301 lines
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301 lines
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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.222.87779" ID-Pensoft-Pub="1314-2003-222-75" ID-Pensoft-UUID="348E6053A6D55787993694B23682FDE9" ModsDocID="1314-2003-222-75" checkinTime="1679617121532" checkinUser="pensoft" docAuthor="Maas, Paul J. M., Maas-van de Kamer, Hiltje, Andre, Thiago, Skinner, David, Valderrama, Eugenio & Specht, Chelsea D." docDate="2023" docId="5B398AFCE2A95824B19C58E6A19DE6EB" docLanguage="en" docName="PhytoKeys 222: 75-127" docOrigin="PhytoKeys 222" docPubDate="2023-03-22" docSource="http://dx.doi.org/10.3897/phytokeys.222.87779" docTitle="Costus alticolus Maas & H. Maas 2023, sp. nov." docType="treatment" docVersion="1" id="348E6053A6D55787993694B23682FDE9" lastPageNumber="75" masterDocId="348E6053A6D55787993694B23682FDE9" masterDocTitle="Eighteen new species of Neotropical Costaceae (Zingiberales)" masterLastPageNumber="127" masterPageNumber="75" pageNumber="75" updateTime="1679617121532" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Eighteen new species of Neotropical Costaceae (Zingiberales)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Maas, Paul J. M.</mods:namePart>
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<mods:affiliation>Naturalis Biodiversity Centre, Botany, P. O. Box 9517, 2300 RA Leiden, Netherlands</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Maas-van de Kamer, Hiltje</mods:namePart>
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<mods:affiliation>Naturalis Biodiversity Centre, Botany, P. O. Box 9517, 2300 RA Leiden, Netherlands</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Andre, Thiago</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4148-3662</mods:nameIdentifier>
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<mods:affiliation>Universidade de Brasilia, Departamento de Botanica, Campus Universitario Darcy Ribeiro, Asa Norte, Brasilia (DF), Brazil</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Skinner, David</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4585-925X</mods:nameIdentifier>
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<mods:affiliation>Le Jardin Ombrage, Tallahassee, (Private botanical garden, Botanic Gardens Conservation International - BGCI - registration ID 50148), Florida, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Valderrama, Eugenio</mods:namePart>
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<mods:affiliation>Cornell University, Section of Plant Biology and the L. H. Bailey Hortorium, School of Integrative Plant Science, Ithaca, NY, USA</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Specht, Chelsea D.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7746-512X</mods:nameIdentifier>
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<mods:affiliation>Cornell University, Section of Plant Biology and the L. H. Bailey Hortorium, School of Integrative Plant Science, Ithaca, NY, USA</mods:affiliation>
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<mods:nameIdentifier type="email">cdspecht@cornell.edu</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-03-22</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>222</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>75</mods:start>
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<mods:end>127</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.222.87779</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.222.87779</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-222-75</mods:identifier>
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<mods:identifier type="Pensoft-UUID">348E6053A6D55787993694B23682FDE9</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:5B398AFCE2A95824B19C58E6A19DE6EB" httpUri="http://treatment.plazi.org/id/5B398AFCE2A95824B19C58E6A19DE6EB" lastPageNumber="75" pageId="0" pageNumber="75">
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<subSubSection pageId="0" pageNumber="75" type="nomenclature">
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<paragraph pageId="0" pageNumber="75">
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<taxonomicName LSID="5B398AFC-E2A9-5824-B19C-58E6A19DE6EB" authority="Maas & H. Maas" authorityName="Maas & H. Maas" authorityYear="2023" class="Liliopsida" family="Costaceae" genus="Costus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Costus alticolus" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="alticolus" status="sp. nov.">Costus alticolus Maas & H.Maas</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="75">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="75" type="diagnosis">
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<paragraph pageId="0" pageNumber="75">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="75">
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<taxonomicName authorityName="Maas & H. Maas" authorityYear="2023" class="Liliopsida" family="Costaceae" genus="Costus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Costus alticolus" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="alticolus">
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<emphasis italics="true" pageId="0" pageNumber="75">Costus alticolus</emphasis>
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</taxonomicName>
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sp. nov. (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Costus alticolus Maas & H. Maas A inflorescence with flower, showing stamen (st) extending far beyond labellum, fully exposing the thecae B, C plant in habitat D flower and bract details E flowering shoot. Photos in habitat in Chinantla region near Santa Cruz Tepetotutla, Oaxaca, Mexico by Dave Skinner." figureDoi="10.3897/phytokeys.222.87779.figure4" httpUri="https://binary.pensoft.net/fig/828343" pageId="0" pageNumber="75">4</figureCitation>
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) can be recognised by its leafless flowering shoots up to ca. 1 m long, combined with large, red and unappendaged bracts, tubular and yellow flowers, and stamen far exceeding the labellum like in
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<taxonomicName class="Liliopsida" family="Costaceae" kingdom="Plantae" lsidName="C. pulverulentus" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="pulverulentus">
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<emphasis italics="true" pageId="0" pageNumber="75">C. pulverulentus</emphasis>
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</taxonomicName>
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C.Presl.
