337 lines
55 KiB
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337 lines
55 KiB
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<document id="2AB357141883D6D77981D3536CDE5625" ID-CLB-Dataset="27734" ID-DOI="10.11646/zootaxa.4532.4.4" ID-GBIF-Dataset="4809e341-99ce-476f-a3a8-5229de62f5eb" ID-ISSN="1175-5326" ID-Zenodo-Dep="2615629" ID-ZooBank="A35A92D5-2628-4203-8C94-DDBFFFCDC79E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553788686464" checkinUser="plazi" docAuthor="Anisyutkin, Leonid N. & Telnov, Dmitry" docDate="2018" docId="03F887BAFFA8FFD8FF37F9DA1B84FA21" docLanguage="en" docName="zootaxa.4532.4.4.pdf" docOrigin="Zootaxa 4532 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Neostylopyga yemenica Anisyutkin & Telnov 2018, comb. nov." docType="treatment" docVersion="8" lastPageNumber="536" masterDocId="FFC1FFC2FFA2FFD5FFA0FFB4184CFE79" masterDocTitle="New data on the cockroaches previously attributed to the genus Duchailluia Rehn, 1933 (Dictyoptera: Blattidae: Blattinae)" masterLastPageNumber="538" masterPageNumber="523" pageNumber="533" updateTime="1698715127610" updateUser="ExternalLinkService">
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<mods:mods id="E945FEF6E4768B015DA01CBBA92C678D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="2009C07F2E6EA3B9D97E5301F08924B9">
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<mods:title id="1E0B561AA1A8D1CED1F6CED99F0E213A">New data on the cockroaches previously attributed to the genus Duchailluia Rehn, 1933 (Dictyoptera: Blattidae: Blattinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="3E019BBFA831952A95FEF10DD359F476" type="personal">
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<mods:roleTerm id="E5F731A72949B79C8EADBEADA443105E">Author</mods:roleTerm>
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<mods:namePart id="B08B950F1C5A01D04B360948A188B048">Anisyutkin, Leonid N.</mods:namePart>
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</mods:role>
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<mods:namePart id="C4E5601CEAC96E3383C483E216244C9C">Telnov, Dmitry</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="AD5B109465C80B8CCA4C0A8EC17CACC9">text</mods:typeOfResource>
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<mods:titleInfo id="C8408CE2F17999163E564834ECFFE170">
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<mods:title id="475CC073D792DA06F2050A26A78092CA">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="A1D939771A7D4FF1F168B1A30906AED0">
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<mods:date id="895CAD7ECEBA009C14FE191CBD82F278">2018</mods:date>
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<mods:detail id="55EA590266C66FF0142AF94CF9F2C937" type="pubDate">
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<mods:number id="0253B1955681ED62D4794F0E9B9B62E8">2018-12-20</mods:number>
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</mods:detail>
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<mods:detail id="A13C2F91ACB141D204182CA81B3CFBE3" type="volume">
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<mods:number id="6BC76EDF46A32B5F050A4E6CFDCAE9D3">4532</mods:number>
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<mods:detail id="FDEB78BF072590D195DF408F45A0001B" type="issue">
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<mods:number id="EFEDA2B0A74C1A9FB406E2912832A6E2">4</mods:number>
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<mods:classification id="A8C15104DD4D0486D33E1026FC16A9CE">journal article</mods:classification>
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<mods:identifier id="1313E4BBA77C55439BF9D950FC595C07" type="CLB-Dataset">27734</mods:identifier>
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<mods:identifier id="26B79342258E3C663FB8D4B30C940329" type="DOI">10.11646/zootaxa.4532.4.4</mods:identifier>
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</mods:mods>
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<treatment id="03F887BAFFA8FFD8FF37F9DA1B84FA21" ID-DOI="http://doi.org/10.5281/zenodo.5976588" ID-GBIF-Taxon="154521469" ID-Zenodo-Dep="5976588" LSID="urn:lsid:plazi:treatment:03F887BAFFA8FFD8FF37F9DA1B84FA21" httpUri="http://treatment.plazi.org/id/03F887BAFFA8FFD8FF37F9DA1B84FA21" lastPageId="13" lastPageNumber="536" pageId="10" pageNumber="533">
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<subSubSection id="C34B6527FFA8FFDFFF37F9DA1B7FF8F1" box="[151,819,1646,1672]" pageId="10" pageNumber="533" type="nomenclature">
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<paragraph id="8BEE36ACFFA8FFDFFF37F9DA1B7FF8F1" blockId="10.[151,819,1646,1706]" box="[151,819,1646,1672]" pageId="10" pageNumber="533">
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<heading id="D0A681C0FFA8FFDFFF37F9DA1B7FF8F1" bold="true" box="[151,819,1646,1672]" fontSize="11" level="1" pageId="10" pageNumber="533" reason="1">
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<emphasis id="B925EABEFFA8FFDFFF37F9DA1B7FF8F1" bold="true" box="[151,819,1646,1672]" pageId="10" pageNumber="533">
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<taxonomicName id="4C514D2FFFA8FFDFFF37F9DA1AEAF8F1" ID-CoL="8N9LS" authority="(Bey-Bienko, 1969)" authorityName="Anisyutkin & Telnov" authorityYear="2018" baseAuthorityName="Bey-Bienko" baseAuthorityYear="1969" box="[151,678,1646,1672]" class="Insecta" family="Blattidae" genus="Neostylopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="10" pageNumber="533" phylum="Arthropoda" rank="species" species="yemenica" status="comb. nov.">
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<emphasis id="B925EABEFFA8FFDFFF37F9DA19FDF8F1" bold="true" box="[151,433,1646,1672]" italics="true" pageId="10" pageNumber="533">Neostylopyga yemenica</emphasis>
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(
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<bibRefCitation id="EFC04B5DFFA8FFDFFE60F9DA1AD1F8F1" author="Bey-Bienko, G. Ya." box="[448,669,1646,1672]" pageId="10" pageNumber="533" pagination="831 - 862" refId="ref7112" refString="Bey-Bienko, G. Ya. (1969) New genera and species of cockroaches (Blattoptera) from tropical and subtropical Asia. Entomologicheskoe Obozrenie, 48 (4), 831 - 862. [in Russian, English abstract]" type="journal article" year="1969">Bey-Bienko, 1969</bibRefCitation>
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)
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</taxonomicName>
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<taxonomicNameLabel id="A21657C5FFA8FFDFFD0DF9DA1B7FF8F1" box="[685,819,1646,1672]" pageId="10" pageNumber="533" rank="species">comb. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34B6527FFA8FFDFFF37F92519F1F8D3" box="[151,445,1681,1706]" pageId="10" pageNumber="533" type="description">
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<paragraph id="8BEE36ACFFA8FFDFFF37F92519F1F8D3" blockId="10.[151,819,1646,1706]" box="[151,445,1681,1706]" pageId="10" pageNumber="533">
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(
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<figureCitation id="136A2A29FFA8FFDFFF3FF92518A6F8D3" box="[159,234,1681,1706]" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. A–D—Duchailluia ivindo sp. nov.; E–H—Distylopyga beccalonii sp. nov.; I–L—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = A–D; b = E–H; c = I– L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="10" pageNumber="533">Figs. 3</figureCitation>
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I–L, 6A–E, 7A–E)
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</paragraph>
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</subSubSection>
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<subSubSection id="C34B6527FFA8FFDFFF37F96D1C11F88B" box="[151,1117,1753,1778]" pageId="10" pageNumber="533" type="materials_examined">
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<paragraph id="8BEE36ACFFA8FFDFFF37F96D1C11F88B" blockId="10.[151,1436,1753,2030]" box="[151,1117,1753,1778]" pageId="10" pageNumber="533">
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<emphasis id="B925EABEFFA8FFDFFF37F96D18B3F88B" bold="true" box="[151,255,1753,1778]" pageId="10" pageNumber="533">Material</emphasis>
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:
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<materialsCitation id="3B393CF1FFA8FFDFFEAEF96D1C11F88B" ID-GBIF-Occurrence="2120084144" box="[270,1117,1753,1778]" collectionCode="ZIN" collectorName="K. Suvorova" country="Yemen" location="Sana'a" pageId="10" pageNumber="533" specimenCount="1" stateProvince="San'a'" typeStatus="holotype">
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<typeStatus id="54EA880EFFA8FFDFFEAEF96D1936F88B" box="[270,378,1753,1778]" pageId="10" pageNumber="533" type="holotype">Holotype</typeStatus>
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. male,
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<collectingCountry id="F346763CFFA8FFDFFE68F96E1A60F888" box="[456,556,1754,1777]" name="Yemen" pageId="10" pageNumber="533">YEMEN</collectingCountry>
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,
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<collectingRegion id="4995F84EFFA8FFDFFD97F96E1A33F88B" box="[567,639,1754,1778]" country="Yemen" name="San'a'" pageId="10" pageNumber="533">Sana'a</collectingRegion>
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city, spring 1960, leg.
