<documentID-CLB-Dataset="21450"ID-DOI="http://dx.doi.org/10.3897/zookeys.950.51298"ID-GBIF-Dataset="21975627-96b7-45c7-9c80-0a73a850169a"ID-PMC="PMC7387366"ID-Pensoft-Pub="1313-2970-950-51"ID-Pensoft-UUID="7F08092C01585A70A7948EA57A2A8E2E"ID-PubMed="32774101"ID-ZooBank="3E805885D3354FB3AE8DFA443FAD82AE"ModsDocID="1313-2970-950-51"checkinTime="1595285263451"checkinUser="pensoft"docAuthor="Takaoka, Hiroyuki, Srisuka, Wichai, Fukuda, Masako & Saeung, Atiporn"docDate="2020"docId="8E6F2C6FF5875C75B991D960297CD504"docLanguage="en"docName="ZooKeys 950: 51-152"docOrigin="ZooKeys 950"docSource="http://dx.doi.org/10.3897/zookeys.950.51298"docTitle="Simulium (Gomphostilbia) maehongsonense Takaoka, Srisuka & Saeung 2020, sp. nov."docType="treatment"docUuid="BD4BF904-907D-460F-9276-168D189BC582"docUuidSource="ZooBank"docVersion="6"id="7F08092C01585A70A7948EA57A2A8E2E"lastPageNumber="51"masterDocId="7F08092C01585A70A7948EA57A2A8E2E"masterDocTitle="Twenty-one new species of the Simulium (Gomphostilbia) asakoae species group (Diptera, Simuliidae) in Thailand, with their genetic relationships"masterLastPageNumber="152"masterPageNumber="51"pageNumber="51"updateTime="1732680320779"updateUser="ExternalLinkService">
<mods:titleid="00EE0097DFA134DF239CB3939586A4E7">Twenty-one new species of the Simulium (Gomphostilbia) asakoae species group (Diptera, Simuliidae) in Thailand, with their genetic relationships</mods:title>
<mods:affiliationid="B2F899008929C06E3E5FE2C774EB649F">Tropical Infectious Diseases Research and Education Centre (TIDREC), University of Malaya, 50603, Kuala Lumpur, Malaysia</mods:affiliation>
<mods:affiliationid="C30F5F6D05D250D72CFCA9AE39E998B1">Entomology Section, Queen Sirikit Botanic Garden, P. O. Box 7, Maerim, Chiang Mai 50180, Thailand</mods:affiliation>
<mods:affiliationid="5747E11C866F3B744C19DCFB247468F2">Center of Insect Vector Study, Department of Parasitology, Faculty of Medicine, Chiang Mai University, Chiang Mai 50200, Thailand</mods:affiliation>
<figureCitationid="33EC1A7A0E4B48CF9CB71D08C50497CD"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. Female, male, pupa and larva of S. maehongsonense sp. nov. A female B-F male G-I pupa J larva. A mandible (right side) B hind basitarsus and second tarsomere (left side; lateral view) C coxites, styles and ventral plate (ventral view) D style (right side; ventrolateral view) E ventral plate and median sclerite (lateral view) F ventral plate (caudal view) G gill filaments (left side; lateral view) H terminal hooks (caudal view) I cocoon (dorsal view) J head capsule (ventral view). Scale bars: 1.0 mm (I); 0.1 mm (B, G, J); 0.02 mm (C-F); 0.01 mm (A, H)."figureDoi="10.3897/zookeys.950.51298.figure14"httpUri="https://binary.pensoft.net/fig/433878"pageId="0"pageNumber="51">Figs 14</figureCitation>
<figureCitationid="12564139E1869ACF2D3D58A18C0A3020"captionStart="Figure 25"captionStartId="F25"captionText="Figure 25. Schematic illustrations of larval body color patterns (dorsal view). A S. teerachanense sp. nov. B S. pitasawatae sp. nov. C S. sutheppuiense sp. nov. D S. asakoae E S. klonglanense sp. nov. F S. namdanense sp. nov. G S. chaowaense sp. nov. H S. loeiense sp. nov. I S. huaimorense sp. nov. J S. nanoiense sp. nov. K S. junkumae sp. nov. L S. myanmarense M S. maewongense sp. nov. N S. phapeungense sp. nov. O S. muangpanense sp. nov. P S. maehongsonense sp. nov. Q S. thungchangense sp. nov. R S. chiangdaoense S S. inthanonense (morph 1) T S. inthanonense (morph 2)."figureDoi="10.3897/zookeys.950.51298.figure25"httpUri="https://binary.pensoft.net/fig/433889"pageId="0"pageNumber="51">, 25P</figureCitation>
