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<document id="D53B11D89C5E7A178D370124C82FFDED" ID-CLB-Dataset="46304" ID-DOI="10.5281/zenodo.205904" ID-GBIF-Dataset="62376e09-f30b-4fb0-a635-8f8f43e008fd" ID-ISSN="1175-5326" ID-Zenodo-Dep="205904" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460391967158" checkinUser="plazi" docAuthor="Sasakawa, Mitsuhiro" docDate="2011" docId="03A787A1FFCAFFA2FF5EFE1FEB2A1A65" docLanguage="en" docName="zt03038p028.pdf" docOrigin="Zootaxa 3038" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tetrameringia maculata Sasakawa, 2011, sp. nov." docType="treatment" docVersion="14" lastPageNumber="16" masterDocId="FF9EFFD9FFC4FFADFFC9FFB5EF5D1878" masterDocTitle="Oriental species of clusiid flies (Diptera: Clusiidae: Clusiinae)" masterLastPageNumber="28" masterPageNumber="1" pageNumber="15" updateTime="1698298315006" updateUser="ExternalLinkService">
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<mods:titleInfo id="783629711AC2079F4A86072157F24BAA">
<mods:title id="42B3ECDABBF61A42D7F40CBE70676706">Oriental species of clusiid flies (Diptera: Clusiidae: Clusiinae)</mods:title>
</mods:titleInfo>
<mods:name id="EE17E80350A323E7582CED63A50B82ED" type="personal">
<mods:role id="F0B534A07E777C47FD7EDED8C49DEC6C">
<mods:roleTerm id="F86BB7CCFF33A990C296836F3DC6091C">Author</mods:roleTerm>
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<mods:namePart id="F4ECCC434074EF69FF18AF7993838BDF">Sasakawa, Mitsuhiro</mods:namePart>
</mods:name>
<mods:typeOfResource id="333BB2C2D43CD435B0EB1285E5DF10FC">text</mods:typeOfResource>
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<mods:title id="6C811610819E0B55E4A0F8DC8361C030">Zootaxa</mods:title>
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<mods:part id="3C511664C71EFDC6BBC09F4292EB92B4">
<mods:date id="FBDF489E8F1A03F0B422BFD0C4AD83A6">2011</mods:date>
<mods:detail id="00AF329AD1485CE62233D60E93BB4062" type="volume">
<mods:number id="B1E249F0AF8EB42200CB4CC0C736286F">3038</mods:number>
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<mods:classification id="FB847FB48D3D64D52C994934AD3462C7">journal article</mods:classification>
<mods:identifier id="59FD36D183E95306D513F4F3AD739FF1" type="CLB-Dataset">46304</mods:identifier>
<mods:identifier id="A684A3D67DFA206C637B6E8759E7A120" type="DOI">10.5281/zenodo.205904</mods:identifier>
<mods:identifier id="53FC864E25A9D650FB916F73230E39EE" type="GBIF-Dataset">62376e09-f30b-4fb0-a635-8f8f43e008fd</mods:identifier>
<mods:identifier id="99FD4A604B2AA3BBF58888ADE5D6219B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="54F271706EDDBF65714C67365BD66D5C" type="Zenodo-Dep">205904</mods:identifier>
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<treatment id="03A787A1FFCAFFA2FF5EFE1FEB2A1A65" ID-DOI="http://doi.org/10.5281/zenodo.3509788" ID-GBIF-Taxon="119416980" ID-Zenodo-Dep="3509788" LSID="urn:lsid:plazi:treatment:03A787A1FFCAFFA2FF5EFE1FEB2A1A65" httpUri="http://treatment.plazi.org/id/03A787A1FFCAFFA2FF5EFE1FEB2A1A65" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<subSubSection id="C314653CFFCAFFA3FF5EFE1FEE7A199F" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="8BB136B7FFCAFFA3FF5EFE1FED5019BB" blockId="14.[151,525,426,487]" box="[151,525,426,451]" pageId="14" pageNumber="15">