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<collectorName id="26A4537AFFA8FFDFFC20F96E1C41F88B" box="[896,1037,1754,1778]" pageId="10" pageNumber="533">K. Suvorova</collectorName>
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(
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<collectionCode id="ED40AE69FFA8FFDFFBBCF96E1C1CF88B" box="[1052,1104,1754,1778]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="10" pageNumber="533" type="Museum">ZIN</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34B6527FFA8FFD8FF67F9491B84FA21" lastPageId="13" lastPageNumber="536" pageId="10" pageNumber="533" type="description">
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<paragraph id="8BEE36ACFFA8FFDFFF67F9491910F943" blockId="10.[151,1436,1753,2030]" pageId="10" pageNumber="533">
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<emphasis id="B925EABEFFA8FFDFFF67F949199CF96F" bold="true" box="[199,464,1789,1814]" pageId="10" pageNumber="533">Redescription of male</emphasis>
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(
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<typeStatus id="54EA880EFFA8FFDFFE40F9491A04F96F" box="[480,584,1789,1814]" pageId="10" pageNumber="533" type="holotype">holotype</typeStatus>
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). The original description (
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<bibRefCitation id="EFC04B5DFFA8FFDFFCDDF9491C05F96F" author="Bey-Bienko, G. Ya." box="[893,1097,1789,1814]" pageId="10" pageNumber="533" pagination="831 - 862" refId="ref7112" refString="Bey-Bienko, G. Ya. (1969) New genera and species of cockroaches (Blattoptera) from tropical and subtropical Asia. Entomologicheskoe Obozrenie, 48 (4), 831 - 862. [in Russian, English abstract]" type="journal article" year="1969">Bey-Bienko 1969</bibRefCitation>
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) to be supplemented with the following details:
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</paragraph>
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<paragraph id="8BEE36ACFFA8FFDEFF67F8F11CB0FC61" blockId="10.[151,1436,1753,2030]" lastBlockId="11.[151,1437,151,536]" lastPageId="11" lastPageNumber="534" pageId="10" pageNumber="533">
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General color light reddish brown, ocellar spots and anteclypeus yellow. Head wide and rounded, slightly longer than wide (
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<figureCitation id="136A2A29FFA8FFDFFEC1F8DD19F7F9FB" box="[353,443,1897,1922]" captionStart="FIGURE 6" captionStartId="11.[151,249,1908,1930]" captionTargetBox="[190,1396,592,1885]" captionTargetId="figure@11.[172,1416,581,1898]" captionTargetPageId="11" captionText="FIGURE 6. Neostylopyga yemenica (Bey-Bienko, 1969), male, holotype. A—facial part of head; B—head, thorax and 1–6th abdominal tergites from above; C—7th abdominal tergite from above; D—abdominal apex from above, cerci broken off; Ehypandrium from below, styli broken off. Dotted areas shows membranous parts. Abbreviations: I–VIII, X—1st–10th abdominal tergites respectively—see text. Scale bar 1 mm: a = A; b = B; c = C, D; d = E." httpUri="https://zenodo.org/record/2615643/files/figure.png" pageId="10" pageNumber="533">Fig. 6A</figureCitation>
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); epicranial sutures absent; eyes small; ocellar spots small, but distinct; distance between eyes 1.2 times eye length; distance between antennal sockets 2.1 times of the scape length (about
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<quantity id="4CA99B49FFA8FFDFFA90F8391DDCF9DF" box="[1328,1424,1933,1958]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="533" unit="mm" value="0.5">0.5 mm</quantity>
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); approximate length ratio of 3rd–5th maxillary palpomeres 1.1: 1.0: 1.2. Pronotum campaniform, with caudal margin slightly angulate (
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<figureCitation id="136A2A29FFA8FFDFFE1AF8611A58F997" box="[442,532,2005,2030]" captionStart="FIGURE 6" captionStartId="11.[151,249,1908,1930]" captionTargetBox="[190,1396,592,1885]" captionTargetId="figure@11.[172,1416,581,1898]" captionTargetPageId="11" captionText="FIGURE 6. Neostylopyga yemenica (Bey-Bienko, 1969), male, holotype. A—facial part of head; B—head, thorax and 1–6th abdominal tergites from above; C—7th abdominal tergite from above; D—abdominal apex from above, cerci broken off; Ehypandrium from below, styli broken off. Dotted areas shows membranous parts. Abbreviations: I–VIII, X—1st–10th abdominal tergites respectively—see text. Scale bar 1 mm: a = A; b = B; c = C, D; d = E." httpUri="https://zenodo.org/record/2615643/files/figure.png" pageId="10" pageNumber="533">Fig. 6B</figureCitation>
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). Tegmina reduced to lateral flaps, with traces of venation, marginated laterally and rounded apically (
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<figureCitation id="136A2A29FFA9FFDEFE0BFF231A41FEC9" box="[427,525,151,176]" captionStart="FIGURE 6" captionStartId="11.[151,249,1908,1930]" captionTargetBox="[190,1396,592,1885]" captionTargetId="figure@11.[172,1416,581,1898]" captionTargetPageId="11" captionText="FIGURE 6. Neostylopyga yemenica (Bey-Bienko, 1969), male, holotype. A—facial part of head; B—head, thorax and 1–6th abdominal tergites from above; C—7th abdominal tergite from above; D—abdominal apex from above, cerci broken off; Ehypandrium from below, styli broken off. Dotted areas shows membranous parts. Abbreviations: I–VIII, X—1st–10th abdominal tergites respectively—see text. Scale bar 1 mm: a = A; b = B; c = C, D; d = E." httpUri="https://zenodo.org/record/2615643/files/figure.png" pageId="11" pageNumber="534">Fig. 6B</figureCitation>
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). Wings absent. Mesonotum and metanotum transverse, with weak medial impressions and caudal margins slightly angulate (
