treatments-xml/data/90/3C/80/903C80318771595EFEFEBEC9FAB7FDEB.xml

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<document id="DF888F6C0164B99FF031574E047A9C11" ID-ISSN="2201-4349" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629580321750" checkinUser="felipe" docAuthor="Cairns, S. D." docDate="2004" docId="903C80318771595EFEFEBEC9FAB7FDEB" docLanguage="en" docName="RecAustMus.56.3.259-329.pdf" docOrigin="Records of the Australian Museum 56" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Conocyathus formosus Cairns 2004, n.sp." docType="treatment" docVersion="2" lastPageNumber="290" masterDocId="6C05F849876E5941FFF1BE50FF96FF9F" masterDocTitle="The Azooxanthellate Scleractinia (Coelenterata: Anthozoa) of Australia" masterLastPageNumber="329" masterPageNumber="259" pageNumber="290" updateTime="1699080756492" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="8A24508C2068124856930EF239825208">The Azooxanthellate Scleractinia (Coelenterata: Anthozoa) of Australia</mods:title>
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<mods:namePart id="E70F58DB8EEE2FC93611D83C35FFFF5D">Cairns, S. D.</mods:namePart>
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<mods:title id="BCF846C2AB56C960F7EE290E43C72E4C">Records of the Australian Museum</mods:title>
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<mods:part id="3A8FDE97C01F4BA791F34C51AE3119E1">
<mods:date id="DBC7489F6662989E31F12619D7B426EE">2004</mods:date>
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<paragraph id="182A31278771595EFEFEBEC9FDC0FF2E" blockId="31.[271,598,153,177]" box="[271,598,153,177]" pageId="31" pageNumber="290">
<heading id="4362864B8771595EFEFEBEC9FDC0FF2E" box="[271,598,153,177]" centered="true" fontSize="11" level="2" pageId="31" pageNumber="290" reason="2">
<emphasis id="2AE1ED358771595EFEFEBEC9FDC0FF2E" bold="true" box="[271,598,153,177]" pageId="31" pageNumber="290">
<taxonomicName id="DF954AA48771595EFEFEBEC9FD8EFF2E" ID-CoL="XQPZ" authority="Cairns, 2004" authorityName="Cairns" authorityYear="2004" box="[271,536,153,177]" class="Anthozoa" family="Turbinoliidae" genus="Conocyathus" kingdom="Animalia" order="Scleractinia" pageId="31" pageNumber="290" phylum="Cnidaria" rank="species" species="formosus" status="sp. nov.">
<emphasis id="2AE1ED358771595EFEFEBEC9FD8EFF2E" bold="true" box="[271,536,153,177]" italics="true" pageId="31" pageNumber="290">Conocyathus formosus</emphasis>
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<subSubSection id="508F62AC8771595EFEBABE92FB22FE27" pageId="31" pageNumber="290" type="description">
<paragraph id="182A31278771595EFEBABE92FD8CFF46" blockId="31.[331,538,194,217]" box="[331,538,194,217]" pageId="31" pageNumber="290">
<figureCitation id="80AE2DA28771595EFEBABE92FE5AFF46" box="[331,460,194,217]" captionStart="Fig" captionStartId="30.[166,202,1793,1814]" captionTargetBox="[166,1467,151,1776]" captionTargetId="figure-6@30.[166,1467,151,1776]" captionTargetPageId="30" captionText="Fig. 5. (A, B), Trochocyathus wellsi, holotype, AM G16704, stereo calicular and lateral views, ×6.4, ×3.8, respectively (GCD=7.8 mm). (C, D), Vaughanella oreophila, Franklin 06/88/x, AM G16707, calicular and lateral views, both ×3.9 (CD=13.1 mm); (E), Alatotrochus rubescens, USNM 1008301 (SEM 996), juvenile corallum showing pronounced edge and basal crests, ×11.8 (GCD=3.6 mm). (FH), Conocyathus formosus, holotype,AM G16743: (F), inner view of septa, ×35; (G), stereo calicular view, ×19.1; (H), lateral view, ×17.1 (CD=2.76 mm)." pageId="31" pageNumber="290">Figs. 5FH</figureCitation>