<heading id="D0F981DBFFCAFFA3FF5EFE1FED5019BB" bold="true" box="[151,525,426,451]" fontSize="10" level="2" pageId="14" pageNumber="15" reason="2">
<emphasis id="B97AEAA5FFCAFFA3FF5EFE1FED5019BB" bold="true" box="[151,525,426,451]" pageId="14" pageNumber="15">
<taxonomicName id="4C0E4D34FFCAFFA3FF5EFE1FEEF019BB" ID-CoL="8FDSY" box="[151,429,426,451]" class="Insecta" family="Clusiidae" genus="Tetrameringia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="maculata" status="sp. nov.">
<emphasis id="B97AEAA5FFCAFFA3FF5EFE1FEEF019BB" bold="true" box="[151,429,426,451]" italics="true" pageId="14" pageNumber="15">Tetrameringia maculata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24957DEFFCAFFA3FE7DFE1EED5019BB" box="[436,525,427,451]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB136B7FFCAFFA3FF5EFE7AEE7A199F" blockId="14.[151,525,426,487]" box="[151,295,463,487]" pageId="14" pageNumber="15">
(
<figureCitation id="13352A32FFCAFFA3FF56FE7AEE43199F" box="[159,286,463,487]" captionStart="FIGURES 19 – 24" captionStartId="13.[151,264,1976,1999]" captionTargetBox="[194,1382,198,1918]" captionTargetId="figure@13.[186,1393,193,1928]" captionTargetPageId="13" captionText="FIGURES 19 – 24. Male genitalia of Sobarocephala megastylis sp. nov. (19 – 22, holotype) and Tetrameringia maculata sp. nov. (23, 24, paratype). See Figs 1 – 3 for explanation of parts. Scale = 0.1 mm." httpUri="https://zenodo.org/record/205908/files/figure.png" pageId="14" pageNumber="15">Figs 23, 24</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C314653CFFCAFFA3FF54FDA5EEC51AEC" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="8BB136B7FFCAFFA3FF54FDA5EEC51AEC" blockId="14.[151,1437,527,2036]" pageId="14" pageNumber="15">
<emphasis id="B97AEAA5FFCAFFA3FF54FDA5EE161A50" bold="true" box="[157,331,527,552]" pageId="14" pageNumber="15">
<typeStatus id="54B58815FFCAFFA3FF54FDA5EF881A50" box="[157,213,528,552]" pageId="14" pageNumber="15">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54B58815FFCAFFA3FE98FDA5EEE41A50" box="[337,441,528,552]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="ED1FAE72FFCAFFA3FDCDFDA5ED0E1A50" LSID="urn:lsid:biocol.org:col:34697" box="[516,595,528,552]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34697" name="Bishop Museum" pageId="14" pageNumber="15">BPBM</collectionCode>
17246),
<collectingCountry id="F3197627FFCAFFA3FD7EFDA4EDAD1A50" box="[695,752,529,552]" name="Laos" pageId="14" pageNumber="15">Laos</collectingCountry>
, Vientiane Prov., Ban Van Eue,
<date id="FFB01077FFCAFFA3FB9AFDA5EB821A50" box="[1107,1247,528,552]" pageId="14" pageNumber="15" value="1966-04-29">29. IV. 1966</date>
, native collector, light trap (Rondon-Bishop collection).