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<figureCitation id="136A2A29FFA9FFDEFD57FF081B1AFEAC" box="[759,854,188,213]" captionStart="FIGURE 6" captionStartId="11.[151,249,1908,1930]" captionTargetBox="[190,1396,592,1885]" captionTargetId="figure@11.[172,1416,581,1898]" captionTargetPageId="11" captionText="FIGURE 6. Neostylopyga yemenica (Bey-Bienko, 1969), male, holotype. A—facial part of head; B—head, thorax and 1–6th abdominal tergites from above; C—7th abdominal tergite from above; D—abdominal apex from above, cerci broken off; Ehypandrium from below, styli broken off. Dotted areas shows membranous parts. Abbreviations: I–VIII, X—1st–10th abdominal tergites respectively—see text. Scale bar 1 mm: a = A; b = B; c = C, D; d = E." httpUri="https://zenodo.org/record/2615643/files/figure.png" pageId="11" pageNumber="534">Fig. 6B</figureCitation>
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). Thoracal tergites and abdominal tergites II–VI marginated laterally (
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<figureCitation id="136A2A29FFA9FFDEFE27FF6B1993FE81" box="[391,479,223,248]" captionStart="FIGURE 6" captionStartId="11.[151,249,1908,1930]" captionTargetBox="[190,1396,592,1885]" captionTargetId="figure@11.[172,1416,581,1898]" captionTargetPageId="11" captionText="FIGURE 6. Neostylopyga yemenica (Bey-Bienko, 1969), male, holotype. A—facial part of head; B—head, thorax and 1–6th abdominal tergites from above; C—7th abdominal tergite from above; D—abdominal apex from above, cerci broken off; Ehypandrium from below, styli broken off. Dotted areas shows membranous parts. Abbreviations: I–VIII, X—1st–10th abdominal tergites respectively—see text. Scale bar 1 mm: a = A; b = B; c = C, D; d = E." httpUri="https://zenodo.org/record/2615643/files/figure.png" pageId="11" pageNumber="534">Fig. 6B</figureCitation>
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). Anterior margin of fore femur armed as in the
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<typeStatus id="54EA880EFFA9FFDEFC5EFF541C62FE81" box="[1022,1070,224,248]" pageId="11" pageNumber="534">type</typeStatus>
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A, with 14–15 spines, including 2 apical ones. Fore tibiae not thickened distally. Tibial spines well developed. Tarsi of mid and hind legs broken off. Fore tarsi: Basal tarsomere of fore tarsi (metatarsus) shorter than other segments combined, with 2 more or less equal rows of spines along lower margin; all euplantulae absent or vestigial; claws symmetrical, simple; arolium vestigial. Abdomen without visible glandular specializations; posterolateral angles of tergites attenuate caudally; tergite VI with caudal margin sinusoidally curved (
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<figureCitation id="136A2A29FFA9FFDEFD4AFE201B09FFD4" box="[746,837,404,429]" captionStart="FIGURE 6" captionStartId="11.[151,249,1908,1930]" captionTargetBox="[190,1396,592,1885]" captionTargetId="figure@11.[172,1416,581,1898]" captionTargetPageId="11" captionText="FIGURE 6. Neostylopyga yemenica (Bey-Bienko, 1969), male, holotype. A—facial part of head; B—head, thorax and 1–6th abdominal tergites from above; C—7th abdominal tergite from above; D—abdominal apex from above, cerci broken off; Ehypandrium from below, styli broken off. Dotted areas shows membranous parts. Abbreviations: I–VIII, X—1st–10th abdominal tergites respectively—see text. Scale bar 1 mm: a = A; b = B; c = C, D; d = E." httpUri="https://zenodo.org/record/2615643/files/figure.png" pageId="11" pageNumber="534">Fig. 6B</figureCitation>
|
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|
|
); tergite VII roundly projected caudally, with small median incision (
|
|||
|
|
<figureCitation id="136A2A29FFA9FFDEFEC2FE0319F3FFA9" box="[354,447,439,464]" captionStart="FIGURE 6" captionStartId="11.[151,249,1908,1930]" captionTargetBox="[190,1396,592,1885]" captionTargetId="figure@11.[172,1416,581,1898]" captionTargetPageId="11" captionText="FIGURE 6. Neostylopyga yemenica (Bey-Bienko, 1969), male, holotype. A—facial part of head; B—head, thorax and 1–6th abdominal tergites from above; C—7th abdominal tergite from above; D—abdominal apex from above, cerci broken off; Ehypandrium from below, styli broken off. Dotted areas shows membranous parts. Abbreviations: I–VIII, X—1st–10th abdominal tergites respectively—see text. Scale bar 1 mm: a = A; b = B; c = C, D; d = E." httpUri="https://zenodo.org/record/2615643/files/figure.png" pageId="11" pageNumber="534">Fig. 6C</figureCitation>
|
|||
|
|
). Anal plate (tergite X) caudally protruded, with distinct median incision on caudal margin (
|
|||
|
|
<figureCitation id="136A2A29FFA9FFDEFF58FE681918FF8C" box="[248,340,476,501]" captionStart="FIGURE 6" captionStartId="11.[151,249,1908,1930]" captionTargetBox="[190,1396,592,1885]" captionTargetId="figure@11.[172,1416,581,1898]" captionTargetPageId="11" captionText="FIGURE 6. Neostylopyga yemenica (Bey-Bienko, 1969), male, holotype. A—facial part of head; B—head, thorax and 1–6th abdominal tergites from above; C—7th abdominal tergite from above; D—abdominal apex from above, cerci broken off; Ehypandrium from below, styli broken off. Dotted areas shows membranous parts. Abbreviations: I–VIII, X—1st–10th abdominal tergites respectively—see text. Scale bar 1 mm: a = A; b = B; c = C, D; d = E." httpUri="https://zenodo.org/record/2615643/files/figure.png" pageId="11" pageNumber="534">Fig. 6D</figureCitation>
|
|||
|
|
). Cerci broken off. Hypandrium weakly asymmetrical, wide, antero-lateral parts (lateral sternal apodemes or apophyses) elongated; caudal margin angulate between styli (
|
|||
|
|
<figureCitation id="136A2A29FFA9FFDEFC7CFE4B1C78FC61" box="[988,1076,511,536]" captionStart="FIGURE 6" captionStartId="11.[151,249,1908,1930]" captionTargetBox="[190,1396,592,1885]" captionTargetId="figure@11.[172,1416,581,1898]" captionTargetPageId="11" captionText="FIGURE 6. Neostylopyga yemenica (Bey-Bienko, 1969), male, holotype. A—facial part of head; B—head, thorax and 1–6th abdominal tergites from above; C—7th abdominal tergite from above; D—abdominal apex from above, cerci broken off; Ehypandrium from below, styli broken off. Dotted areas shows membranous parts. Abbreviations: I–VIII, X—1st–10th abdominal tergites respectively—see text. Scale bar 1 mm: a = A; b = B; c = C, D; d = E." httpUri="https://zenodo.org/record/2615643/files/figure.png" pageId="11" pageNumber="534">Fig. 6E</figureCitation>
|
|||
|
|
); styli broken off.