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</paragraph>
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<caption doi="10.3897/phytokeys.222.87779.figure4" httpUri="https://binary.pensoft.net/fig/828343" pageId="0" pageNumber="75" start="Figure 4" startId="F4">
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<paragraph pageId="0" pageNumber="75">
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<emphasis bold="true" pageId="0" pageNumber="75">Figure 4.</emphasis>
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<taxonomicName authorityName="Maas & H. Maas" authorityYear="2023" class="Liliopsida" family="Costaceae" genus="Costus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Costus alticolus" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="alticolus">
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<emphasis italics="true" pageId="0" pageNumber="75">Costus alticolus</emphasis>
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</taxonomicName>
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Maas & H.Maas
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<emphasis bold="true" pageId="0" pageNumber="75">A</emphasis>
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inflorescence with flower, showing stamen (st) extending far beyond labellum, fully exposing the thecae
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<emphasis bold="true" pageId="0" pageNumber="75">B, C</emphasis>
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plant in habitat
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<emphasis bold="true" pageId="0" pageNumber="75">D</emphasis>
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flower and bract details
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<emphasis bold="true" pageId="0" pageNumber="75">E</emphasis>
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flowering shoot. Photos in habitat in Chinantla region near Santa Cruz Tepetotutla, Oaxaca, Mexico by Dave Skinner.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="75" type="materials_examined">
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<paragraph pageId="0" pageNumber="75">
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<typeStatus>Type</typeStatus>
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.
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</paragraph>
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<paragraph pageId="0" pageNumber="75">
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<materialsCitation collectingDate="1984-01-16" collectorName="Solheim, Reisfeld" country="Mexico" county="Sierra Mazateca" elevation="1200" location="Cuauhtemoc" municipality="Santa Maria Chilchotla" specimenCount="1" stateProvince="Oaxaca" typeStatus="holotype">
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<collectingCountry name="Mexico">Mexico</collectingCountry>
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.
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<collectingRegion country="Mexico" name="Oaxaca">Oaxaca</collectingRegion>
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:
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<collectingCounty>Sierra Mazateca</collectingCounty>
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, Mun.
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<collectingMunicipality>
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Santa
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<normalizedToken originalValue="María">Maria</normalizedToken>
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Chilchotla
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</collectingMunicipality>
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,
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<location LSID="urn:lsid:plazi:treatment:5B398AFCE2A95824B19C58E6A19DE6EB:38F274D80E06997D6510729D62AD6770" country="Mexico" county="Sierra Mazateca" municipality="Santa Maria Chilchotla" name="Cuauhtemoc" stateProvince="Oaxaca">Cuauhtemoc</location>
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, ca.
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<locationDeviation location="Santa Maria Chilchotla">
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<quantity metricMagnitude="3" metricUnit="m" metricValue="4.0" unit="km" value="4.0">4 km</quantity>
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NE of Santa
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<normalizedToken originalValue="María">Maria</normalizedToken>
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Chilchotla
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</locationDeviation>
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, near
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<location LSID="urn:lsid:plazi:treatment:5B398AFCE2A95824B19C58E6A19DE6EB:1A9B822FDBFC1FF8A8251D5BC662EFFD" country="Mexico" county="Sierra Mazateca" municipality="Santa Maria Chilchotla" name="Clemencia" stateProvince="Oaxaca">Clemencia</location>
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and
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<location LSID="urn:lsid:plazi:treatment:5B398AFCE2A95824B19C58E6A19DE6EB:9D0A13C78FEE8EAE20927E707565786C" country="Mexico" county="Sierra Mazateca" municipality="Santa Maria Chilchotla" name="Santa Rosa" stateProvince="Oaxaca">
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<location LSID="urn:lsid:plazi:treatment:5B398AFCE2A95824B19C58E6A19DE6EB:FC69E5493F71CEB6773582098144D474" country="Mexico" county="Sierra Mazateca" municipality="Santa Maria Chilchotla" name="Santa" stateProvince="Oaxaca">Santa</location>
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Rosa
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</location>
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,
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<quantity metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">
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<elevation metricMagnitude="3" metricUnit="m" metricValue="1.2" unit="m" value="1200.0">1200 m</elevation>