<typeStatus id="54B58815FFCAFFA3FD9BFD80ED9E1A35" box="[594,707,565,589]" pageId="14" pageNumber="15" type="paratype">Paratypes</typeStatus>
: 13, 5Ƥ, same data as
<typeStatus id="54B58815FFCAFFA3FC06FD81EB681A34" box="[975,1077,564,588]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
; 6Ƥ, same locality as
<typeStatus id="54B58815FFCAFFA3FAFAFD81EAC51A34" box="[1331,1432,564,588]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
,
<date id="FFB01077FFCAFFA3FF5EFDEDEE191A08" box="[151,324,600,624]" pageId="14" pageNumber="15" value="1965-05-15" valueMax="1965-05-31" valueMin="1965-05-15">15–31. V. 1965</date>
,
<date id="FFB01077FFCAFFA3FE86FDEDEE8E1A08" box="[335,467,600,624]" pageId="14" pageNumber="15" value="1967-05-15">15. V. 1967</date>
, and
<date id="FFB01077FFCAFFA3FDC4FDEDEDC31A08" box="[525,670,600,624]" pageId="14" pageNumber="15" value="1967-09-15">15. IX. 1967</date>
, native collector; Ƥ, same locality as
<typeStatus id="54B58815FFCAFFA3FB8BFDEDEBFA1A08" box="[1090,1191,600,624]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
,
<date id="FFB01077FFCAFFA3FB7BFDEDEA631A08" box="[1202,1342,600,624]" pageId="14" pageNumber="15" value="1966-04-29">29. IV. 1966</date>
, native collector, Malaise trap.
</paragraph>
</subSubSection>
<subSubSection id="C314653CFFCAFFA3FF0EFD2AEC191AA4" pageId="14" pageNumber="15" type="diagnosis">
<paragraph id="8BB136B7FFCAFFA3FF0EFD2AEC191AA4" blockId="14.[151,1437,527,2036]" pageId="14" pageNumber="15">
<emphasis id="B97AEAA5FFCAFFA3FF0EFD2AEE1C1AC0" bold="true" box="[199,321,671,696]" pageId="14" pageNumber="15">Diagnosis.</emphasis>
This species is characterized by having two dark spots on the mesoscutum, a largely black anepisternum, three pairs of sc, and an inverted-L-shaped surstylus.
</paragraph>
</subSubSection>
<subSubSection id="C314653CFFCAFFA3FF0EFD52EEEC1F8C" pageId="14" pageNumber="15" type="description">
<paragraph id="8BB136B7FFCAFFA3FF0EFD52ED0F1C3F" blockId="14.[151,1437,527,2036]" pageId="14" pageNumber="15">
<emphasis id="B97AEAA5FFCAFFA3FF0EFD52EE041B78" bold="true" box="[199,345,743,768]" pageId="14" pageNumber="15">Description.</emphasis>
MALE. Head yellow; frons shining, brown-tinged on dorsal half; ocellar triangle brown. Antenna and palpus brownish yellow, first antennal flagellomere brown-tinged surrounding base of black arista. Thorax brownish yellow; mesoscutum weakly shining, with pair of brownish-black, oval spots between dc- and salines (spots 2/3–3/4 as long as distance between transverse and scutoscutellar sutures, extending slightly inside of dc-lines but not extending to sa- and pa-bases); anepisternum posteriorly with shining black spot (this 2/3 as high as and 1/2 as wide as sclerite), anatergite and katatergite brown; scutellum yellow, with pale brown, anterolateral spots before bases of lsc. Wing faintly infuscated on apical 1/3, fading behind M1 or CuA1; calypter with margin and fringe yellowish brown; halter yellow. Legs yellow. Abdomen entirely black, shining (
<typeStatus id="54B58815FFCAFFA3FB55FC52EA591B87" box="[1180,1284,999,1023]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
), but in
<typeStatus id="54B58815FFCAFFA3FAABFC5DEF9A1C5C" pageId="14" pageNumber="15" type="paratype">paratype</typeStatus>
male, T1 and T2 yellowish brown, T3–5 brown and blackened laterally, T6 and S8 black. Bristles on head and thorax brown, those on abdomen black.