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF2E6624FFA9FFDEFF37F8C01C51F99F" httpUri="https://zenodo.org/record/2615643/files/figure.png" pageId="11" pageNumber="534" startId="11.[151,249,1908,1930]" targetBox="[190,1396,592,1885]" targetPageId="11">
|
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|
|
<paragraph id="8BEE36ACFFA9FFDEFF37F8C01C51F99F" blockId="11.[151,1436,1908,2022]" pageId="11" pageNumber="534">
|
|||
|
|
<emphasis id="B925EABEFFA9FFDEFF37F8C0195FF9F3" bold="true" box="[151,275,1908,1930]" pageId="11" pageNumber="534">FIGURE 6.</emphasis>
|
|||
|
|
<taxonomicName id="4C514D2FFFA9FFDEFEBBF8C01AACF9F3" authority="(Bey-Bienko, 1969)" authorityName="Anisyutkin & Telnov" authorityYear="2018" baseAuthorityName="Bey-Bienko" baseAuthorityYear="1969" box="[283,736,1908,1931]" class="Insecta" family="Blattidae" genus="Neostylopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="11" pageNumber="534" phylum="Arthropoda" rank="species" species="yemenica">
|
|||
|
|
<emphasis id="B925EABEFFA9FFDEFEBBF8C01A40F9F2" box="[283,524,1908,1931]" italics="true" pageId="11" pageNumber="534">Neostylopyga yemenica</emphasis>
|
|||
|
|
(Bey-Bienko, 1969)
|
|||
|
|
</taxonomicName>
|
|||
|
|
, male, holotype. A—facial part of head; B—head, thorax and 1–6th abdominal tergites from above; C—7th abdominal tergite from above; D—abdominal apex from above, cerci broken off; Ehypandrium from below, styli broken off. Dotted areas shows membranous parts. Abbreviations:
|
|||
|
|
<emphasis id="B925EABEFFA9FFDEFB6AF807199FF99F" italics="true" pageId="11" pageNumber="534">I–VIII, X—1st–10th abdominal tergites respectively</emphasis>
|
|||
|
|
—see text. Scale bar 1 mm: a = A; b = B; c = C, D; d = E.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="DF2E6624FFAEFFD9FF37F8241C77F993" httpUri="https://zenodo.org/record/2615645/files/figure.png" pageId="12" pageNumber="535" startId="12.[151,249,1936,1958]" targetBox="[166,1407,200,1908]" targetPageId="12">
|
|||
|
|
<paragraph id="8BEE36ACFFAEFFD9FF37F8241C77F993" blockId="12.[151,1436,1936,2026]" pageId="12" pageNumber="535">
|
|||
|
|
<emphasis id="B925EABEFFAEFFD9FF37F8241958F9DF" bold="true" box="[151,276,1936,1958]" pageId="12" pageNumber="535">FIGURE 7.</emphasis>
|
|||
|
|
<taxonomicName id="4C514D2FFFAEFFD9FEBCF8241AA9F9DF" authority="(Bey-Bienko, 1969)" authorityName="Anisyutkin & Telnov" authorityYear="2018" baseAuthorityName="Bey-Bienko" baseAuthorityYear="1969" box="[284,741,1936,1958]" class="Insecta" family="Blattidae" genus="Neostylopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="12" pageNumber="535" phylum="Arthropoda" rank="species" species="yemenica">
|
|||
|
|
<emphasis id="B925EABEFFAEFFD9FEBCF8241A43F9DF" box="[284,527,1936,1958]" italics="true" pageId="12" pageNumber="535">Neostylopyga yemenica</emphasis>
|
|||
|
|
(Bey-Bienko, 1969)
|
|||
|
|
</taxonomicName>
|
|||
|
|
, male genitalia, holotype. A–C—right phallomere from above (A), below (B) and outside (C); D, E—ventral phallomere from below (D) and side (E). Dotted areas shows membranous parts. Abbreviations:
|
|||
|
|
<emphasis id="B925EABEFFAEFFD9FE94F8611A11F993" box="[308,605,2004,2026]" italics="true" pageId="12" pageNumber="535">a.t., c.out., R1G, R1H, R2, R3</emphasis>
|
|||
|
|
—see text. Scale bar 1 mm: a = A–C; b = D, E.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8BEE36ACFFAFFFD8FF67FF231902FC61" blockId="13.[151,1437,151,1112]" pageId="13" pageNumber="536">
|
|||
|
|
Male genitalia (
|
|||
|
|
<figureCitation id="136A2A29FFAFFFD8FE20FF231983FEC9" box="[384,463,151,176]" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. A–D—Duchailluia ivindo sp. nov.; E–H—Distylopyga beccalonii sp. nov.; I–L—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = A–D; b = E–H; c = I– L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="13" pageNumber="536">Figs. 3</figureCitation>
|
|||
|
|
I–L, 7A–E): left phallomere (
|
|||
|
|
<figureCitation id="136A2A29FFAFFFD8FC8CFF231B3CFEC9" box="[812,880,151,176]" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. A–D—Duchailluia ivindo sp. nov.; E–H—Distylopyga beccalonii sp. nov.; I–L—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = A–D; b = E–H; c = I– L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="13" pageNumber="536">Fig. 3</figureCitation>
|
|||
|
|
I–L) with sclerite L4C (L2D) long and narrow, caudally terminating in bulky process (
|
|||
|
|
<figureCitation id="136A2A29FFAFFFD8FDF2FF081AB5FEAD" box="[594,761,188,213]" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. A–D—Duchailluia ivindo sp. nov.; E–H—Distylopyga beccalonii sp. nov.; I–L—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = A–D; b = E–H; c = I– L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="13" pageNumber="536">
|
|||
|
|
Fig. 3J, K,
|
|||
|
|
<emphasis id="B925EABEFFAFFFD8FD73FF081AB5FEAD" box="[723,761,188,212]" italics="true" pageId="13" pageNumber="536">b.p</emphasis>
|
|||
|
|
</figureCitation>
|
|||
|
|
.); elongated field densely covered with teeth situated along L4C (
|
|||
|
|
<figureCitation id="136A2A29FFAFFFD8FF79FF6B193FFE81" box="[217,371,223,248]" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. A–D—Duchailluia ivindo sp. nov.; E–H—Distylopyga beccalonii sp. nov.; I–L—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = A–D; b = E–H; c = I– L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="13" pageNumber="536">
|
|||
|
|
Fig.
|
|||
|
|
<date id="FFEF106CFFAFFFD8FEAFFF541964FE81" box="[271,296,224,248]" pageId="13" pageNumber="536">3I</date>
|
|||
|
|
, K,
|
|||
|
|
<emphasis id="B925EABEFFAFFFD8FEF9FF54193FFE81" box="[345,371,224,248]" italics="true" pageId="13" pageNumber="536">t.f</emphasis>
|
|||
|
|
</figureCitation>
|
|||
|
|
.); sclerite L3 (L3d) robust, curved in apical part, with directed downward tooth; sclerite L4E, plate-like and curved; sclerite L4F weakly sclerotized, caudally fused with caudal process of sclerite L2; sclerite L4D (L
|
|||
|
|
<date id="FFEF106CFFAFFFD8FF4DFE9C1945FF39" box="[237,265,296,320]" pageId="13" pageNumber="536">3v</date>
|
|||
|
|
) large (
|
|||
|
|
<figureCitation id="136A2A29FFAFFFD8FEFFFE93198AFF39" box="[351,454,295,320]" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. A–D—Duchailluia ivindo sp. nov.; E–H—Distylopyga beccalonii sp. nov.; I–L—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = A–D; b = E–H; c = I– L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="13" pageNumber="536">
|
|||
|
|
Fig.
