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<document id="0452F9C21351431B036041DDAE2933A4" ID-CLB-Dataset="46798" ID-DOI="10.11646/zootaxa.4078.1.22" ID-GBIF-Dataset="f71df488-ad28-4308-aed3-27540ae6ab4e" ID-ISSN="1175-5326" ID-Zenodo-Dep="255899" ID-ZooBank="C45311A2-E7EA-448F-9F38-79E145E53674" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460361558807" checkinUser="plazi" docAuthor="Grossi, Paschoal C., Santos, Mariana D. &amp; Almeida, Lúcia M." docDate="2016" docId="D11887F6C451FFB3C6D082D085E0F86E" docLanguage="en" docName="zootaxa.4078.1.22.pdf" docOrigin="Zootaxa 4078 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cyclocephala machadoi Grossi, Santos &amp; Almeida, 2016, sp. nov." docType="treatment" docVersion="8" lastPageNumber="246" masterDocId="2D21FF8EC450FFB2C64786128148FF9B" masterDocTitle="Two new species of Cyclocephala (Coleoptera: Scarabaeoidea: Melolonthidae) from Minas Gerais State, Brazil" masterLastPageNumber="251" masterPageNumber="245" pageNumber="246" updateTime="1698294628747" updateUser="ExternalLinkService">
<mods:mods id="62252646D91C4F233CD5D2F8EF199CBF" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="39EBCFF36CC0B6F4DA8404DF2E289ECE">Two new species of Cyclocephala (Coleoptera: Scarabaeoidea: Melolonthidae) from Minas Gerais State, Brazil</mods:title>
</mods:titleInfo>
<mods:name id="47D32B08F7F59412B46A6E5CCFCAA46D" type="personal">
<mods:role id="D0F9D32B90824846337CD0344BDABB4A">
<mods:roleTerm id="F78CE73985DC7185271FF6AC12E8AF72">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="06CE1FB4401890225F6398D92D578923">Grossi, Paschoal C.</mods:namePart>
</mods:name>
<mods:name id="28F69F4D7C8B5A62BFF3B300C0714F6E" type="personal">
<mods:role id="CAFB2D122C056926C6DC18D69F27F96E">
<mods:roleTerm id="757B95AE3A9C18492A2A3FFE830B30DD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B379893BDC21678CD4961F81D1405F0F">Santos, Mariana D.</mods:namePart>
</mods:name>
<mods:name id="58BAAB7A73B5A34396B62531BE7ADB7E" type="personal">
<mods:role id="1AD9B48DD0219E175779AE2B3C4EF98C">
<mods:roleTerm id="F980707641E24F816BD123DB595F2116">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="634224010309BCEA02E3DB52A5311400">Almeida, Lúcia M.</mods:namePart>
</mods:name>
<mods:typeOfResource id="5E105DCE84642D83411E18C1B7AD00D8">text</mods:typeOfResource>
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<mods:title id="018D8B84AB93FD0093191DBDD127BDD6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="39F9B8E77C7E57E9EB02BA16DC8ABB9F">
<mods:date id="A45014562C406545C03595D6451061E3">2016</mods:date>
<mods:detail id="5464433A8047E994B7821861B9836BB6" type="volume">
<mods:number id="99AA231E8D8186B9C983B615C2D68A22">4078</mods:number>
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<mods:detail id="20A0DFDB408EE897D461A962A08FE761" type="issue">
<mods:number id="CDBD268B8C2202E4C70EA8D0DF91F2CE">1</mods:number>
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<mods:classification id="666723C2F72F679AC98849ACADCF3398">journal article</mods:classification>
<mods:identifier id="4BDCA765034DE9F8C351025D8F2E913C" type="CLB-Dataset">46798</mods:identifier>
<mods:identifier id="C648A51FD70CDB690B885C039BEC7559" type="DOI">10.11646/zootaxa.4078.1.22</mods:identifier>
<mods:identifier id="C83B5F6418CDF56C31A3DFBF1FD1E11E" type="GBIF-Dataset">f71df488-ad28-4308-aed3-27540ae6ab4e</mods:identifier>