<taxonomicNameid="D71BF58A6EFCEDCE92DE3144EF56579F"authorityName="Takaoka, Srisuka & Saeung"authorityYear="2020"class="Insecta"family="Simuliidae"genus="Simulium"higherTaxonomySource="CoL"kingdom="Animalia"lsidName="Simulium maehongsonense"order="Diptera"pageId="0"pageNumber="51"phylum="Arthropoda"rank="species"species="maehongsonense">
male, QSBG col. no. 107, Thailand, 12-VII-2017, by W. Srisuka", collected from a stream (width 120 cm, depth 2 cm, bed sandy, fast flow, pH 7.6, 21.2 °C, partially shade, elevation 1,034 m,
: Two females (thorax of one female for DNA analysis), seven males (thorax of one male for DNA analysis) (with their associated pupal exuviae and cocoons), and six mature larvae (one mature larva for DNA analysis) (in 80% ethanol), same data as for holotype.
<figureCitationid="7F36B9D690B8C914F17A76C0A185F9FA"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. Female, male, pupa and larva of S. maehongsonense sp. nov. A female B-F male G-I pupa J larva. A mandible (right side) B hind basitarsus and second tarsomere (left side; lateral view) C coxites, styles and ventral plate (ventral view) D style (right side; ventrolateral view) E ventral plate and median sclerite (lateral view) F ventral plate (caudal view) G gill filaments (left side; lateral view) H terminal hooks (caudal view) I cocoon (dorsal view) J head capsule (ventral view). Scale bars: 1.0 mm (I); 0.1 mm (B, G, J); 0.02 mm (C-F); 0.01 mm (A, H)."figureDoi="10.3897/zookeys.950.51298.figure14"httpUri="https://binary.pensoft.net/fig/433878"pageId="0"pageNumber="51">14A</figureCitation>
). Male: number of upper-eye facets in 13 vertical columns and 14 horizontal rows. Pupa: relatively shorter gill filaments (longest 2.1-2.3 mm long) (Fig.
<figureCitationid="2237ABD13B3051C2D4C3D6D3C16C22AC"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. Female, male, pupa and larva of S. maehongsonense sp. nov. A female B-F male G-I pupa J larva. A mandible (right side) B hind basitarsus and second tarsomere (left side; lateral view) C coxites, styles and ventral plate (ventral view) D style (right side; ventrolateral view) E ventral plate and median sclerite (lateral view) F ventral plate (caudal view) G gill filaments (left side; lateral view) H terminal hooks (caudal view) I cocoon (dorsal view) J head capsule (ventral view). Scale bars: 1.0 mm (I); 0.1 mm (B, G, J); 0.02 mm (C-F); 0.01 mm (A, H)."figureDoi="10.3897/zookeys.950.51298.figure14"httpUri="https://binary.pensoft.net/fig/433878"pageId="0"pageNumber="51">14G</figureCitation>
<figureCitationid="DA8F9D5EC71216C570125CBF0697DE08"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. Female, male, pupa and larva of S. maehongsonense sp. nov. A female B-F male G-I pupa J larva. A mandible (right side) B hind basitarsus and second tarsomere (left side; lateral view) C coxites, styles and ventral plate (ventral view) D style (right side; ventrolateral view) E ventral plate and median sclerite (lateral view) F ventral plate (caudal view) G gill filaments (left side; lateral view) H terminal hooks (caudal view) I cocoon (dorsal view) J head capsule (ventral view). Scale bars: 1.0 mm (I); 0.1 mm (B, G, J); 0.02 mm (C-F); 0.01 mm (A, H)."figureDoi="10.3897/zookeys.950.51298.figure14"httpUri="https://binary.pensoft.net/fig/433878"pageId="0"pageNumber="51">14H</figureCitation>