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</quantity>
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,
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<collectingDate value="1984-01-16">16 Jan 1984</collectingDate>
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,
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<emphasis italics="true" pageId="0" pageNumber="75">
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<collectorName>Solheim</collectorName>
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&
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<collectorName>Reisfeld</collectorName>
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1367
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</emphasis>
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(
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<typeStatus>holotype</typeStatus>
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WIS)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="75" type="description">
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<paragraph pageId="0" pageNumber="75">Description.</paragraph>
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<paragraph pageId="0" pageNumber="75">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="75">Herb</emphasis>
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2-3.5 m tall.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="75">Leaves</emphasis>
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sheaths 10-30 mm diam; ligule truncate, 4-12 mm long; petiole 5-20 mm long; sheaths, ligule and petiole densely to sparsely puberulous; lamina narrowly elliptic, 25-47
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<normalizedToken originalValue="×">x</normalizedToken>
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6-14 cm, adaxially glabrous, abaxially densely (particularly along the primary vein) sericeous to almost glabrous, base acute, apex acuminate (acumen 5-10 mm long).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="75">Inflorescence</emphasis>
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cylindric, 11-18
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<normalizedToken originalValue="×">x</normalizedToken>
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5-9 cm, terminating a leafless shoot 20-95 cm long, sheaths obliquely truncate, 4-7 cm long, rather densely puberulous to glabrous; bracts, bracteole, calyx, ovary, and capsule glabrous to densely puberulous.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="75">Flowers</emphasis>
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abaxially oriented; bracts red to bright crimson, coriaceous, ovate-oblong, 4-6
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<normalizedToken originalValue="×">x</normalizedToken>
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3-4 cm, apex obtuse, callus ca. 10 mm long; bracteole boat-shaped, 30-35 mm long; calyx red, 13-21 mm long, lobes deltate to shallowly triangular, 3-6 mm long; corolla yellow to orange, 50-60 mm long, glabrous, lobes narrowly elliptic, ca. 40 mm long; labellum yellow, lateral lobes rolled inwards and forming a slightly curved tube 18-20 mm diam, obovate when spread out, ca. 60 by 35 mm, irregularly lobulate, lobules recurved, ca. 4 mm long; stamen yellow, ca. 65
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<normalizedToken originalValue="×">x</normalizedToken>
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11 mm, far exceeding the labellum, apex acute, anther ca. 8 mm long.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="75">Capsule</emphasis>
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ellipsoid, 17-18 mm long.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="75" type="distribution">
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<paragraph pageId="0" pageNumber="75">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="75">
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Mexico (Oaxaca) (Fig.
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<figureCitation captionStart="Figure 21" captionStartId="F21" captionText="Figure 21. Distribution maps for new species of Costaceae: Chamaecostus - Costus douglasdalyi A Chamaecostus manausensis Maas & H. Maas B Costus alfredoi Maas & H. Maas C Costus alleniopsis Maas & D. Skinner D Costus alticolus Maas & H. Maas E Costus antioquiensis Maas & H. Maas F Costus callosus Maas & H. Maas G Costus cochabambae Maas & H. Maas H Costus convexus Maas & D. Skinner I Costus douglasdalyi Maas & H. Maas." figureDoi="10.3897/phytokeys.222.87779.figure21" httpUri="https://binary.pensoft.net/fig/828360" pageId="0" pageNumber="75">21D</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="75" type="habitat">
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<paragraph pageId="0" pageNumber="75">Habitat and ecology.</paragraph>
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<paragraph pageId="0" pageNumber="75">
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In primary, wet montane to premontane cloud forests, with some mosses on trunks, drier on ridge crests, with
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<taxonomicName class="Magnoliopsida" family="Chloranthaceae" genus="Hedyosmum" kingdom="Plantae" lsidName="Hedyosmum mexicanum" order="Chloranthales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="mexicanum">
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<emphasis italics="true" pageId="0" pageNumber="75">Hedyosmum mexicanum</emphasis>
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</taxonomicName>
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Cordem. Ex Baill.,
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<taxonomicName class="Magnoliopsida" family="Altingiaceae" genus="Liquidambar" kingdom="Plantae" lsidName="Liquidambar macrophylla" order="Saxifragales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="macrophylla">