<mods:identifier id="4091A23B0947874278B6776B2F9E801C" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="2F4CA314E83DEEFB57EC9E45AAB4F0E2" type="Zenodo-Dep">255899</mods:identifier>
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<treatment id="D11887F6C451FFB3C6D082D085E0F86E" ID-DOI="http://doi.org/10.5281/zenodo.5659768" ID-GBIF-Taxon="119409862" ID-Zenodo-Dep="5659768" LSID="urn:lsid:plazi:treatment:D11887F6C451FFB3C6D082D085E0F86E" httpUri="http://treatment.plazi.org/id/D11887F6C451FFB3C6D082D085E0F86E" lastPageNumber="246" pageId="1" pageNumber="246">
<subSubSection id="11AB656BC451FFB3C6D082D0800CFB65" pageId="1" pageNumber="246" type="nomenclature">
<paragraph id="590E36E0C451FFB3C6D082D08355FB46" blockId="1.[151,541,1218,1278]" box="[151,541,1218,1245]" pageId="1" pageNumber="246">
<heading id="0246818CC451FFB3C6D082D08355FB46" bold="true" box="[151,541,1218,1245]" fontSize="11" level="1" pageId="1" pageNumber="246" reason="1">
<emphasis id="6BC5EAF2C451FFB3C6D082D08355FB46" bold="true" box="[151,541,1218,1245]" pageId="1" pageNumber="246">
<taxonomicName id="9EB14D63C451FFB3C6D082D080F0FB47" box="[151,440,1218,1244]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="246" phylum="Arthropoda" rank="species" species="machadoi" status="sp. nov.">
<emphasis id="6BC5EAF2C451FFB3C6D082D080F0FB47" bold="true" box="[151,440,1218,1244]" italics="true" pageId="1" pageNumber="246">Cyclocephala machadoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="70F65789C451FFB3C7F882D18355FB46" box="[447,541,1219,1245]" pageId="1" pageNumber="246" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="590E36E0C451FFB3C6D082F7800CFB65" blockId="1.[151,541,1218,1278]" box="[151,324,1253,1278]" pageId="1" pageNumber="246">
(
<figureCitation id="C18A2A65C451FFB3C6D982F7804CFB65" box="[158,260,1253,1278]" captionStart="FIGURE 1" captionStartId="2.[151,250,1906,1928]" captionTargetBox="[222,1352,203,1875]" captionTargetId="figure@2.[212,1374,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Males of Cyclocephala. A – F. holotype male of C. machadoi sp. nov.; G. C. everardoi sp. nov. holotype male; A. dorsal habitus; B. head, pronotum, and protibiae; C. antennal lamellae; D. anteriorprotarsus; E. parameres in caudal view; F. anterior margin of prosternum (arrows indicating the anterior projected margin); G. anterior margin of prosternum (arrows indicate the projected margin). Scale bars = 1 mm." httpUri="https://zenodo.org/record/255900/files/figure.png" pageId="1" pageNumber="246">Figures 1</figureCitation>
A–F)
</paragraph>
</subSubSection>
<subSubSection id="11AB656BC451FFB3C6D0833F818EF837" pageId="1" pageNumber="246" type="materials_examined">
<paragraph id="590E36E0C451FFB3C6D0833F85AEFAF7" blockId="1.[151,1437,1325,2037]" pageId="1" pageNumber="246">
<emphasis id="6BC5EAF2C451FFB3C6D0833F80C9FADD" bold="true" box="[151,385,1325,1350]" pageId="1" pageNumber="246">Material examined.</emphasis>
<typeStatus id="860A8842C451FFB3C7CD833F80BAFADD" box="[394,498,1325,1350]" pageId="1" pageNumber="246" type="holotype">Holotype</typeStatus>
male labeled: “
<collectingCountry id="21A67670C451FFB3C4E3833F83A3FADD" box="[676,747,1325,1350]" name="Brazil" pageId="1" pageNumber="246">Brasil</collectingCountry>
, MG, São Gonçalo/do Rio Preto, P.E. do/Rio Preto, Porteira, Posto (4)/
<date id="2D0F1020C451FFB3C742834680CDFAF6" box="[261,389,1364,1389]" pageId="1" pageNumber="246" value="2011-12-02">
02.xii.