,
<figureCitation id="80AE2DA28771595EFE29BE92FD8CFF46" box="[472,538,194,217]" captionStart="Fig" captionStartId="32.[166,202,1793,1814]" captionTargetBox="[166,1467,151,1776]" captionTargetId="figure-6@32.[166,1467,151,1776]" captionTargetPageId="32" captionText="Fig. 6. (AC), Conocyathus formosus (A, C, paratype, USNM 1008831, CD=2.5 mm; B, holotype, CD=2.76 mm): lateral, oblique calicular, and lateral basal views, the latter showing the short, disjunct costae, ×15, ×16.5, ×9.6 respectively. (D, E), Endopachys australiae (=Cyathotrochus pileus), holotype, Macleay Museum, lateral and oblique calicular views, both ×3.5 (GCD=12.4 mm). (F J), Dunocyathus wallaceae (F, G, I, holotype, AM G16744, GCD=4.4 mm; (H, J), paratype, USNM 1008835, intercostal width = 0.23 mm): (F, G), lateral and stereo calicular views, both ×10.8; (H), spiny base, ×21; (I), palar and columellar region, ×21; (J), detail of thecal edge showing alternation of septa and costae, ×44." pageId="31" pageNumber="290">6AC</figureCitation>
</paragraph>
<paragraph id="182A31278771595EFF88BEBAFD0DFE73" blockId="31.[121,749,234,492]" pageId="31" pageNumber="290">
<emphasis id="2AE1ED358771595EFF88BEBAFEB6FE9D" bold="true" box="[121,288,234,258]" pageId="31" pageNumber="290">Records/Types</emphasis>
.
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<typeStatus id="C72E8F858771595EFEDDBEBAFE05FE9E" box="[300,403,234,257]" pageId="31" pageNumber="290" type="holotype">Holotype</typeStatus>
:
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03/99/D11,
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<geoCoordinate id="7DA157E08771595EFF88BF58FF68FE80" box="[121,254,264,287]" degrees="151" direction="east" minutes="47.53" orientation="longitude" pageId="31" pageNumber="290" precision="9" value="151.79216">151°47.53'E</geoCoordinate>
(
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,
<collectingRegion id="DA51FFC58771595EFE36BF58FDDDFE80" box="[455,587,264,287]" country="Australia" name="Queensland" pageId="31" pageNumber="290">Queensland</collectingRegion>
),
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, (type locality),
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G16743
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.
<materialsCitation id="A8FD3B7A8771595EFE60BF75FD9FFEC6" collectionCode="USNM" pageId="31" pageNumber="290" specimenCode="USNM 1008829" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
<typeStatus id="C72E8F858771595EFE60BF75FD93FEA3" box="[401,517,293,316]" pageId="31" pageNumber="290" type="paratype">Paratypes</typeStatus>
: Franklin 03/99/D8 (Marion Plateau), 1,
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</materialsCitation>
;
<materialsCitation id="A8FD3B7A8771595EFDE3BF12FEC4FEE8" collectionCode="USNM" pageId="31" pageNumber="290" specimenCode="USNM 1008830" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
Franklin 03/99/D10, 12,
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;
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Franklin 03/99/D11, 48: 43 (including
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stub 1005),
<specimenCode id="4833995C8771595EFEB4BF2DFD96FE0B" box="[325,512,381,404]" collectionCode="USNM" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="31" pageNumber="290" type="Museum">USNM 1008831</specimenCode>
,
<specimenCode id="4833995C8771595EFDFEBF2DFD89FE0B" box="[527,543,381,404]" collectionCode="USNM" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="31" pageNumber="290" type="Museum">2</specimenCode>