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<emphasis italics="true" pageId="0" pageNumber="75">Liquidambar macrophylla</emphasis>
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</taxonomicName>
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Oerst.,
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<taxonomicName class="Magnoliopsida" family="Siparunaceae" genus="Siparuna" kingdom="Plantae" lsidName="Siparuna andina" order="Laurales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="andina">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Siparuna andina</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
A.DC.,
|
|||
|
|
<taxonomicName class="Magnoliopsida" family="Rubiaceae" genus="Posoqueria" kingdom="Plantae" lsidName="Posoqueria latifolia" order="Gentianales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="latifolia">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Posoqueria latifolia</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(Rudge) Roem. & Schult.,
|
|||
|
|
<taxonomicName class="Liliopsida" family="Heliconiaceae" genus="Heliconia" kingdom="Plantae" lsidName="Heliconia" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="genus">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Heliconia</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
sp.,
|
|||
|
|
<taxonomicName class="Magnoliopsida" family="Rubiaceae" genus="Deppea" kingdom="Plantae" lsidName="Deppea" order="Gentianales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="genus">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Deppea</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
sp,
|
|||
|
|
<taxonomicName class="Liliopsida" family="Arecaceae" genus="Chamaedorea" kingdom="Plantae" lsidName="Chamaedorea" order="Arecales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="genus">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Chamaedorea</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
sp.,
|
|||
|
|
<taxonomicName class="Magnoliopsida" family="Gesneriaceae" genus="Solenophora" kingdom="Plantae" lsidName="Solenophora" order="Lamiales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="genus">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Solenophora</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
sp., and
|
|||
|
|
<taxonomicName class="Magnoliopsida" family="Lamiaceae" genus="Salvia" kingdom="Plantae" lsidName="Salvia divinorum" order="Lamiales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="divinorum">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Salvia divinorum</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
Epling &
|
|||
|
|
<normalizedToken originalValue="Játiva">Jativa</normalizedToken>
|
|||
|
|
, at elevations of 1040-1900 m. Flowering year-round.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="0" pageNumber="75" type="etymology">
|
|||
|
|
<paragraph pageId="0" pageNumber="75">Etymology.</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="75">
|
|||
|
|
<taxonomicName authorityName="Maas & H. Maas" authorityYear="2023" class="Liliopsida" family="Costaceae" genus="Costus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Costus alticolus" order="Zingiberales" pageId="0" pageNumber="75" phylum="Tracheophyta" rank="species" species="alticolus">
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Costus alticolus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
sp. nov. occurs at quite high elevations up to 1900 m, hence the specific epithet alticolus (
|
|||
|
|
<normalizedToken originalValue="‘altus’">'altus'</normalizedToken>
|
|||
|
|
means
|
|||
|
|
<normalizedToken originalValue="‘high’">'high'</normalizedToken>
|
|||
|
|
in Latin;
|
|||
|
|
<normalizedToken originalValue="‘colere’">'colere'</normalizedToken>
|
|||
|
|
means 'to
|
|||
|
|
<normalizedToken originalValue="live’">live'</normalizedToken>
|
|||
|
|
in Latin).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="0" pageNumber="75" type="paratypes">
|
|||
|
|
<paragraph pageId="0" pageNumber="75">Paratypes.</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="75">
|
|||
|
|
<emphasis bold="true" pageId="0" pageNumber="75">Mexico. Oaxaca</emphasis>
|
|||
|
|
: Distr. and Mun.
|
|||
|
|
<normalizedToken originalValue="Ixtlán">Ixtlan</normalizedToken>
|
|||
|
|
, near abandoned community of Tarabundi, S side of
|
|||
|
|
<normalizedToken originalValue="Río">Rio</normalizedToken>
|
|||
|
|
<normalizedToken originalValue="Soyolapán">Soyolapan</normalizedToken>
|
|||
|
|
, access via La Luz, primary forest toward
|
|||
|
|
<normalizedToken originalValue="Río">Rio</normalizedToken>
|
|||
|
|
Langaro, 1230-1260 m, 16 Nov 1993,
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Boyle & Massart 2556</emphasis>
|
|||
|
|
(MO, U); Mun. San Felipe Usila, cuenca del
|
|||
|
|
<normalizedToken originalValue="Río">Rio</normalizedToken>
|
|||
|
|
Perfume, 4.4 km SE of Santa Cruz Tepetotutla, 1400 m, 29 Mar 1995,
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">
|
|||
|
|
<normalizedToken originalValue="Rincón">Rincon</normalizedToken>
|
|||
|
|
<normalizedToken originalValue="Guttiérrez">Guttierrez</normalizedToken>
|
|||
|
|
et al. 592
|
|||
|
|
</emphasis>
|
|||
|
|
(MO); Distr. Mixe, Mun. Totontepec, Totontepec, 1900 m, 9 Aug 1990,
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Rivera Reyes & Martin 1610</emphasis>
|
|||
|
|
(CAS), Hwy 175, km 169, near La Esperanza, 1075 m, 12 Aug 2017,
|
|||
|
|
<emphasis italics="true" pageId="0" pageNumber="75">Skinner R3400</emphasis>
|
|||
|
|
(BH).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection pageId="0" pageNumber="75" type="notes">
|
|||
|
|
<paragraph pageId="0" pageNumber="75">Notes.</paragraph>
|
|||
|
|
<paragraph pageId="0" pageNumber="75">
|
|||
|
|
We are greatly indebted to Manuel
|
|||
|
|
<normalizedToken originalValue="Gutiérrez">Gutierrez</normalizedToken>
|
|||
|
|
, who assisted Dave Skinner in obtaining permission from the indigenous people of Santa Cruz Tepetotutla, Oaxaca, Mexico, to study this new species common in the Chinantla region of Oaxaca.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|