</paragraph>
<paragraph id="8BB136B7FFCAFFA3FF0EFBE1EC9E1D67" blockId="14.[151,1437,527,2036]" pageId="14" pageNumber="15">
Frons as wide as eye, almost parallel-sided; frontalia sparsely setulose on ventral part; ventral part of parafrontalia at level of antennal base slightly projecting above eye in profile; or three, second or shorter than others; oc and poc subequal in length, slightly shorter than second or; eye 1.2 times as high as broad; gena 1/7–1/5 as high as eye;
<date id="FFB01077FFCAFFA3FF5EFB0AEF9F1CAF" box="[151,194,1215,1239]" pageId="14" pageNumber="15">vi 2</date>
/3 as long as ivt: pm six. First antennal flagellomere shorter than broad, with long, whitish pile; arista plumose but more sparsely so on apical 1/3, with longest pubescence about 1/3 as long as width of first flagellomere; palpus slightly wider than genal height, nearly 2/3 as wide as first flagellomere.
</paragraph>
<paragraph id="8BB136B7FFCAFFA3FF0EFA99EE8E1E65" blockId="14.[151,1437,527,2036]" pageId="14" pageNumber="15">Mesoscutum with three dc, first dc slightly longer than acr and 1/3 as long as third dc, second dc 1/2–2/3 as long as third dc and situated behind level of sa; acr in six or seven irregular rows; prs subequal to or shorter than second dc; prsc 1/2 as long as third dc; scutellum with three pairs of sc: dsc nearly twice as long as lsc, asc twice as long as dsc. Wing 2.9–3.0 (3.0) mm long; costa with three sections in proportion of 67: 15: 10, r-m before middle level of cell dm (15: 20), M1 ratio about 2.5, ultimate section of CuA1 slightly shorter than 1/2 length of penultimate section (8: 18). Fore femur with several short fb; fore and mid tibiae each with very short dp, mid tibia with one long and one short spur.</paragraph>
<paragraph id="8BB136B7FFCAFFA3FF0EF99CEFAE1EA9" blockId="14.[151,1437,527,2036]" pageId="14" pageNumber="15">
Epandrium with surstylus (
<figureCitation id="13352A32FFCAFFA3FE33F99CED101E39" box="[506,589,1577,1601]" captionStart="FIGURES 19 – 24" captionStartId="13.[151,264,1976,1999]" captionTargetBox="[194,1382,198,1918]" captionTargetId="figure@13.[186,1393,193,1928]" captionTargetPageId="13" captionText="FIGURES 19 – 24. Male genitalia of Sobarocephala megastylis sp. nov. (19 – 22, holotype) and Tetrameringia maculata sp. nov. (23, 24, paratype). See Figs 1 – 3 for explanation of parts. Scale = 0.1 mm." httpUri="https://zenodo.org/record/205908/files/figure.png" pageId="14" pageNumber="15">Fig. 24</figureCitation>
) inverted-L-shaped, bearing minute apical spinule; cercus small; proctiger Y-shaped. Phallapodeme 400 µm long; pregonite with seven setulae apically, postgonite with six setulae. Basiphallus (
<figureCitation id="13352A32FFCAFFA3FF01F9C4EE471EF1" box="[200,282,1649,1673]" captionStart="FIGURES 19 – 24" captionStartId="13.[151,264,1976,1999]" captionTargetBox="[194,1382,198,1918]" captionTargetId="figure@13.[186,1393,193,1928]" captionTargetPageId="13" captionText="FIGURES 19 – 24. Male genitalia of Sobarocephala megastylis sp. nov. (19 – 22, holotype) and Tetrameringia maculata sp. nov. (23, 24, paratype). See Figs 1 – 3 for explanation of parts. Scale = 0.1 mm." httpUri="https://zenodo.org/record/205908/files/figure.png" pageId="14" pageNumber="15">Fig. 23</figureCitation>
) short; mesophallus rather long, curved at distal end, striated on distal part; distiphallus 460 µm long, almost membranous, with microscopic scales on basal part; paraphallus very narrow, with right sclerite longer than left one.