<figureCitationid="6AD5E04758D6B094D3930B6136B01648"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. Female, male, pupa and larva of S. maehongsonense sp. nov. A female B-F male G-I pupa J larva. A mandible (right side) B hind basitarsus and second tarsomere (left side; lateral view) C coxites, styles and ventral plate (ventral view) D style (right side; ventrolateral view) E ventral plate and median sclerite (lateral view) F ventral plate (caudal view) G gill filaments (left side; lateral view) H terminal hooks (caudal view) I cocoon (dorsal view) J head capsule (ventral view). Scale bars: 1.0 mm (I); 0.1 mm (B, G, J); 0.02 mm (C-F); 0.01 mm (A, H)."figureDoi="10.3897/zookeys.950.51298.figure14"httpUri="https://binary.pensoft.net/fig/433878"pageId="0"pageNumber="51">14J</figureCitation>
<figureCitationid="DAF9266CC1F16806FFCE99AE7ABBDED7"captionStart="Figure 25"captionStartId="F25"captionText="Figure 25. Schematic illustrations of larval body color patterns (dorsal view). A S. teerachanense sp. nov. B S. pitasawatae sp. nov. C S. sutheppuiense sp. nov. D S. asakoae E S. klonglanense sp. nov. F S. namdanense sp. nov. G S. chaowaense sp. nov. H S. loeiense sp. nov. I S. huaimorense sp. nov. J S. nanoiense sp. nov. K S. junkumae sp. nov. L S. myanmarense M S. maewongense sp. nov. N S. phapeungense sp. nov. O S. muangpanense sp. nov. P S. maehongsonense sp. nov. Q S. thungchangense sp. nov. R S. chiangdaoense S S. inthanonense (morph 1) T S. inthanonense (morph 2)."figureDoi="10.3897/zookeys.950.51298.figure25"httpUri="https://binary.pensoft.net/fig/433889"pageId="0"pageNumber="51">25P</figureCitation>
Frontal ratio 1.9:1.0:2.1; frons:head ratio 1.0:4.2. Labrum 0.52 times length of clypeus. Maxillary palpus: proportional length of third, fourth and fifth palpal segments 1:1.1:2.2; sensory vesicle medium-long (0.26 times length of third palpal segment. Lacinia with 9-11 inner and 11-13 outer teeth. Mandible (Fig.
<figureCitationid="B71499FE66D48A3CB528BE9A68A59499"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. Female, male, pupa and larva of S. maehongsonense sp. nov. A female B-F male G-I pupa J larva. A mandible (right side) B hind basitarsus and second tarsomere (left side; lateral view) C coxites, styles and ventral plate (ventral view) D style (right side; ventrolateral view) E ventral plate and median sclerite (lateral view) F ventral plate (caudal view) G gill filaments (left side; lateral view) H terminal hooks (caudal view) I cocoon (dorsal view) J head capsule (ventral view). Scale bars: 1.0 mm (I); 0.1 mm (B, G, J); 0.02 mm (C-F); 0.01 mm (A, H)."figureDoi="10.3897/zookeys.950.51298.figure14"httpUri="https://binary.pensoft.net/fig/433878"pageId="0"pageNumber="51">14A</figureCitation>
Foreleg: tibia yellowish white except apical three-tenths brownish black; basitarsus moderately dilated, 5.5 times as long as its greatest width. Midleg: tarsus dark brown though basal half of basitarsus yellow (its border not well defined). Hind leg: coxa light brown; tibia yellowish white on basal two-thirds and light brown to brownish black on rest; basitarsus 6.7 times as long as wide, and 0.6 and 0.5 times as wide as greatest widths of tibia and femur, respectively; calcipala 0.5 times as wide as greatest width of basitarsus; claw with large basal tooth 0.46 times length of claw.