<quantity id="9E499B05C451FFB3C70D834680CDFAF6" box="[330,389,1364,1389]" metricMagnitude="6" metricUnit="m" metricValue="2.011847" pageId="1" pageNumber="246" unit="m" value="2011847.0">2011</quantity>
</date>
, 847m, black/ligth, –
<geoCoordinate id="3C855027C451FFB3C43483468382FAF7" box="[627,714,1360,1388]" direction="south" orientation="latitude" pageId="1" pageNumber="246" precision="555" value="-18.09">18.09o S</geoCoordinate>
, –
<geoCoordinate id="3C855027C451FFB3C4A28346820EFAF6" box="[741,838,1360,1389]" direction="west" orientation="longitude" pageId="1" pageNumber="246" precision="555" value="-43.34">43.34o W</geoCoordinate>
/Oliveira &amp; Ferreira legs.” (
<collectionCode id="3FA0AE25C451FFB3C23B83468592FAF7" box="[1148,1242,1364,1388]" pageId="1" pageNumber="246">CERPE</collectionCode>
).
</paragraph>
<paragraph id="590E36E0C451FFB3C680836A818EF837" blockId="1.[151,1437,1325,2037]" pageId="1" pageNumber="246">
<emphasis id="6BC5EAF2C451FFB3C680836A8033FA0A" bold="true" box="[199,379,1400,1425]" pageId="1" pageNumber="246">
<typeStatus id="860A8842C451FFB3C680836A807CFA0A" box="[199,308,1400,1425]" pageId="1" pageNumber="246" type="holotype">Holotype</typeStatus>
male
</emphasis>
.
<emphasis id="6BC5EAF2C451FFB3C7D7836A836EFA0A" box="[400,550,1400,1425]" italics="true" pageId="1" pageNumber="246">Total length:</emphasis>
9.0 mm.
<emphasis id="6BC5EAF2C451FFB3C4E3836A82D7FA0A" box="[676,927,1400,1425]" italics="true" pageId="1" pageNumber="246">Total pronotal width:</emphasis>
<quantity id="9E499B05C451FFB3C5EA836A8553FA0B" box="[941,1051,1400,1425]" metricMagnitude="-3" metricUnit="m" metricValue="3.61" pageId="1" pageNumber="246" unit="mm" value="3.61">3.61 mm</quantity>
.
<emphasis id="6BC5EAF2C451FFB3C26B836A8527FA0A" box="[1068,1135,1400,1425]" italics="true" pageId="1" pageNumber="246">Body:</emphasis>
Dorsal surface glabrous, ventrally moderately setose, setae distinctly elongate. Color almost completely testaceous, but head black except clypeus, antennae light pale whitish yellow, and pronotum and legs slightly darker (
<figureCitation id="C18A2A65C451FFB3C27883D28536FA43" box="[1087,1150,1472,1497]" captionStart="FIGURE 1" captionStartId="2.[151,250,1906,1928]" captionTargetBox="[222,1352,203,1875]" captionTargetId="figure@2.[212,1374,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Males of Cyclocephala. A – F. holotype male of C. machadoi sp. nov.; G. C. everardoi sp. nov. holotype male; A. dorsal habitus; B. head, pronotum, and protibiae; C. antennal lamellae; D. anteriorprotarsus; E. parameres in caudal view; F. anterior margin of prosternum (arrows indicating the anterior projected margin); G. anterior margin of prosternum (arrows indicate the projected margin). Scale bars = 1 mm." httpUri="https://zenodo.org/record/255900/files/figure.png" pageId="1" pageNumber="246">Fig. 1</figureCitation>
A).