</paragraph>
<paragraph id="8BB136B7FFCAFFA3FF0EF968ED7F1E8D" blockId="14.[151,1437,527,2036]" box="[199,546,1757,1781]" pageId="14" pageNumber="15">Body length 3.0–3.5 (3.5) mm.</paragraph>
<paragraph id="8BB136B7FFCAFFA3FF0EF8B7EEEC1F8C" blockId="14.[151,1437,527,2036]" pageId="14" pageNumber="15">
FEMALE. Similar to male, but dark spots on mesoscutum broader than those of male, extending to lateral margins of scutum; anespisternum almost entirely black, with anterodorsal corner yellow; brown area of anatergite and katatergite extending anteriorly to anepimeron and posteriorly to lateral margin of mediotergite; hind tibia pale brown on basal half; T1 and T2 yellowish brown but blackened laterally, T3 and T4 black with posterior margins yellowish, T5 and T6 entirely black, T7 brown at base and yellowish posteriorly; ovipositor brownish yellow, with base brown; wing length
<quantity id="4CF69B52FFCAFFA3FE71F800ED1E1FB5" box="[440,579,1973,1997]" metricMagnitude="-3" metricUnit="m" metricValue="3.5999999999999996" metricValueMax="3.8" metricValueMin="3.4" pageId="14" pageNumber="15" unit="mm" value="3.5999999999999996" valueMax="3.8" valueMin="3.4">3.4–3.8 mm</quantity>
, second and third costal sections in proportion of 75: 15, M1 ratio 2.4–2.9; body length
<quantity id="4CF69B52FFCAFFA3FEEAF869EEF01F8C" box="[291,429,2012,2036]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="4.3" metricValueMin="3.2" pageId="14" pageNumber="15" unit="mm" value="3.75" valueMax="4.3" valueMin="3.2">3.2–4.3 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C314653CFFCBFFA2FF0EFF22EEF818D7" box="[199,421,151,176]" pageId="15" pageNumber="16" type="distribution">
<paragraph id="8BB136B7FFCBFFA2FF0EFF22EEF818D7" blockId="15.[151,1436,151,542]" box="[199,421,151,176]" pageId="15" pageNumber="16">
<emphasis id="B97AEAA5FFCBFFA2FF0EFF22EE3D18C8" bold="true" box="[199,352,151,176]" pageId="15" pageNumber="16">Distribution.</emphasis>
<collectingCountry id="F3197627FFCBFFA2FEAEFF2DEEFC18D7" box="[359,417,152,175]" name="Laos" pageId="15" pageNumber="16">Laos</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C314653CFFCBFFA2FF0EFF09EC321981" pageId="15" pageNumber="16" type="discussion">
<paragraph id="8BB136B7FFCBFFA2FF0EFF09EC321981" blockId="15.[151,1436,151,542]" pageId="15" pageNumber="16">
<emphasis id="B97AEAA5FFCBFFA2FF0EFF09EE6618AC" bold="true" box="[199,315,188,212]" pageId="15" pageNumber="16">Remarks.</emphasis>
The male genitalia of this new species are almost similar in appearance to those of its known congeners. However, this species is clearly distinguished from the Australian
<taxonomicName id="4C0E4D34FFCBFFA2FC75FF54EBE11880" box="[956,1212,224,248]" class="Insecta" family="Clusiidae" genus="Tetrameringia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="ustulata">
<emphasis id="B97AEAA5FFCBFFA2FC75FF54EBE11880" box="[956,1212,224,248]" italics="true" pageId="15" pageNumber="16">Tetrameringia ustulata</emphasis>
</taxonomicName>
and the Japanese
<taxonomicName id="4C0E4D34FFCBFFA2FA4EFF54EEF51964" authority="Sueyoshi, 2006" authorityName="Sueyoshi" authorityYear="2006" class="Insecta" family="Clusiidae" genus="Tetrameringia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="borealis">
<emphasis id="B97AEAA5FFCBFFA2FA4EFF54EFAF1964" italics="true" pageId="15" pageNumber="16">T. borealis</emphasis>