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,
<materialsCitation id="A8FD3B7A8771595EFDDFBF2DFF46FE2E" accessionNumber="Z20515" collectionCode="WAM" pageId="31" pageNumber="290" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
ZMUZ, 3,
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Z20515
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;
<materialsCitation id="A8FD3B7A8771595EFF2DBFCAFD10FE2E" box="[220,646,410,433]" collectionCode="USNM" pageId="31" pageNumber="290" specimenCode="USNM 1008832" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
Franklin, 03/99/12, 3,
<specimenCode id="4833995C8771595EFE3EBFCAFD10FE2E" box="[463,646,410,433]" collectionCode="USNM" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="31" pageNumber="290" type="Museum">USNM 1008832</specimenCode>
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;
<materialsCitation id="A8FD3B7A8771595EFD60BFCAFEA9FE73" collectionCode="USNM" latitude="-12.471666" longLatPrecision="92" longitude="130.84917" pageId="31" pageNumber="290" specimenCode="USNM 1008833" specimenCount="1" stateProvince="Queensland" typeStatus="paratype">
Franklin 03/99/D13, 1,
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;
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<geoCoordinate id="7DA157E08771595EFD95BFE8FD7BFE50" box="[612,749,440,463]" degrees="130" direction="east" minutes="50.95" orientation="longitude" pageId="31" pageNumber="290" precision="9" value="130.84917">130°50.95'E</geoCoordinate>
(Darwin Harbour)
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,
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depth unknown, 3,
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.
</paragraph>
<paragraph id="182A31278771595EFF88BFADFD36FB05" blockId="31.[121,749,509,1413]" pageId="31" pageNumber="290">
<emphasis id="2AE1ED358771595EFF88BFADFF68FD8A" bold="true" box="[121,254,509,533]" pageId="31" pageNumber="290">Description</emphasis>
. Corallum conical and fairly slender (H:D = 1.621.88), with a circular calice and blunt base.
<typeStatus id="C72E8F858771595EFD79BC4BFD7AFDAD" box="[648,748,539,562]" pageId="31" pageNumber="290" type="holotype">Holotype</typeStatus>
(and largest specimen)
<quantity id="DF6D9CC28771595EFE8FBC68FE77FDD0" box="[382,481,568,591]" metricMagnitude="-3" metricUnit="m" metricValue="2.76" pageId="31" pageNumber="290" unit="mm" value="2.76">2.76 mm</quantity>
in CD and
<quantity id="DF6D9CC28771595EFD9BBC68FD58FDD0" box="[618,718,568,591]" metricMagnitude="-3" metricUnit="m" metricValue="4.55" pageId="31" pageNumber="290" unit="mm" value="4.55">4.55 mm</quantity>
in height. C12 extend from base to calice; C3 originate independently about
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above the base, the C4 about
<quantity id="DF6D9CC28771595EFF88BCC0FF5AFD38" box="[121,204,656,679]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="31" pageNumber="290" unit="mm" value="1.7">1.7 mm</quantity>
above the base. Proximal to the origin of each C3 is a short costal ridge about
<quantity id="DF6D9CC28771595EFE54BCFEFDD7FD5A" box="[421,577,686,709]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" metricValueMax="4.5" metricValueMin="3.5" pageId="31" pageNumber="290" unit="mm" value="0.4" valueMax="0.45" valueMin="0.35">0.350.45 mm</quantity>
in length that is initially aligned with the C3 but distally is curved outward toward its adjacent S1, terminating in the intercostal groove that will be aligned with the future C4, altogether resulting in a zone of 12 such short costal ridges at a height of 1.0