<emphasis id="6BC5EAF2C451FFB3C2EB83D285BAFA42" box="[1196,1266,1472,1497]" italics="true" pageId="1" pageNumber="246">Head:</emphasis>
Dorsal surface finely punctate, more punctate on frons; clypeus almost impunctate, sides parallel at base, then weakly convergent to trapezoidal, reflexed apex (
<figureCitation id="C18A2A65C451FFB3C7BA801A830EF9BB" box="[509,582,1544,1569]" captionStart="FIGURE 1" captionStartId="2.[151,250,1906,1928]" captionTargetBox="[222,1352,203,1875]" captionTargetId="figure@2.[212,1374,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Males of Cyclocephala. A – F. holotype male of C. machadoi sp. nov.; G. C. everardoi sp. nov. holotype male; A. dorsal habitus; B. head, pronotum, and protibiae; C. antennal lamellae; D. anteriorprotarsus; E. parameres in caudal view; F. anterior margin of prosternum (arrows indicating the anterior projected margin); G. anterior margin of prosternum (arrows indicate the projected margin). Scale bars = 1 mm." httpUri="https://zenodo.org/record/255900/files/figure.png" pageId="1" pageNumber="246">Figs 1</figureCitation>
A–B). Frontoclypeal suture obsolete. Antennae with lamellae extremely developed, moderately and densely punctate (
<figureCitation id="C18A2A65C451FFB3C4E7803E83A9F9DF" box="[672,737,1580,1605]" captionStart="FIGURE 1" captionStartId="2.[151,250,1906,1928]" captionTargetBox="[222,1352,203,1875]" captionTargetId="figure@2.[212,1374,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Males of Cyclocephala. A – F. holotype male of C. machadoi sp. nov.; G. C. everardoi sp. nov. holotype male; A. dorsal habitus; B. head, pronotum, and protibiae; C. antennal lamellae; D. anteriorprotarsus; E. parameres in caudal view; F. anterior margin of prosternum (arrows indicating the anterior projected margin); G. anterior margin of prosternum (arrows indicate the projected margin). Scale bars = 1 mm." httpUri="https://zenodo.org/record/255900/files/figure.png" pageId="1" pageNumber="246">Fig. 1</figureCitation>
C); first lamella externally with longitudinal row of scattered setae.
<emphasis id="6BC5EAF2C451FFB3C69980428021F9F2" box="[222,361,1616,1641]" italics="true" pageId="1" pageNumber="246">Mouthparts:</emphasis>
Mandibles slender, apices upturned and weakly curved externally; last maxillary palpomere as long as preceding 2 together.
<emphasis id="6BC5EAF2C451FFB3C7B780678320F917" box="[496,616,1653,1676]" italics="true" pageId="1" pageNumber="246">Pronotum:</emphasis>
Surface moderately and uniformly punctate, smooth along center line, posterior margin with a weak bead at middle. Scutellum totally punctate, punctures moderately large.
<emphasis id="6BC5EAF2C451FFB3C30B808A84D4F92A" box="[1356,1436,1688,1713]" italics="true" pageId="1" pageNumber="246">Elytra:</emphasis>
Surface distinctly punctate; striae well defined but weakly marked, punctures fine.
<emphasis id="6BC5EAF2C451FFB3C20380AF85CBF94F" box="[1092,1155,1725,1748]" italics="true" pageId="1" pageNumber="246">Legs:</emphasis>
Protibiae tridentate with basal tooth nearly obsolete (
<figureCitation id="C18A2A65C451FFB3C79580F2835DF963" box="[466,533,1760,1785]" captionStart="FIGURE 1" captionStartId="2.[151,250,1906,1928]" captionTargetBox="[222,1352,203,1875]" captionTargetId="figure@2.[212,1374,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Males of Cyclocephala. A – F. holotype male of C. machadoi sp. nov.; G. C. everardoi sp. nov. holotype male; A. dorsal habitus; B. head, pronotum, and protibiae; C. antennal lamellae; D. anteriorprotarsus; E. parameres in caudal view; F. anterior margin of prosternum (arrows indicating the anterior projected margin); G. anterior margin of prosternum (arrows indicate the projected margin). Scale bars = 1 mm." httpUri="https://zenodo.org/record/255900/files/figure.png" pageId="1" pageNumber="246">Figs 1</figureCitation>
A–B); inner claw widely bifid, forming a right angle (
<figureCitation id="C18A2A65C451FFB3C23080F285FEF963" box="[1143,1206,1760,1785]" captionStart="FIGURE 1" captionStartId="2.[151,250,1906,1928]" captionTargetBox="[222,1352,203,1875]" captionTargetId="figure@2.[212,1374,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Males of Cyclocephala. A – F. holotype male of C. machadoi sp. nov.; G. C. everardoi sp. nov. holotype male; A. dorsal habitus; B. head, pronotum, and protibiae; C. antennal lamellae; D. anteriorprotarsus; E. parameres in caudal view; F. anterior margin of prosternum (arrows indicating the anterior projected margin); G. anterior margin of prosternum (arrows indicate the projected margin). Scale bars = 1 mm." httpUri="https://zenodo.org/record/255900/files/figure.png" pageId="1" pageNumber="246">Fig. 1</figureCitation>
D).