<bibRefCitation id="EF9F4B46FFCBFFA2FF31FEB1EEF51964" author="Sueyoshi" box="[248,424,260,284]" pageId="15" pageNumber="16" refString="Sueyoshi, M. (2006) Species diversity of Japanese Clusiidae (Diptera: Acalyptrata) with description of 12 new species. Annales de la Societe Entomologique de France (n. s.), 42, 1 - 26." type="journal article" year="2006">Sueyoshi, 2006</bibRefCitation>
</taxonomicName>
, by the coloration of the mesoscutum and wing, the numbers of or and sc, and the plumosity of the arista. In
<taxonomicName id="4C0E4D34FFCBFFA2FEA6FE9CEEBB1938" box="[367,486,296,320]" class="Insecta" family="Clusiidae" genus="Tetrameringia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="ustulata">
<emphasis id="B97AEAA5FFCBFFA2FEA6FE9CEEBB1938" box="[367,486,296,320]" italics="true" pageId="15" pageNumber="16">T. ustulata</emphasis>
</taxonomicName>
, the mesoscutum is unmarked and the wing has a blackish mark extending from the costa to CuA1 across the end of cell dm in addition to the anteroapical mark, or are four, lsc are three, and the longest pubescence on the arista is about 1/5 as long as the width of the first flagellomere (
<bibRefCitation id="EF9F4B46FFCBFFA2FBA7FEC7EAD019F2" author="McAlpine" box="[1134,1421,370,394]" pageId="15" pageNumber="16" refString="McAlpine, D. K. (1960) A review of the Australian species of Clusiidae (Diptera, Acalyptrata). Records of the Australian Museum, 25, 63 - 94." type="journal article" year="1960">McAlpine 1960: figs 1, 5</bibRefCitation>
); in
<taxonomicName id="4C0E4D34FFCBFFA2FF7CFE2DEE7119D7" box="[181,300,407,431]" class="Insecta" family="Clusiidae" genus="Tetrameringia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="borealis">
<emphasis id="B97AEAA5FFCBFFA2FF7CFE2DEF9F19D7" box="[181,194,408,431]" italics="true" pageId="15" pageNumber="16">T</emphasis>
.
<emphasis id="B97AEAA5FFCBFFA2FF1BFE22EE7119D7" box="[210,300,407,431]" italics="true" pageId="15" pageNumber="16">borealis</emphasis>
</taxonomicName>
, the mesoscutum is brown bivittate along lateral margin of the scutum, with the marking extending onto the scutellum, the anteroapical marking on the wing is smaller (not beyond R4+5;
<bibRefCitation id="EF9F4B46FFCBFFA2FB93FE0FEA5919AA" author="Sueyoshi" box="[1114,1284,442,466]" pageId="15" pageNumber="16" refString="Sueyoshi, M. (2006) Species diversity of Japanese Clusiidae (Diptera: Acalyptrata) with description of 12 new species. Annales de la Societe Entomologique de France (n. s.), 42, 1 - 26." type="journal article" year="2006">Sueyoshi 2006</bibRefCitation>
: fig. 9H), bsc and lsc are absent, and the arista is sparsely and shortly plumose.
</paragraph>
</subSubSection>
<subSubSection id="C314653CFFCBFFA2FF0EFDB3EB2A1A65" box="[199,1143,517,542]" pageId="15" pageNumber="16" type="etymology">
<paragraph id="8BB136B7FFCBFFA2FF0EFDB3EB2A1A65" blockId="15.[151,1436,151,542]" box="[199,1143,517,542]" pageId="15" pageNumber="16">
<emphasis id="B97AEAA5FFCBFFA2FF0EFDB3EE131A66" bold="true" box="[199,334,518,542]" pageId="15" pageNumber="16">Etymology.</emphasis>
The specific name refers to the spotted mesoscutum (Latin:
<emphasis id="B97AEAA5FFCBFFA2FC3AFDB0EB3B1A65" box="[1011,1126,517,541]" italics="true" pageId="15" pageNumber="16">maculatus</emphasis>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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