<quantity id="DF6D9CC28771595EFF88BD10FF58FCC8" box="[121,206,832,855]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="31" pageNumber="290" unit="mm" value="1.4">1.4 mm</quantity>
above the base (
<figureCitation id="80AE2DA28771595EFE74BD10FE4FFCC8" box="[389,473,832,855]" captionStart="Fig" captionStartId="32.[166,202,1793,1814]" captionTargetBox="[166,1467,151,1776]" captionTargetId="figure-6@32.[166,1467,151,1776]" captionTargetPageId="32" captionText="Fig. 6. (AC), Conocyathus formosus (A, C, paratype, USNM 1008831, CD=2.5 mm; B, holotype, CD=2.76 mm): lateral, oblique calicular, and lateral basal views, the latter showing the short, disjunct costae, ×15, ×16.5, ×9.6 respectively. (D, E), Endopachys australiae (=Cyathotrochus pileus), holotype, Macleay Museum, lateral and oblique calicular views, both ×3.5 (GCD=12.4 mm). (F J), Dunocyathus wallaceae (F, G, I, holotype, AM G16744, GCD=4.4 mm; (H, J), paratype, USNM 1008835, intercostal width = 0.23 mm): (F, G), lateral and stereo calicular views, both ×10.8; (H), spiny base, ×21; (I), palar and columellar region, ×21; (J), detail of thecal edge showing alternation of septa and costae, ×44." pageId="31" pageNumber="290">Fig. 6C</figureCitation>
). In upper corallum C13 equal in width (about 65 µm), whereas C4 are about half this width, but all costae equal in height and exsertness. But in the basal part of the corallum, below the origin of the C4, the C1 3 usually bear thin, continuous lateral ridges that project into the intercostal groove nearly obscuring it and essentially doubling the width of these costae. Intercostal grooves about 75 µm in width near calicular edge and bridged by regularly spaced bars, each bar 6070 µm in width, delimiting rather deep, elliptical pits that are up to 100×70 µm in diameter, the greater diameter aligned with the groove. Approximately 20 pits occur in each intercostal groove adjacent to a C1.
</paragraph>
<paragraph id="182A31278771595EFF66BAF0FB22FE27" blockId="31.[121,749,509,1413]" lastBlockId="31.[794,1422,153,440]" pageId="31" pageNumber="290">
<collectingRegion id="DA51FFC58771595EFF66BAF0FF45FB28" box="[151,211,1184,1207]" country="Spain" name="Ceuta" pageId="31" pageNumber="290">Septa</collectingRegion>
hexamerally arranged in 3 complete cycles (24 septa) having the septal formula: S1&gt;S2&gt;S3. S1 highly exsert (up to
<quantity id="DF6D9CC28771595EFEF6BA8BFEC8FB6D" box="[263,350,1243,1266]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="31" pageNumber="290" unit="mm" value="0.3">0.3 mm</quantity>
), with extremely sinuous axial edges (
<figureCitation id="80AE2DA28771595EFF70BAA8FF47FA90" box="[129,209,1272,1295]" captionStart="Fig" captionStartId="30.[166,202,1793,1814]" captionTargetBox="[166,1467,151,1776]" captionTargetId="figure-6@30.[166,1467,151,1776]" captionTargetPageId="30" captionText="Fig. 5. (A, B), Trochocyathus wellsi, holotype, AM G16704, stereo calicular and lateral views, ×6.4, ×3.8, respectively (GCD=7.8 mm). (C, D), Vaughanella oreophila, Franklin 06/88/x, AM G16707, calicular and lateral views, both ×3.9 (CD=13.1 mm); (E), Alatotrochus rubescens, USNM 1008301 (SEM 996), juvenile corallum showing pronounced edge and basal crests, ×11.8 (GCD=3.6 mm). (FH), Conocyathus formosus, holotype,AM G16743: (F), inner view of septa, ×35; (G), stereo calicular view, ×19.1; (H), lateral view, ×17.1 (CD=2.76 mm)." pageId="31" pageNumber="290">Fig. 5F</figureCitation>
