treatments-xml/data/AD/4D/90/AD4D9054CD746A75F06641B6FD69FACD.xml

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<document id="22480DA3D9B4512BC72E2D841B87E683" ID-CLB-Dataset="22319" ID-DOI="10.5852/ejt.2013.41" ID-GBIF-Dataset="386010a0-6da4-4fa3-9d1a-4b3b5427cac5" ID-ISSN="2118-9773" ID-Zenodo-Dep="3814221" ID-ZooBank="A2F1B9ED-870A-466E-B35E-BD5DA782476E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588881240389" checkinUser="carolina" docAuthor="Kröger, Björn" docDate="2013" docId="AD4D9054CD746A75F06641B6FD69FACD" docLanguage="en" docName="ejt-2013-41.pdf.imf" docOrigin="European Journal of Taxonomy 41" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Beloitoceras sinuososeptatum" docType="treatment" docVersion="9" lastPageNumber="20" masterDocId="5174E82CCD646A66F260472AFFA5FFBD" masterDocTitle="The cephalopods of the Boda Limestone, Late Ordovician, of Dalarna, Sweden" masterLastPageNumber="110" masterPageNumber="1" pageNumber="17" updateTime="1698838944441" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="916E075F4A363D2A603F08DBE339578F">The cephalopods of the Boda Limestone, Late Ordovician, of Dalarna, Sweden</mods:title>
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<mods:title id="76339F55A8BF97D663BD26EE48E8DA8D">European Journal of Taxonomy</mods:title>
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<mods:date id="02DAC9B67AE433869F41E185087892DA">2013</mods:date>
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<treatment id="AD4D9054CD746A75F06641B6FD69FACD" ID-DOI="http://doi.org/10.5281/zenodo.3815104" ID-GBIF-Taxon="163977181" ID-Zenodo-Dep="3815104" LSID="urn:lsid:plazi:treatment:AD4D9054CD746A75F06641B6FD69FACD" httpUri="http://treatment.plazi.org/id/AD4D9054CD746A75F06641B6FD69FACD" lastPageId="19" lastPageNumber="20" pageId="16" pageNumber="17">
<subSubSection id="6DFE72C9CD746A76F06641B6FB8BF90A" box="[518,1070,1692,1719]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="255B2142CD746A76F06641B6FB8BF90A" blockId="16.[518,1070,1692,1754]" box="[518,1070,1692,1719]" pageId="16" pageNumber="17">
<heading id="7E13962ECD746A76F06641B6FB8BF90A" box="[518,1070,1692,1719]" centered="true" fontSize="11" level="2" pageId="16" pageNumber="17" reason="2">
<taxonomicName id="E2E45AC1CD746A76F06641B6FB8BF90A" authority="(Roemer, 1861)" baseAuthorityName="Roemer" baseAuthorityYear="1861" box="[518,1070,1692,1719]" class="Cephalopoda" family="Oncoceratidae" genus="Beloitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oncocerida" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="sinuososeptatum">
<emphasis id="1790FD50CD746A76F06641B6FCC8F90B" bold="true" box="[518,877,1692,1719]" italics="true" pageId="16" pageNumber="17">Beloitoceras sinuososeptatum</emphasis>
(
<bibRefCitation id="41755CB3CD746A76F11D41B6FB80F90A" author="Roemer C. F." box="[893,1061,1692,1719]" pageId="16" pageNumber="17" refId="ref49074" refString="Roemer C. F. 1861. Die fossile Fauna der silurischen Diluvial-Geschiebe von Sadewitz bei Oels in Nieder-Schlsien. Breslau." type="book" year="1861">Roemer, 1861</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="6DFE72C9CD746A76F0A74195FCC9F967" box="[711,876,1727,1754]" pageId="16" pageNumber="17" type="description">
<paragraph id="255B2142CD746A76F0A74195FCC9F967" blockId="16.[518,1070,1692,1754]" box="[711,876,1727,1754]" pageId="16" pageNumber="17">