Sternite 8 bare medially, with 20-22 medium-long to long hairs together with two or three slender short hairs on each side. Ovipositor valves each moderately covered with microsetae interspersed with two or three short hairs. Paraproct in lateral view 0.6 times as long as wide, with 18 medium-long to long hairs on ventral and lateral surfaces. Spermatheca ellipsoid, 1.3 times as long as its greatest width.
Slightly wider than thorax. Upper eye dark brown, consisting of large facets in 13 vertical columns and 14 horizontal rows on each side. Maxillary palpus: proportional lengths of third, fourth, and fifth palpal segments 1.0:1.2:2.8; sensory vesicle 0.17 times length of third palpal segment.
Foreleg: tibia yellowish white except l apical one-third dark brown, and inner and lateral surfaces of subbasal portion light brown; basitarsus moderately dilated, 6.5-7.3 times as long as its greatest width. Hind leg: coxa light brown; tibia dark brown to brownish black except little less than basal half yellowish white; tarsus dark brown except little less than basal half of basitarsus and basal one-third of second tarsomere yellowish; basitarsus (Fig.
<figureCitationid="08783EADEF720FC40ED5D45C7C6EEA16"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. Female, male, pupa and larva of S. maehongsonense sp. nov. A female B-F male G-I pupa J larva. A mandible (right side) B hind basitarsus and second tarsomere (left side; lateral view) C coxites, styles and ventral plate (ventral view) D style (right side; ventrolateral view) E ventral plate and median sclerite (lateral view) F ventral plate (caudal view) G gill filaments (left side; lateral view) H terminal hooks (caudal view) I cocoon (dorsal view) J head capsule (ventral view). Scale bars: 1.0 mm (I); 0.1 mm (B, G, J); 0.02 mm (C-F); 0.01 mm (A, H)."figureDoi="10.3897/zookeys.950.51298.figure14"httpUri="https://binary.pensoft.net/fig/433878"pageId="0"pageNumber="51">14B</figureCitation>
<figureCitationid="9B01C8781C599AA6BE3CE4B8F47AF9A4"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. Female, male, pupa and larva of S. maehongsonense sp. nov. A female B-F male G-I pupa J larva. A mandible (right side) B hind basitarsus and second tarsomere (left side; lateral view) C coxites, styles and ventral plate (ventral view) D style (right side; ventrolateral view) E ventral plate and median sclerite (lateral view) F ventral plate (caudal view) G gill filaments (left side; lateral view) H terminal hooks (caudal view) I cocoon (dorsal view) J head capsule (ventral view). Scale bars: 1.0 mm (I); 0.1 mm (B, G, J); 0.02 mm (C-F); 0.01 mm (A, H)."figureDoi="10.3897/zookeys.950.51298.figure14"httpUri="https://binary.pensoft.net/fig/433878"pageId="0"pageNumber="51">14B</figureCitation>
<figureCitationid="06B163ADD12EC35C080D19FC0211A99E"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. Female, male, pupa and larva of S. maehongsonense sp. nov. A female B-F male G-I pupa J larva. A mandible (right side) B hind basitarsus and second tarsomere (left side; lateral view) C coxites, styles and ventral plate (ventral view) D style (right side; ventrolateral view) E ventral plate and median sclerite (lateral view) F ventral plate (caudal view) G gill filaments (left side; lateral view) H terminal hooks (caudal view) I cocoon (dorsal view) J head capsule (ventral view). Scale bars: 1.0 mm (I); 0.1 mm (B, G, J); 0.02 mm (C-F); 0.01 mm (A, H)."figureDoi="10.3897/zookeys.950.51298.figure14"httpUri="https://binary.pensoft.net/fig/433878"pageId="0"pageNumber="51">14C</figureCitation>