|
|||
|
|
<date id="FFEF106CFFAFFFD8FE33FE9C19E1FF39" box="[403,429,296,320]" pageId="13" pageNumber="536">3I</date>
|
|||
|
|
, J
|
|||
|
|
</figureCitation>
|
|||
|
|
); sclerite L2 (L
|
|||
|
|
<date id="FFEF106CFFAFFFD8FDD5FE9C1ADEFF39" box="[629,658,296,320]" pageId="13" pageNumber="536">2v</date>
|
|||
|
|
) large, widened cranially (
|
|||
|
|
<figureCitation id="136A2A29FFAFFFD8FC1EFE931C7BFF39" box="[958,1079,295,320]" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. A–D—Duchailluia ivindo sp. nov.; E–H—Distylopyga beccalonii sp. nov.; I–L—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = A–D; b = E–H; c = I– L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="13" pageNumber="536">Fig. 3K, L</figureCitation>
|
|||
|
|
), terminating in apically curved partly membranous caudal process (
|
|||
|
|
<figureCitation id="136A2A29FFAFFFD8FD87FEF81A82FF1D" box="[551,718,332,357]" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. A–D—Duchailluia ivindo sp. nov.; E–H—Distylopyga beccalonii sp. nov.; I–L—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = A–D; b = E–H; c = I– L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="13" pageNumber="536">
|
|||
|
|
Fig. 3K, L,
|
|||
|
|
<emphasis id="B925EABEFFAFFFD8FD09FEF91A82FF1D" box="[681,718,333,356]" italics="true" pageId="13" pageNumber="536">a.p</emphasis>
|
|||
|
|
</figureCitation>
|
|||
|
|
.); 2 partly sclerotized lobes situated above L2: ventral lobe (
|
|||
|
|
<figureCitation id="136A2A29FFAFFFD8FACFFEF818AAFFF1" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. A–D—Duchailluia ivindo sp. nov.; E–H—Distylopyga beccalonii sp. nov.; I–L—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = A–D; b = E–H; c = I– L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="13" pageNumber="536">
|
|||
|
|
Fig. 3K,
|
|||
|
|
<emphasis id="B925EABEFFAFFFD8FF6AFEC418AAFFF1" box="[202,230,368,392]" italics="true" pageId="13" pageNumber="536">v.l</emphasis>
|
|||
|
|
</figureCitation>
|
|||
|
|
.) with short toothed outgrowth at caudo-ventral angle and dorsal lobe (
|
|||
|
|
<figureCitation id="136A2A29FFAFFFD8FB81FEDB1CE7FFF1" box="[1057,1195,367,392]" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. A–D—Duchailluia ivindo sp. nov.; E–H—Distylopyga beccalonii sp. nov.; I–L—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = A–D; b = E–H; c = I– L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="13" pageNumber="536">
|
|||
|
|
Fig. 3K,
|
|||
|
|
<emphasis id="B925EABEFFAFFFD8FB2BFEC41CE7FFF1" box="[1163,1195,368,392]" italics="true" pageId="13" pageNumber="536">d.l</emphasis>
|
|||
|
|
</figureCitation>
|
|||
|
|
.), dorsally with setal brush; sclerite L1 small. Ventral phallomere L4G (VP) large, as in
|
|||
|
|
<figureCitation id="136A2A29FFAFFFD8FC23FE201BB7FFD5" box="[899,1019,404,429]" captionStart="FIGURE 7" captionStartId="12.[151,249,1936,1958]" captionTargetBox="[166,1407,200,1908]" captionTargetId="figure@12.[151,1436,193,1915]" captionTargetPageId="12" captionText="FIGURE 7. Neostylopyga yemenica (Bey-Bienko, 1969), male genitalia, holotype. A–C—right phallomere from above (A), below (B) and outside (C); D, E—ventral phallomere from below (D) and side (E). Dotted areas shows membranous parts. Abbreviations: a.t., c.out., R1G, R1H, R2, R3—see text. Scale bar 1 mm: a = A–C; b = D, E." httpUri="https://zenodo.org/record/2615645/files/figure.png" pageId="13" pageNumber="536">Fig. 7D, E</figureCitation>
|
|||
|
|
. Right phallomere complex in shape, as in
|
|||
|
|
<figureCitation id="136A2A29FFAFFFD8FF79FE031951FFA9" box="[217,285,439,464]" captionStart="FIGURE 7" captionStartId="12.[151,249,1936,1958]" captionTargetBox="[166,1407,200,1908]" captionTargetId="figure@12.[151,1436,193,1915]" captionTargetPageId="12" captionText="FIGURE 7. Neostylopyga yemenica (Bey-Bienko, 1969), male genitalia, holotype. A–C—right phallomere from above (A), below (B) and outside (C); D, E—ventral phallomere from below (D) and side (E). Dotted areas shows membranous parts. Abbreviations: a.t., c.out., R1G, R1H, R2, R3—see text. Scale bar 1 mm: a = A–C; b = D, E." httpUri="https://zenodo.org/record/2615645/files/figure.png" pageId="13" pageNumber="536">Fig. 7</figureCitation>
|
|||
|
|
A–C; basal sclerite (R2) large; sclerite R1H plate-like, with 2 medially directed spines and narrow cranial outgrowth (
|
|||
|
|
<figureCitation id="136A2A29FFAFFFD8FECCFE681990FF8D" box="[364,476,476,501]" captionStart="FIGURE 7" captionStartId="12.[151,249,1936,1958]" captionTargetBox="[166,1407,200,1908]" captionTargetId="figure@12.[151,1436,193,1915]" captionTargetPageId="12" captionText="FIGURE 7. Neostylopyga yemenica (Bey-Bienko, 1969), male genitalia, holotype. A–C—right phallomere from above (A), below (B) and outside (C); D, E—ventral phallomere from below (D) and side (E). Dotted areas shows membranous parts. Abbreviations: a.t., c.out., R1G, R1H, R2, R3—see text. Scale bar 1 mm: a = A–C; b = D, E." httpUri="https://zenodo.org/record/2615645/files/figure.png" pageId="13" pageNumber="536">
|
|||
|
|
Fig. 7A,
|
|||
|
|
<emphasis id="B925EABEFFAFFFD8FE6FFE691990FF8D" box="[463,476,477,500]" italics="true" pageId="13" pageNumber="536">c</emphasis>
|
|||
|
|
</figureCitation>
|
|||
|
|
.out.); sclerite R1G large, with single curved apical tooth (
|
|||
|
|
<figureCitation id="136A2A29FFAFFFD8FBC4FE681CE9FF8D" box="[1124,1189,476,501]" captionStart="FIGURE 7" captionStartId="12.[151,249,1936,1958]" captionTargetBox="[166,1407,200,1908]" captionTargetId="figure@12.[151,1436,193,1915]" captionTargetPageId="12" captionText="FIGURE 7. Neostylopyga yemenica (Bey-Bienko, 1969), male genitalia, holotype. A–C—right phallomere from above (A), below (B) and outside (C); D, E—ventral phallomere from below (D) and side (E). Dotted areas shows membranous parts. Abbreviations: a.t., c.out., R1G, R1H, R2, R3—see text. Scale bar 1 mm: a = A–C; b = D, E." httpUri="https://zenodo.org/record/2615645/files/figure.png" pageId="13" pageNumber="536">Fig. 7</figureCitation>
|
|||
|
|
A–C,
|
|||
|
|
<emphasis id="B925EABEFFAFFFD8FB48FE691D40FF8D" box="[1256,1292,477,500]" italics="true" pageId="13" pageNumber="536">a.t.</emphasis>
|
|||
|
|
); sclerite R3 large, plate-like.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BEE36ACFFAFFFD8FF67FD901B2EFC19" blockId="13.[151,1437,151,1112]" pageId="13" pageNumber="536">
|
|||
|
|
<emphasis id="B925EABEFFAFFFD8FF67FD90193AFC44" bold="true" box="[199,374,548,573]" pageId="13" pageNumber="536">Measurements</emphasis>
|
|||
|
|
(mm). Head length: 4.8, width 4.5, pronotum length 6.5, width 8.9, mesonotum length 3.0, tegmina length 4.8, width 3.2, metanotum length 2.4, width 9.8.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BEE36ACFFAFFFD8FF67FDD819F3FCB4" blockId="13.[151,1437,151,1112]" pageId="13" pageNumber="536">
|
|||
|
|
<emphasis id="B925EABEFFAFFFD8FF67FDD81946FCFC" bold="true" box="[199,266,620,645]" pageId="13" pageNumber="536">Notes</emphasis>
|
|||
|
|
. This species was described from the basis of two damaged specimens (male
|
|||
|
|
<typeStatus id="54EA880EFFAFFFD8FB0EFDD81D5CFCFC" box="[1198,1296,620,645]" pageId="13" pageNumber="536" type="holotype">holotype</typeStatus>
|
|||
|
|
and female
|
|||
|
|
<typeStatus id="54EA880EFFAFFFD8FF37FD2418B5FCD1" box="[151,249,656,680]" pageId="13" pageNumber="536" type="paratype">paratype</typeStatus>
|
|||
|
|
) and provisionally ascribed to the genus
|
|||
|
|
<taxonomicName id="4C514D2FFFAFFFD8FD73FD241B15FCD1" baseAuthorityName="Roth" baseAuthorityYear="1988" box="[723,857,656,680]" class="Insecta" family="Blattidae" genus="Duchailluia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="13" pageNumber="536" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B925EABEFFAFFFD8FD73FD241B15FCD1" box="[723,857,656,680]" italics="true" pageId="13" pageNumber="536">Duchailluia</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(see
|
|||
|
|
<bibRefCitation id="EFC04B5DFFAFFFD8FC38FD3B1C28FCD1" author="Bey-Bienko, G. Ya." box="[920,1124,655,680]" pageId="13" pageNumber="536" pagination="831 - 862" refId="ref7112" refString="Bey-Bienko, G. Ya. (1969) New genera and species of cockroaches (Blattoptera) from tropical and subtropical Asia. Entomologicheskoe Obozrenie, 48 (4), 831 - 862. [in Russian, English abstract]" type="journal article" year="1969">Bey-Bienko 1969</bibRefCitation>
|
|||
|
|
). The structure of the male genitalia were not studied.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BEE36ACFFAFFFD8FF67FD631C75FDB1" blockId="13.[151,1437,151,1112]" pageId="13" pageNumber="536">
|
|||
|
|
The structure of the male genitalia of
|
|||
|
|
<taxonomicName id="4C514D2FFFAFFFD8FD35FD6D1B68FC96" authorityName="Anisyutkin & Telnov" authorityYear="2018" baseAuthorityName="Bey-Bienko" baseAuthorityYear="1969" box="[661,804,728,752]" class="Insecta" family="Blattidae" genus="Neostylopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="13" pageNumber="536" phylum="Arthropoda" rank="species" species="yemenica">
|
|||
|
|
<emphasis id="B925EABEFFAFFFD8FD35FD6D1AE4FC89" box="[661,680,729,752]" italics="true" pageId="13" pageNumber="536">N</emphasis>
|
|||
|
|
.
|
|||
|
|
<emphasis id="B925EABEFFAFFFD8FD1BFD6C1B68FC96" box="[699,804,728,751]" italics="true" pageId="13" pageNumber="536">yemenica</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is dissimilar to those of the genera
|
|||
|
|
<taxonomicName id="4C514D2FFFAFFFD8FB41FD6C1D2BFC89" baseAuthorityName="Roth" baseAuthorityYear="1988" box="[1249,1383,728,752]" class="Insecta" family="Blattidae" genus="Duchailluia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="13" pageNumber="536" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B925EABEFFAFFFD8FB41FD6C1D2BFC89" box="[1249,1383,728,752]" italics="true" pageId="13" pageNumber="536">Duchailluia</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C514D2FFFAFFFD8FF37FD481952FD6D" authorityName="Bey-Bienko" authorityYear="1965" box="[151,286,764,788]" class="Insecta" family="Blattidae" genus="Distylopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="13" pageNumber="536" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B925EABEFFAFFFD8FF37FD481952FD6D" box="[151,286,764,788]" italics="true" pageId="13" pageNumber="536">Distylopyga</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(see above). On the contrary, the structure of the male genitalia of
|
|||
|
|
<taxonomicName id="4C514D2FFFAFFFD8FBABFD491CDAFD6D" authorityName="Anisyutkin & Telnov" authorityYear="2018" baseAuthorityName="Bey-Bienko" baseAuthorityYear="1969" box="[1035,1174,765,788]" class="Insecta" family="Blattidae" genus="Neostylopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="13" pageNumber="536" phylum="Arthropoda" rank="species" species="yemenica">
|
|||
|
|
<emphasis id="B925EABEFFAFFFD8FBABFD491C52FD6D" box="[1035,1054,765,788]" italics="true" pageId="13" pageNumber="536">N</emphasis>
|
|||
|
|
.
|
|||
|
|
<emphasis id="B925EABEFFAFFFD8FB8CFD491CDAFD6D" box="[1068,1174,765,788]" italics="true" pageId="13" pageNumber="536">yemenica</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
generally resembles
|
|||
|
|
<taxonomicName id="4C514D2FFFAFFFD8FA22FD491957FD41" authorityName="Stoll" authorityYear="1813" class="Insecta" family="Blattidae" genus="Blatta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="13" pageNumber="536" phylum="Arthropoda" rank="species" species="rhombifolia">
|
|||
|
|
<emphasis id="B925EABEFFAFFFD8FA22FD491DD9FD6D" box="[1410,1429,765,788]" italics="true" pageId="13" pageNumber="536">N</emphasis>
|
|||
|
|
.
|
|||
|
|
<emphasis id="B925EABEFFAFFFD8FF37FC941957FD41" box="[151,283,800,824]" italics="true" pageId="13" pageNumber="536">rhombifolia</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, the
|
|||
|
|
<typeStatus id="54EA880EFFAFFFD8FEFCFC9419C0FD41" box="[348,396,800,824]" pageId="13" pageNumber="536">type</typeStatus>
|
|||
|
|
species of the genus
|
|||
|
|
<taxonomicName id="4C514D2FFFAFFFD8FD33FC941B67FD41" authorityName="Shelford" authorityYear="1911" box="[659,811,800,824]" class="Insecta" family="Blattidae" genus="Neostylopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="13" pageNumber="536" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B925EABEFFAFFFD8FD33FC941B67FD41" box="[659,811,800,824]" italics="true" pageId="13" pageNumber="536">Neostylopyga</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. There are mutual characters in
|
|||
|
|
<taxonomicName id="4C514D2FFFAFFFD8FB13FC951D0DFD4E" authorityName="Anisyutkin & Telnov" authorityYear="2018" baseAuthorityName="Bey-Bienko" baseAuthorityYear="1969" box="[1203,1345,800,824]" class="Insecta" family="Blattidae" genus="Neostylopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="13" pageNumber="536" phylum="Arthropoda" rank="species" species="yemenica">
|
|||
|
|
<emphasis id="B925EABEFFAFFFD8FB13FC951C8AFD41" box="[1203,1222,801,824]" italics="true" pageId="13" pageNumber="536">N</emphasis>
|
|||
|
|
.
|
|||
|
|
<emphasis id="B925EABEFFAFFFD8FB78FC941D0DFD4E" box="[1240,1345,800,823]" italics="true" pageId="13" pageNumber="536">yemenica</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C514D2FFFAFFFD8FA22FC951957FD25" authorityName="Stoll" authorityYear="1813" class="Insecta" family="Blattidae" genus="Blatta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="13" pageNumber="536" phylum="Arthropoda" rank="species" species="rhombifolia">
|
|||
|
|
<emphasis id="B925EABEFFAFFFD8FA22FC951DD9FD41" box="[1410,1429,801,824]" italics="true" pageId="13" pageNumber="536">N</emphasis>
|
|||
|
|
.