<figureCitation id="BDDF3DC7CD746A76F0A74195FCAAF967" box="[711,783,1727,1754]" captionStart="Fig" captionStartId="17.[188,231,1717,1743]" captionTargetBox="[188,1396,541,1671]" captionTargetId="figure@17.[188,1399,541,1676]" captionTargetPageId="17" captionText="Fig. 9. Oncerida of the Boda Limestone. Note the high variability in conch curvature and angle of expansion in Beloitoceras. A-B. Beloitoceras sinuososeptatum (Roemer, 1861). A. PMU 24769. B. PMU 26638, Kallholn. C-D. Beloitoceras siljanense Frye, 1987. C. PMU 26644. D. PMU 26645, Kallholn. E-F. Apex of Beloitoceras sp., PMU 26646, Kallholn. E. Adapical view. F. Lateral view. G-H. Cyrtorizoceras thorslundi sp. nov., PMU 26657, Osmundsberget, mature body chamber. G. Lateral view. H. Adapical view. I. Beloitoceras siljanense Frye, 1987, PMU 26643, Kallholn, with major conch repair at adoral part. J. Cyrtorizoceras thorslundi sp. nov., PMU 26647, holotype, Kallholn, complete juvenile specimen. Scale bars: A-D, G-J = 10 mm; E-F = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3814241" httpUri="https://zenodo.org/record/3814241/files/figure.png" pageId="16" pageNumber="17">Figs 9</figureCitation>
A-B, 11
</paragraph>
</subSubSection>
<subSubSection id="6DFE72C9CD746A76F2DD402BFB53F819" pageId="16" pageNumber="17" type="reference_group">
<paragraph id="255B2142CD746A76F2DD402BFCD8F8A6" blockId="16.[189,923,1793,1854]" box="[189,893,1793,1819]" pageId="16" pageNumber="17">
<taxonomicName id="E2E45AC1CD746A76F2DD402BFD6DF8A6" authority="Roemer, 1861" authorityName="Roemer" authorityYear="1861" box="[189,712,1793,1819]" class="Liliopsida" family="Orchidaceae" genus="Orthoceras" kingdom="Plantae" order="Asparagales" pageId="16" pageNumber="17" phylum="Tracheophyta" rank="species" species="sinuoso-septatum">
<emphasis id="1790FD50CD746A76F2DD402BFDB2F8A6" box="[189,535,1793,1819]" italics="true" pageId="16" pageNumber="17">Orthoceras sinuoso-septatum</emphasis>
<bibRefCitation id="41755CB3CD746A76F07F402BFD6DF8A6" author="Roemer C. F." box="[543,712,1793,1819]" pageId="16" pageNumber="17" refId="ref49074" refString="Roemer C. F. 1861. Die fossile Fauna der silurischen Diluvial-Geschiebe von Sadewitz bei Oels in Nieder-Schlsien. Breslau." type="book" year="1861">Roemer, 1861</bibRefCitation>
</taxonomicName>
: pl. 6, fig. 3a-c.
</paragraph>
<paragraph id="255B2142CD746A76F2DD400EFC3EF883" blockId="16.[189,923,1793,1854]" box="[189,923,1828,1854]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="05F4066CCD746A76F2DD400EFC3EF883" box="[189,923,1828,1854]" pageId="16" pageNumber="17">
<treatmentCitation id="A4450753CD746A76F2DD400EFD7BF883" author="Strand T." box="[189,734,1828,1854]" page="76" pageId="16" pageNumber="17" year="1934">
<taxonomicName id="E2E45AC1CD746A76F2DD400EFD7BF883" authority="Strand, 1934: 76" authorityName="Strand" authorityPageNumber="76" authorityYear="1934" box="[189,734,1828,1854]" class="Cephalopoda" family="Oncoceratidae" genus="Beloitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oncocerida" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="heterocurvatum">
<emphasis id="1790FD50CD746A76F2DD400EFDB7F883" box="[189,530,1828,1854]" italics="true" pageId="16" pageNumber="17">Beloitoceras heterocurvatum</emphasis>
<bibRefCitation id="41755CB3CD746A76F079400EFD7BF883" author="Strand T." box="[537,734,1828,1854]" pageId="16" pageNumber="17" pagination="1 - 117" refId="ref49308" refString="Strand T. 1934. The Upper Ordovician Cephalopods of the Oslo Area. Norsk Geologiske Tidsskrift 14: 1 - 117." type="journal article" year="1934">Strand, 1934: 76</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. 10, figs 8-9.