<figureCitationid="A9ECCBB0188F88BF8450F63BF69C029A"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. Female, male, pupa and larva of S. maehongsonense sp. nov. A female B-F male G-I pupa J larva. A mandible (right side) B hind basitarsus and second tarsomere (left side; lateral view) C coxites, styles and ventral plate (ventral view) D style (right side; ventrolateral view) E ventral plate and median sclerite (lateral view) F ventral plate (caudal view) G gill filaments (left side; lateral view) H terminal hooks (caudal view) I cocoon (dorsal view) J head capsule (ventral view). Scale bars: 1.0 mm (I); 0.1 mm (B, G, J); 0.02 mm (C-F); 0.01 mm (A, H)."figureDoi="10.3897/zookeys.950.51298.figure14"httpUri="https://binary.pensoft.net/fig/433878"pageId="0"pageNumber="51">14C</figureCitation>
<figureCitationid="CF85DD55751DF54F7CE898C2FFCEC78B"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. Female, male, pupa and larva of S. maehongsonense sp. nov. A female B-F male G-I pupa J larva. A mandible (right side) B hind basitarsus and second tarsomere (left side; lateral view) C coxites, styles and ventral plate (ventral view) D style (right side; ventrolateral view) E ventral plate and median sclerite (lateral view) F ventral plate (caudal view) G gill filaments (left side; lateral view) H terminal hooks (caudal view) I cocoon (dorsal view) J head capsule (ventral view). Scale bars: 1.0 mm (I); 0.1 mm (B, G, J); 0.02 mm (C-F); 0.01 mm (A, H)."figureDoi="10.3897/zookeys.950.51298.figure14"httpUri="https://binary.pensoft.net/fig/433878"pageId="0"pageNumber="51">14D</figureCitation>
<figureCitationid="B2BF34F4FEF0BED01ABF63321F86A06B"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. Female, male, pupa and larva of S. maehongsonense sp. nov. A female B-F male G-I pupa J larva. A mandible (right side) B hind basitarsus and second tarsomere (left side; lateral view) C coxites, styles and ventral plate (ventral view) D style (right side; ventrolateral view) E ventral plate and median sclerite (lateral view) F ventral plate (caudal view) G gill filaments (left side; lateral view) H terminal hooks (caudal view) I cocoon (dorsal view) J head capsule (ventral view). Scale bars: 1.0 mm (I); 0.1 mm (B, G, J); 0.02 mm (C-F); 0.01 mm (A, H)."figureDoi="10.3897/zookeys.950.51298.figure14"httpUri="https://binary.pensoft.net/fig/433878"pageId="0"pageNumber="51">14C</figureCitation>
<figureCitationid="182B9888B5761721AE550A55605B2F71"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. Female, male, pupa and larva of S. maehongsonense sp. nov. A female B-F male G-I pupa J larva. A mandible (right side) B hind basitarsus and second tarsomere (left side; lateral view) C coxites, styles and ventral plate (ventral view) D style (right side; ventrolateral view) E ventral plate and median sclerite (lateral view) F ventral plate (caudal view) G gill filaments (left side; lateral view) H terminal hooks (caudal view) I cocoon (dorsal view) J head capsule (ventral view). Scale bars: 1.0 mm (I); 0.1 mm (B, G, J); 0.02 mm (C-F); 0.01 mm (A, H)."figureDoi="10.3897/zookeys.950.51298.figure14"httpUri="https://binary.pensoft.net/fig/433878"pageId="0"pageNumber="51">14F</figureCitation>
<figureCitationid="93A488083152079A31D3D83A901F6C8D"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. Female, male, pupa and larva of S. maehongsonense sp. nov. A female B-F male G-I pupa J larva. A mandible (right side) B hind basitarsus and second tarsomere (left side; lateral view) C coxites, styles and ventral plate (ventral view) D style (right side; ventrolateral view) E ventral plate and median sclerite (lateral view) F ventral plate (caudal view) G gill filaments (left side; lateral view) H terminal hooks (caudal view) I cocoon (dorsal view) J head capsule (ventral view). Scale bars: 1.0 mm (I); 0.1 mm (B, G, J); 0.02 mm (C-F); 0.01 mm (A, H)."figureDoi="10.3897/zookeys.950.51298.figure14"httpUri="https://binary.pensoft.net/fig/433878"pageId="0"pageNumber="51">14G</figureCitation>
) composed of eight slender thread-like filaments, arranged as (3+3) +2 or [3+(1+2)]+2 from dorsal to ventral; common basal stalk 0.5-0.7 times length of interspiracular trunk; dorsal triplet with extremely short stalk; stalk of ventral pair of filaments 0.9-1.5 times length of common basal stalk, and 0.5-0.8 times length of interspiracular trunk; primary stalk of dorsal triplet lying against that of lower pair at angle of 70° when viewed laterally; filaments of dorsal triplet subequal in length (1.1-1.3 mm) and thickness to one another; filaments of middle triplet subequal in length (1.5-1.7 mm) and thickness to one another; two filaments of ventral pair subequal in length (2.1-2.3 mm) and thickness to each other and 1.6 times as thick as six other filaments of dorsal and middle triplets when compared basally.
<figureCitationid="6C9A685229764AEED0DF286093CDA6B0"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. Female, male, pupa and larva of S. maehongsonense sp. nov. A female B-F male G-I pupa J larva. A mandible (right side) B hind basitarsus and second tarsomere (left side; lateral view) C coxites, styles and ventral plate (ventral view) D style (right side; ventrolateral view) E ventral plate and median sclerite (lateral view) F ventral plate (caudal view) G gill filaments (left side; lateral view) H terminal hooks (caudal view) I cocoon (dorsal view) J head capsule (ventral view). Scale bars: 1.0 mm (I); 0.1 mm (B, G, J); 0.02 mm (C-F); 0.01 mm (A, H)."figureDoi="10.3897/zookeys.950.51298.figure14"httpUri="https://binary.pensoft.net/fig/433878"pageId="0"pageNumber="51">14H</figureCitation>
<figureCitationid="9F067B797449FC1465F71DCF94D92116"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. Female, male, pupa and larva of S. maehongsonense sp. nov. A female B-F male G-I pupa J larva. A mandible (right side) B hind basitarsus and second tarsomere (left side; lateral view) C coxites, styles and ventral plate (ventral view) D style (right side; ventrolateral view) E ventral plate and median sclerite (lateral view) F ventral plate (caudal view) G gill filaments (left side; lateral view) H terminal hooks (caudal view) I cocoon (dorsal view) J head capsule (ventral view). Scale bars: 1.0 mm (I); 0.1 mm (B, G, J); 0.02 mm (C-F); 0.01 mm (A, H)."figureDoi="10.3897/zookeys.950.51298.figure14"httpUri="https://binary.pensoft.net/fig/433878"pageId="0"pageNumber="51">14I</figureCitation>
). Pale yellow, slipper-shaped, roughly to moderately woven, moderately extended ventrolaterally; anterior margin thickly woven medially, without bulge or short projection; 3.0-3.5 mm long by 2.0-2.9 mm wide.
= 5). Body length 4.5-5.5 mm. Body light ochreous with following color markings: thoracic segment 1 encircled with distinct greyish band (though disconnected ventromedially), thoracic segment 2 greyish and thoracic segment 3 ochreous on ventral surface; abdominal segment 1 light grey, abdominal segments 5 and 6 each with distinct reddish brown spot of various size and shape dorsomedially, and dorsal and dorsolateral surface of abdominal segments 4-8 faintly to moderately covered with pinkish to reddish brown pigments (Fig.
<figureCitationid="26C416FC767D970D7BA90BBB02204192"captionStart="Figure 25"captionStartId="F25"captionText="Figure 25. Schematic illustrations of larval body color patterns (dorsal view). A S. teerachanense sp. nov. B S. pitasawatae sp. nov. C S. sutheppuiense sp. nov. D S. asakoae E S. klonglanense sp. nov. F S. namdanense sp. nov. G S. chaowaense sp. nov. H S. loeiense sp. nov. I S. huaimorense sp. nov. J S. nanoiense sp. nov. K S. junkumae sp. nov. L S. myanmarense M S. maewongense sp. nov. N S. phapeungense sp. nov. O S. muangpanense sp. nov. P S. maehongsonense sp. nov. Q S. thungchangense sp. nov. R S. chiangdaoense S S. inthanonense (morph 1) T S. inthanonense (morph 2)."figureDoi="10.3897/zookeys.950.51298.figure25"httpUri="https://binary.pensoft.net/fig/433889"pageId="0"pageNumber="51">25P</figureCitation>
Head spots moderately positive. Antenna: proportional lengths of first, second, and third articles 1.0:0.8:0.7-0.9. Labral fan with 38-40 primary rays. Hypostoma with four to six hypostomal bristles per side lying nearly parallel to lateral margin. Postgenal cleft (Fig.
<figureCitationid="4D1B6485070DC90A04D4EAEF8FB0956A"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. Female, male, pupa and larva of S. maehongsonense sp. nov. A female B-F male G-I pupa J larva. A mandible (right side) B hind basitarsus and second tarsomere (left side; lateral view) C coxites, styles and ventral plate (ventral view) D style (right side; ventrolateral view) E ventral plate and median sclerite (lateral view) F ventral plate (caudal view) G gill filaments (left side; lateral view) H terminal hooks (caudal view) I cocoon (dorsal view) J head capsule (ventral view). Scale bars: 1.0 mm (I); 0.1 mm (B, G, J); 0.02 mm (C-F); 0.01 mm (A, H)."figureDoi="10.3897/zookeys.950.51298.figure14"httpUri="https://binary.pensoft.net/fig/433878"pageId="0"pageNumber="51">14J</figureCitation>
) by having the same number of upper-eye large facets, but is distinguished by the fore basitarsus 6.5-7.3 times as long as its greatest width (8.3 times in
<figureCitationid="0B1BE884FB00A41387BAA0D94C19452D"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. Female, male, pupa and larva of S. maehongsonense sp. nov. A female B-F male G-I pupa J larva. A mandible (right side) B hind basitarsus and second tarsomere (left side; lateral view) C coxites, styles and ventral plate (ventral view) D style (right side; ventrolateral view) E ventral plate and median sclerite (lateral view) F ventral plate (caudal view) G gill filaments (left side; lateral view) H terminal hooks (caudal view) I cocoon (dorsal view) J head capsule (ventral view). Scale bars: 1.0 mm (I); 0.1 mm (B, G, J); 0.02 mm (C-F); 0.01 mm (A, H)."figureDoi="10.3897/zookeys.950.51298.figure14"httpUri="https://binary.pensoft.net/fig/433878"pageId="0"pageNumber="51">14F</figureCitation>
<figureCitationid="F414B37AC57850E280409CBFE8773D97"captionStart="Figure 14"captionStartId="F14"captionText="Figure 14. Female, male, pupa and larva of S. maehongsonense sp. nov. A female B-F male G-I pupa J larva. A mandible (right side) B hind basitarsus and second tarsomere (left side; lateral view) C coxites, styles and ventral plate (ventral view) D style (right side; ventrolateral view) E ventral plate and median sclerite (lateral view) F ventral plate (caudal view) G gill filaments (left side; lateral view) H terminal hooks (caudal view) I cocoon (dorsal view) J head capsule (ventral view). Scale bars: 1.0 mm (I); 0.1 mm (B, G, J); 0.02 mm (C-F); 0.01 mm (A, H)."figureDoi="10.3897/zookeys.950.51298.figure14"httpUri="https://binary.pensoft.net/fig/433878"pageId="0"pageNumber="51">14I</figureCitation>