|
|||
|
|
<emphasis id="B925EABEFFAFFFD8FF37FCF01957FD25" box="[151,283,836,860]" italics="true" pageId="13" pageNumber="536">rhombifolia</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
: left phallomere with sclerite L4C long and narrow, sclerites L4E and L4F separated, sclerite L2 widened cranially, terminating in caudal process, 2 partly sclerotized lobes situated on inner side of phallomere (compare
|
|||
|
|
<figureCitation id="136A2A29FFAFFFD8FEA8FC381904FDDD" box="[264,328,908,933]" captionStart="FIGURE 3" captionStartId="6.[151,249,1878,1900]" captionTargetBox="[230,1361,198,1854]" captionTargetId="figure@6.[225,1367,188,1868]" captionTargetPageId="6" captionText="FIGURE 3. Left phallomere of the male genitalia (holotypes): from above (A, E, I), outside (B, F, J), inside (C, G, K) and below (D, H); L—left phallomere from below, partly. A–D—Duchailluia ivindo sp. nov.; E–H—Distylopyga beccalonii sp. nov.; I–L—Neostylopyga yemenica (Bey-Bienko, 1969). Dotted areas shows membranous parts. Abbreviations: a.c.p., a.e.s., a.p., b.p., d.c.p., d.l., f.c.p., L2, L3, L4C, L4D, L4F, m.l., r.m.l., s.l., t.f., v.l.—see text. Scale bar 1 mm: a = A–D; b = E–H; c = I– L." httpUri="https://zenodo.org/record/2615637/files/figure.png" pageId="13" pageNumber="536">Fig. 3</figureCitation>
|
|||
|
|
I–L and
|
|||
|
|
<figureCitation id="136A2A29FFAFFFD8FE09FC3819A6FDDC" box="[425,490,908,933]" captionStart="FIGURE 5" captionStartId="9.[151,249,1939,1961]" captionTargetBox="[191,1409,352,1920]" captionTargetId="figure@9.[171,1424,339,1925]" captionTargetPageId="9" captionText="FIGURE 5. Distylopyga beccalonii Anisyutkin, sp. nov., male genitalia, holotype. A–C—right phallomere from above (A), below (B) and outside (C); D, E—ventral phallomere from below (D) and side (E). Dotted areas shows membranous parts. Abbreviations: a.p., a.t., R1G, R1Hi., R1Hout., R2, R3—see text. Scale bar 1 mm." httpUri="https://zenodo.org/record/2615641/files/figure.png" pageId="13" pageNumber="536">
|
|||
|
|
Fig.
|
|||
|
|
<quantity id="4CA99B49FFAFFFD8FE7DFC3819A6FDDC" box="[477,490,908,933]" metricMagnitude="-1" metricUnit="m" metricValue="1.651" metricValueMax="2.032" metricValueMin="1.27" pageId="13" pageNumber="536" unit="in" value="6.5" valueMax="8.0" valueMin="5.0">5</quantity>
|
|||
|
|
</figureCitation>
|
|||
|
|
–8 in
|
|||
|
|
<bibRefCitation id="EFC04B5DFFAFFFD8FD8CFC381AA1FDDD" author="Anisyutkin, L. N." box="[556,749,908,933]" pageId="13" pageNumber="536" pagination="871 - 876" refId="ref6769" refString="Anisyutkin, L. N. (2010) New data on the genus Neostylopyga Shelford, 1911 (Dictyoptera, Blattidae) with description of a new species from Laos. Entomologicheskoe Obozrenie, 89 (3), 596 - 602. [in Russian]. English translation: Anisyutkin, L. N. (2010) New data on the genus Neostylopyga Shelford, 1911 (Dictyoptera, Blattidae) with description of a new species from Laos. Entomological Review, 90 (7), 871 - 876. https: // doi. org / 10.1134 / S 0013873810070055" type="journal article" year="2010">Anisyutkin 2010</bibRefCitation>
|
|||
|
|
); right phallomere with sclerite R1H plate-like, with 2 spines and narrow cranial outgrowth (compare
|
|||
|
|
<figureCitation id="136A2A29FFAFFFD8FDFBFC1B1AD7FDB1" box="[603,667,943,968]" captionStart="FIGURE 7" captionStartId="12.[151,249,1936,1958]" captionTargetBox="[166,1407,200,1908]" captionTargetId="figure@12.[151,1436,193,1915]" captionTargetPageId="12" captionText="FIGURE 7. Neostylopyga yemenica (Bey-Bienko, 1969), male genitalia, holotype. A–C—right phallomere from above (A), below (B) and outside (C); D, E—ventral phallomere from below (D) and side (E). Dotted areas shows membranous parts. Abbreviations: a.t., c.out., R1G, R1H, R2, R3—see text. Scale bar 1 mm: a = A–C; b = D, E." httpUri="https://zenodo.org/record/2615645/files/figure.png" pageId="13" pageNumber="536">Fig. 7</figureCitation>
|
|||
|
|
A–C and
|
|||
|
|
<figureCitation id="136A2A29FFAFFFD8FCA7FC1B1B05FDB1" box="[775,841,943,968]" captionStart="FIGURE 4" captionStartId="7.[151,249,1811,1833]" captionTargetBox="[219,1368,193,1791]" captionTargetId="figure@7.[212,1380,183,1798]" captionTargetPageId="7" captionText="FIGURE 4. Distylopyga beccalonii Anisyutkin, sp. nov., male, holotype (A–K) and female, paratype (L–N). A, L—facial part of head; B—head, thorax and six abdominal tergites from above; C—left tegmen from side; D—left hind tarsus from outside; E—1st–4th segments of left hind tarsus from below; F—arolium and claws of left hind tarsus; G—abdominal apex from above, only bases of cerci are shown; H—right cercus from inside; I—hypandrium from below; J, K—caudal margin of hypandrium from above (J) and caudal (K); M—general view from above; N—genital plate from below. Dotted areas shows membranous parts. Dotted line in Fig. 4A, L delimits areas of ocellar spots. Abbreviations: I–X—1st–10th abdominal tergites respectively, a.s., par., pv.– see text. Scale bar 1 mm: a = A, L; b = B; c = C; d = D; e = E; f = F; g = G, H; i = J, K; j = M; k = N." httpUri="https://zenodo.org/record/2615639/files/figure.png" pageId="13" pageNumber="536">
|
|||
|
|
Fig.
|
|||
|
|
<quantity id="4CA99B49FFAFFFD8FC9BFC041B05FDB1" box="[827,841,944,968]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="13" pageNumber="536" unit="in" value="4.0">4</quantity>
|
|||
|
|
</figureCitation>
|
|||
|
|
in
|
|||
|
|
<bibRefCitation id="EFC04B5DFFAFFFD8FCCCFC1B1C61FDB1" author="Anisyutkin, L. N." box="[876,1069,943,968]" pageId="13" pageNumber="536" pagination="871 - 876" refId="ref6769" refString="Anisyutkin, L. N. (2010) New data on the genus Neostylopyga Shelford, 1911 (Dictyoptera, Blattidae) with description of a new species from Laos. Entomologicheskoe Obozrenie, 89 (3), 596 - 602. [in Russian]. English translation: Anisyutkin, L. N. (2010) New data on the genus Neostylopyga Shelford, 1911 (Dictyoptera, Blattidae) with description of a new species from Laos. Entomological Review, 90 (7), 871 - 876. https: // doi. org / 10.1134 / S 0013873810070055" type="journal article" year="2010">Anisyutkin 2010</bibRefCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BEE36ACFFAFFFD8FF67FC601D79FD95" blockId="13.[151,1437,151,1112]" box="[199,1333,980,1005]" pageId="13" pageNumber="536">
|
|||
|
|
Taking into consideration the aforesaid, we herewith transfer
|
|||
|
|
<taxonomicName id="4C514D2FFFAFFFD8FCD6FC611C4DFD95" authorityName="Anisyutkin & Telnov" authorityYear="2018" baseAuthorityName="Bey-Bienko" baseAuthorityYear="1969" box="[886,1025,981,1004]" class="Insecta" family="Blattidae" genus="Neostylopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="13" pageNumber="536" phylum="Arthropoda" rank="species" species="yemenica">
|
|||
|
|
<emphasis id="B925EABEFFAFFFD8FCD6FC611BC5FD95" box="[886,905,981,1004]" italics="true" pageId="13" pageNumber="536">N</emphasis>
|
|||
|
|
.