</treatmentCitationGroup>
</paragraph>
<paragraph id="255B2142CD746A76F2DD404CFB3AF8C2" blockId="16.[189,1270,1893,1956]" box="[189,1183,1893,1920]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="05F4066CCD746A76F2DD404CFB3AF8C2" box="[189,1183,1893,1920]" pageId="16" pageNumber="17">
<taxonomicName id="E2E45AC1CD746A76F2DD404CFDF8F8C2" baseAuthorityName="Roemer" baseAuthorityYear="1861" box="[189,605,1894,1920]" class="Cephalopoda" family="Nautilidae" genus="Richardsonoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Nautilida" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="sinuososeptatum">
<emphasis id="1790FD50CD746A76F2DD404CFDF8F8C2" box="[189,605,1894,1920]" italics="true" pageId="16" pageNumber="17">Richardsonoceras sinuososeptatum</emphasis>
</taxonomicName>
<treatmentCitation id="A4450753CD746A76F01A404FFC97F83D" author="Dzik J." box="[634,818,1893,1920]" page="188" pageId="16" pageNumber="17" year="1984">
<bibRefCitation id="41755CB3CD746A76F01A404FFC97F83D" author="Dzik J." box="[634,818,1893,1920]" pageId="16" pageNumber="17" refId="ref47068" refString="Dzik J. 1984. Phylogeny of the Nautiloidea. Palaeontologia Polonica, Vol. 45, 203 pp." type="book" year="1984">Dzik 1984: 188</bibRefCitation>
</treatmentCitation>
, text-fig. 16a-b, pl. 8, fig. 1a-d.
</treatmentCitationGroup>
</paragraph>
<paragraph id="255B2142CD746A76F2DD40A3FB53F819" blockId="16.[189,1270,1893,1956]" box="[189,1270,1929,1956]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="05F4066CCD746A76F2DD40A3FB53F819" box="[189,1270,1929,1956]" pageId="16" pageNumber="17">
<taxonomicName id="E2E45AC1CD746A76F2DD40A3FDB7F81E" authorityName="Strand" authorityYear="1934" box="[189,530,1929,1955]" class="Cephalopoda" family="Oncoceratidae" genus="Beloitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oncocerida" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="heterocurvatum">
<emphasis id="1790FD50CD746A76F2DD40A3FDB7F81E" box="[189,530,1929,1955]" italics="true" pageId="16" pageNumber="17">Beloitoceras heterocurvatum</emphasis>
</taxonomicName>
<treatmentCitation id="A4450753CD746A76F04F40A3FD73F81E" author="Frye M. W." box="[559,726,1929,1955]" page="84" pageId="16" pageNumber="17" year="1987">
<bibRefCitation id="41755CB3CD746A76F04F40A3FD73F81E" author="Frye M. W." box="[559,726,1929,1955]" pageId="16" pageNumber="17" pagination="83 - 99" refId="ref47793" refString="Frye M. W. 1987. Upper Ordovocian (Harjuan) oncoceratid nautiloids from Boda Limestone, Siljan District, Sweden. Geologiska Foreningens i Stockholm Forhandlingar 109: 83 - 99. http: // dx. doi. org / 10.1080 / 11035898709454748" type="journal article" year="1987">Frye 1987: 84</bibRefCitation>
</treatmentCitation>
, figs 3a-d. ―
<treatmentCitation id="A4450753CD746A76F11B40A3FB25F81E" author="Kroger B. &amp; Ebbestad J. O. R. &amp; Hogstrom A. E. S. &amp; Frisk A. M." box="[891,1152,1929,1955]" page="38" pageId="16" pageNumber="17" year="2011">
<bibRefCitation id="41755CB3CD746A76F11B40A3FB25F81E" author="Kroger B. &amp; Ebbestad J. O. R. &amp; Hogstrom A. E. S. &amp; Frisk A. M." box="[891,1152,1929,1955]" pageId="16" pageNumber="17" pagination="35 - 53" refId="ref48453" refString="Kroger B., Ebbestad J. O. R., Hogstrom A. E. S. &amp; