
|
|
(Šඏൾർ 2017). Both species are similar in the shape of body, shape of head broadest at posterior part of eyes, feebly roundly emarginate clypeus, in dark antennal club with a little lighter colour of antennomere 11, both species are similar also in the length of A3 that is distinctly longer than A2.
|
|||
|
|
<taxonomicName authority="Švec, 2022" authorityName="Švec" authorityYear="2022" box="[828,1083,1921,1944]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="159" phylum="Arthropoda" rank="species" species="cornigerum" status="sp. nov.">
|
|||
|
|
<emphasis box="[828,1083,1921,1944]" italics="true" pageId="4" pageNumber="159">Agathidium cornigerum</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel box="[1090,1173,1921,1944]" pageId="4" pageNumber="159" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
differs from
|
|||
|
|
<taxonomicName authorityName="Svec" authorityYear="2017" box="[1312,1436,1921,1944]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="159" phylum="Arthropoda" rank="species" species="pokornyi">
|
|||
|
|
<emphasis box="[1312,1436,1921,1944]" italics="true" pageId="4" pageNumber="159">A. pokornyi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in lighter colouring of the dorsum, in the drop-shaped eyes that are parabola slice-shaped in
|
|||
|
|
<taxonomicName authorityName="Svec" authorityYear="2017" box="[1167,1289,1985,2008]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="159" phylum="Arthropoda" rank="species" species="pokornyi">
|
|||
|
|
<emphasis box="[1167,1289,1985,2008]" italics="true" pageId="4" pageNumber="159">A. pokornyi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and mainly in the median lobe constricted before rounded tip while the tegmen is linden leaf-shaped in
|
|||
|
|
<taxonomicName authorityName="Svec" authorityYear="2017" box="[1154,1274,2049,2072]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="159" phylum="Arthropoda" rank="species" species="pokornyi">
|
|||
|
|
<emphasis box="[1154,1274,2049,2072]" italics="true" pageId="4" pageNumber="159">A. pokornyi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. Also, the parameres are quite different as paramera is lacking appendix in
|
|||
|
|
<taxonomicName authorityName="Svec" authorityYear="2017" box="[150,274,1241,1264]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="160" phylum="Arthropoda" rank="species" species="pokornyi">
|
|||
|
|
<emphasis box="[150,274,1241,1264]" italics="true" pageId="5" pageNumber="160">A. pokornyi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. The shape of the parameres bearing appendices is very distinctive and very specific, distinguishing the new species from the known
|
|||
|
|
<taxonomicName box="[448,651,1305,1328]" pageId="5" pageNumber="160">
|
|||
|
|
<emphasis box="[448,570,1305,1328]" italics="true" pageId="5" pageNumber="160">Agathidium</emphasis>
|
|||
|
|
species
|
|||
|
|
</taxonomicName>
|
|||
|
|
except for
|
|||
|
|
<taxonomicName authority="Švec, 2022" authorityName="Švec" authorityYear="2022" box="[150,277,1337,1360]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="160" phylum="Arthropoda" rank="species" species="apiforme" status="sp. nov.">
|
|||
|
|
<emphasis box="[150,277,1337,1360]" italics="true" pageId="5" pageNumber="160">A. apiforme</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel box="[284,368,1337,1360]" pageId="5" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
that also possesses apical parameral appendices, but beside the shape of genitalia both species differ also in the morphological characters mentioned in the key above. Spermatheca is S-shaped in
|
|||
|
|
<taxonomicName authority="Švec, 2022" authorityName="Švec" authorityYear="2022" box="[606,759,1433,1456]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="160" phylum="Arthropoda" rank="species" species="cornigerum" status="sp. nov.">
|
|||
|
|
<emphasis box="[606,759,1433,1456]" italics="true" pageId="5" pageNumber="160">A. cornigerum</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel box="[150,234,1465,1488]" pageId="5" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
while the same has semi-circular basal part with curved cylindrical distal part in
|
|||
|
|
<taxonomicName authorityName="Svec" authorityYear="2017" box="[490,615,1497,1520]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="160" phylum="Arthropoda" rank="species" species="pokornyi">
|
|||
|
|
<emphasis box="[490,615,1497,1520]" italics="true" pageId="5" pageNumber="160">A. pokornyi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<caption ID-DOI="http://doi.org/10.5281/zenodo.7161308" ID-Zenodo-Dep="7161308" httpUri="https://zenodo.org/record/7161308/files/figure.png" pageId="5" pageNumber="160" startId="5.[149,186,1096,1115]" targetBox="[150,1435,189,1072]" targetPageId="5">
|
|||
|
|
<paragraph blockId="5.[149,1436,1096,1142]" pageId="5" pageNumber="160">
|
|||
|
|
Figs 13–16. 13, 15, 16 – head dorsally; 14 – head laterally. 13 –
|
|||
|
|
<taxonomicName authority="Švec, 2022" authorityName="Švec" authorityYear="2022" box="[680,966,1096,1115]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="160" phylum="Arthropoda" rank="species" species="pilulum" status="sp. nov." subGenus="Agathidium">
|
|||
|
|
<emphasis box="[680,779,1096,1115]" italics="true" pageId="5" pageNumber="160">Agathidium</emphasis>
|
|||
|
|
(
|
|||
|
|
<emphasis box="[790,889,1096,1115]" italics="true" pageId="5" pageNumber="160">Agathidium</emphasis>
|
|||
|
|
)
|
|||
|
|
<emphasis box="[901,966,1096,1115]" italics="true" pageId="5" pageNumber="160">pilulum</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel box="[970,1038,1096,1115]" pageId="5" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
; 14, 16 –
|
|||
|
|
<taxonomicName authority="Švec, 2022" authorityName="Švec" authorityYear="2022" box="[1115,1436,1096,1115]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="160" phylum="Arthropoda" rank="species" species="cornigerum" status="sp. nov." subGenus="Macroceble">
|
|||
|
|
<emphasis box="[1115,1214,1096,1115]" italics="true" pageId="5" pageNumber="160">Agathidium</emphasis>
|
|||
|
|
(
|
|||
|
|
<emphasis box="[1225,1326,1096,1115]" italics="true" pageId="5" pageNumber="160">Macroceble</emphasis>
|
|||
|
|
)
|
|||
|
|
<emphasis box="[1337,1436,1096,1115]" italics="true" pageId="5" pageNumber="160">cornigerum</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel box="[151,220,1123,1142]" pageId="5" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
; 15 –
|
|||
|
|
<taxonomicName authority="Švec, 2022" authorityName="Švec" authorityYear="2022" box="[270,573,1123,1142]" class="Insecta" family="Leiodidae" genus="Agathidium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="160" phylum="Arthropoda" rank="species" species="apiforme" status="sp. nov." subGenus="Macroceble">
|
|||
|
|
<emphasis box="[270,370,1123,1142]" italics="true" pageId="5" pageNumber="160">Agathidium</emphasis>
|
|||
|
|
(
|
|||
|
|
<emphasis box="[382,483,1123,1142]" italics="true" pageId="5" pageNumber="160">Macroceble</emphasis>
|
|||
|
|
)
|
|||
|
|
<emphasis box="[496,573,1123,1142]" italics="true" pageId="5" pageNumber="160">apiforme</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel box="[578,645,1123,1142]" pageId="5" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<subSubSection pageId="5" pageNumber="160" type="etymology">
|
|||
|
|
<paragraph blockId="5.[150,759,1209,1648]" pageId="5" pageNumber="160">
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<emphasis bold="true" box="[150,279,1529,1552]" pageId="5" pageNumber="160">Etymology.</emphasis>
|
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|
The name of the new species draws the attention to the well-developed horn on the left mandible of the
|
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|
<typeStatus box="[150,247,1593,1616]" pageId="5" pageNumber="160">holotype</typeStatus>
|
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|
. The name is derivate from Latin words
|
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|
|
<emphasis box="[677,758,1593,1616]" italics="true" pageId="5" pageNumber="160">cornum</emphasis>
|
|||
|
|
(= horn) and
|
|||
|
|
<emphasis box="[291,337,1625,1648]" italics="true" pageId="5" pageNumber="160">gere</emphasis>
|
|||
|
|
(= bear in English). Adjective.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|