<emphasis id="6BC5EAF2C451FFB3C2A280F3847AF963" box="[1253,1330,1761,1784]" italics="true" pageId="1" pageNumber="246">Venter:</emphasis>
Anterior prosternal margin sinuous and slightly crenulated, distinctly trilobed at middle, with middle lobe more prominent and conspicuous than lateral ones (
<figureCitation id="C18A2A65C451FFB3C47E813A8336F8DB" box="[569,638,1832,1857]" captionStart="FIGURE 1" captionStartId="2.[151,250,1906,1928]" captionTargetBox="[222,1352,203,1875]" captionTargetId="figure@2.[212,1374,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Males of Cyclocephala. A – F. holotype male of C. machadoi sp. nov.; G. C. everardoi sp. nov. holotype male; A. dorsal habitus; B. head, pronotum, and protibiae; C. antennal lamellae; D. anteriorprotarsus; E. parameres in caudal view; F. anterior margin of prosternum (arrows indicating the anterior projected margin); G. anterior margin of prosternum (arrows indicate the projected margin). Scale bars = 1 mm." httpUri="https://zenodo.org/record/255900/files/figure.png" pageId="1" pageNumber="246">Fig. 1</figureCitation>
F). Prosternal process high, but clearly not exceeding coxal height.
<emphasis id="6BC5EAF2C451FFB3C6D0815E8042F8FE" box="[151,266,1868,1893]" italics="true" pageId="1" pageNumber="246">Genitalia:</emphasis>
Parameres symmetrical, moderately elongated, with 2 lateral projections, basal projection larger, middle projection directed laterally; apex not diverging internally, parallel; subapex with small tooth; apex rounded (
<figureCitation id="C18A2A65C451FFB3C328816281EAF837" captionStart="FIGURE 1" captionStartId="2.[151,250,1906,1928]" captionTargetBox="[222,1352,203,1875]" captionTargetId="figure@2.[212,1374,195,1881]" captionTargetPageId="2" captionText="FIGURE 1. Males of Cyclocephala. A – F. holotype male of C. machadoi sp. nov.; G. C. everardoi sp. nov. holotype male; A. dorsal habitus; B. head, pronotum, and protibiae; C. antennal lamellae; D. anteriorprotarsus; E. parameres in caudal view; F. anterior margin of prosternum (arrows indicating the anterior projected margin); G. anterior margin of prosternum (arrows indicate the projected margin). Scale bars = 1 mm." httpUri="https://zenodo.org/record/255900/files/figure.png" pageId="1" pageNumber="246">Fig. 1</figureCitation>
E).
</paragraph>
</subSubSection>
<subSubSection id="11AB656BC451FFB3C68081AA85E0F86E" pageId="1" pageNumber="246" type="etymology">
<paragraph id="590E36E0C451FFB3C68081AA85E0F86E" blockId="1.[151,1437,1325,2037]" pageId="1" pageNumber="246">
<emphasis id="6BC5EAF2C451FFB3C68081AA800EF84A" bold="true" box="[199,326,1976,2001]" pageId="1" pageNumber="246">Etymology</emphasis>
. This species is named in honor of the entomologist and friend, Dr. Angelo Barbosa Monteiro Machado, expert in the taxonomy of dragonflies (Odonata), on occasion of his 80th birthday.
</paragraph>
</subSubSection>
</treatment>
</document>
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