) that reach almost to the cental palar structure; S1 about 60 µm thick at calicular edge. S2 equally exsert, about 90% width of an S1, also having quite sinuous axial edges. S3 less exsert (
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), about 75% the width of an S1, and have sinuous axial edges that fuse with the axial edges of their adjacent S2 deep in the fossa, not easily visible in an intact specimen. Although C4 are well developed, there is no trace of S4. In fact, the region that would correspond to an S4 is slightly grooved internally (
<figureCitation id="80AE2DA28771595EFB34BEA1FA8EFE97" box="[1221,1304,241,264]" captionStart="Fig" captionStartId="30.[166,202,1793,1814]" captionTargetBox="[166,1467,151,1776]" captionTargetId="figure-6@30.[166,1467,151,1776]" captionTargetPageId="30" captionText="Fig. 5. (A, B), Trochocyathus wellsi, holotype, AM G16704, stereo calicular and lateral views, ×6.4, ×3.8, respectively (GCD=7.8 mm). (C, D), Vaughanella oreophila, Franklin 06/88/x, AM G16707, calicular and lateral views, both ×3.9 (CD=13.1 mm); (E), Alatotrochus rubescens, USNM 1008301 (SEM 996), juvenile corallum showing pronounced edge and basal crests, ×11.8 (GCD=3.6 mm). (FH), Conocyathus formosus, holotype,AM G16743: (F), inner view of septa, ×35; (G), stereo calicular view, ×19.1; (H), lateral view, ×17.1 (CD=2.76 mm)." pageId="31" pageNumber="290">Fig. 5F</figureCitation>
). All septa bear tall (up to 50 µm), rounded granules on their faces arranged in rows in the crests of the septal undulations. Centre of fossa occupied by 6 robust, lamellar P2, each about
<quantity id="DF6D9CC28771595EFCEBBF37FCEEFEE1" box="[794,888,359,382]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="31" pageNumber="290" unit="mm" value="0.25">0.25 mm</quantity>
in width, the axial edges of the 6 P2 fused together and altogether forming a single robust axial structure that rises to the level of the calicular edge.
</paragraph>
</subSubSection>
<subSubSection id="508F62AC8771595EFCEBBF99FAB7FDEB" pageId="31" pageNumber="290" type="discussion">
<paragraph id="182A31278771595EFCEBBF99FAB7FDEB" blockId="31.[794,1421,457,628]" pageId="31" pageNumber="290">
<emphasis id="2AE1ED358771595EFCEBBF99FC15FE7E" bold="true" box="[794,899,457,481]" pageId="31" pageNumber="290">Remarks</emphasis>
. As seen in
<tableCitation id="5517049C8771595EFBF3BF9AFBC4FE7E" box="[1026,1106,458,481]" captionStart="Table 5" captionStartId="31.[121,178,1500,1521]" captionText="Table 5. Distinguishing characteristics of the three Recent species of Conocyathus dOrbigny, 1849." pageId="31" pageNumber="290">Table 5</tableCitation>
,
<taxonomicName id="DF954AA48771595EFBACBF9AFB49FE7E" authorityName="Cairns" authorityYear="2004" box="[1117,1247,458,481]" class="Anthozoa" family="Turbinoliidae" genus="Conocyathus" kingdom="Animalia" order="Scleractinia" pageId="31" pageNumber="290" phylum="Cnidaria" rank="species" species="formosus">
<emphasis id="2AE1ED358771595EFBACBF9AFB49FE7E" box="[1117,1247,458,481]" italics="true" pageId="31" pageNumber="290">C. formosus</emphasis>
</taxonomicName>
is distinguished from the two other Recent species in the genus by having flanged costae; tall, but slender C4; sinuous septal axial edges; and pali that rise to the edge of the calice.
<collectingRegion id="DA51FFC58771595EFAD2BC72FADDFDA6" box="[1315,1355,546,569]" country="Maldives" name="Uthuru" pageId="31" pageNumber="290">Not</collectingRegion>
noted in the table is the peculiarity that most specimens bear 12 short, disjunct C3 near the base of the corallum.
</paragraph>
</subSubSection>
</treatment>
</document>