|
|||
|
|
<emphasis id="B925EABEFFAFFFD8FC37FC611C4DFD95" box="[919,1025,981,1004]" italics="true" pageId="13" pageNumber="536">yemenica</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
to the genus
|
|||
|
|
<taxonomicName id="4C514D2FFFAFFFD8FB36FC601D62FD95" authorityName="Shelford" authorityYear="1911" box="[1174,1326,980,1004]" class="Insecta" family="Blattidae" genus="Neostylopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="13" pageNumber="536" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B925EABEFFAFFFD8FB36FC601D62FD95" box="[1174,1326,980,1004]" italics="true" pageId="13" pageNumber="536">Neostylopyga</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BEE36ACFFAFFFD8FF67FC431B84FA21" blockId="13.[151,1437,151,1112]" pageId="13" pageNumber="536">
|
|||
|
|
The sclerites R1H and R1G of
|
|||
|
|
<taxonomicName id="4C514D2FFFAFFFD8FDBEFC431AF6FA69" authorityName="Shelford" authorityYear="1911" box="[542,698,1015,1040]" class="Insecta" family="Blattidae" genus="Neostylopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="13" pageNumber="537" phylum="Arthropoda" rank="genus">Neostylopyga</taxonomicName>
|
|||
|
|
are similar to those of
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<taxonomicName id="4C514D2FFFAFFFD8FC17FC4C1DD4FA69" authority="(Fabricius, 1781)" authorityName="Fabricius" authorityYear="1781" baseAuthorityName="Fabricius" baseAuthorityYear="1781" box="[951,1432,1015,1040]" class="Insecta" family="Blattidae" genus="Deropeltis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="13" pageNumber="536" phylum="Arthropoda" rank="species" species="erytrocephala">
|
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<emphasis id="B925EABEFFAFFFD8FC17FC4C1C82FA69" box="[951,1230,1016,1040]" italics="true" pageId="13" pageNumber="536">Deropeltis erytrocephala</emphasis>
|
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|
|
(
|
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<bibRefCitation id="EFC04B5DFFAFFFD8FB7CFC431DC3FA69" author="Fabricius, I. C." box="[1244,1423,1015,1040]" pageId="13" pageNumber="536" refId="ref7203" refString="Fabricius, I. C. (1781) Species insectorum: exhibentes differentias specificas, synonyma auctorum, loca natalia, metamorphosin adiectis observationibus, descriptionibus. Vol. 1. impensis C. E. Bohnii, Hamburgi et Kilonii, 552 pp." type="book" year="1781">Fabricius, 1781</bibRefCitation>
|
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)
|
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|
|
</taxonomicName>
|
|||
|
|
, the
|
|||
|
|
<typeStatus id="54EA880EFFAFFFD8FF1FFBA918A3FA4C" box="[191,239,1053,1077]" pageId="13" pageNumber="536">type</typeStatus>
|
|||
|
|
species of the genus
|
|||
|
|
<taxonomicName id="4C514D2FFFAFFFD8FE7BFBA81B56FA4D" authority="Burmeister, 1838" authorityName="Burmeister" authorityYear="1838" box="[475,794,1052,1077]" class="Insecta" family="Blattidae" genus="Deropeltis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="13" pageNumber="536" phylum="Arthropoda" rank="genus">
|
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|
|
<emphasis id="B925EABEFFAFFFD8FE7BFBA81A1CFA4D" box="[475,592,1052,1076]" italics="true" pageId="13" pageNumber="536">Deropeltis</emphasis>
|
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|
|
<bibRefCitation id="EFC04B5DFFAFFFD8FDF7FBA81B56FA4D" author="Burmeister, H." box="[599,794,1052,1077]" pageId="13" pageNumber="536" refId="ref7177" refString="Burmeister, H. (1838) Handbuch der Entomologie. Band 2. Abteilung 2. G. Reimer, Berlin, 654 pp." type="book" year="1838">Burmeister, 1838</bibRefCitation>
|
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|
|
</taxonomicName>
|
|||
|
|
, and
|
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|
|
<taxonomicName id="4C514D2FFFAFFFD8FCF3FBA81D75FA4D" authority="Anisyutkin, 2014" authorityName="Anisyutkin" authorityYear="2014" box="[851,1337,1052,1077]" class="Insecta" family="Blattidae" genus="Afrostylopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="13" pageNumber="536" phylum="Arthropoda" rank="species" species="angolensis">
|
|||
|
|
<emphasis id="B925EABEFFAFFFD8FCF3FBA81C22FA4D" box="[851,1134,1052,1076]" italics="true" pageId="13" pageNumber="536">Afrostylopyga angolensis</emphasis>
|
|||
|
|
<bibRefCitation id="EFC04B5DFFAFFFD8FBD4FBA81D75FA4D" author="Anisyutkin, L. N." box="[1140,1337,1052,1077]" pageId="13" pageNumber="536" pagination="33 - 50" refId="ref6876" refString="Anisyutkin, L. N. (2014) New and little known Blattidae (Dictyoptera) from the collection of the Museum d'histoire naturelle de Geneve. Revue suisse de Zoologie, 121 (1), 33 - 50." type="journal article" year="2014">Anisyutkin, 2014</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, the
|
|||
|
|
<typeStatus id="54EA880EFFAFFFD8FACCFBA91DD0FA4C" box="[1388,1436,1053,1077]" pageId="13" pageNumber="536">type</typeStatus>
|
|||
|
|
species of the genus
|
|||
|
|
<taxonomicName id="4C514D2FFFAFFFD8FEDEFBF41AABFA21" authority="Anisyutkin, 2014" authorityName="Anisyutkin" authorityYear="2014" box="[382,743,1087,1112]" class="Insecta" family="Blattidae" genus="Afrostylopyga" higherTaxonomySource="GBIF" kingdom="Animalia" order="Blattodea" pageId="13" pageNumber="536" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B925EABEFFAFFFD8FEDEFBF41A56FA21" box="[382,538,1088,1112]" italics="true" pageId="13" pageNumber="536">Afrostylopyga</emphasis>
|
|||
|
|
<bibRefCitation id="EFC04B5DFFAFFFD8FD82FB8B1AABFA21" author="Anisyutkin, L. N." box="[546,743,1087,1112]" pageId="13" pageNumber="536" pagination="33 - 50" refId="ref6876" refString="Anisyutkin, L. N. (2014) New and little known Blattidae (Dictyoptera) from the collection of the Museum d'histoire naturelle de Geneve. Revue suisse de Zoologie, 121 (1), 33 - 50." type="journal article" year="2014">Anisyutkin, 2014</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, (
|
|||
|
|
<bibRefCitation id="EFC04B5DFFAFFFD8FD59FB8B1BF0FA21" author="Anisyutkin, L. N." box="[761,956,1087,1112]" pageId="13" pageNumber="536" pagination="33 - 50" refId="ref6876" refString="Anisyutkin, L. N. (2014) New and little known Blattidae (Dictyoptera) from the collection of the Museum d'histoire naturelle de Geneve. Revue suisse de Zoologie, 121 (1), 33 - 50." type="journal article" year="2014">Anisyutkin 2014</bibRefCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|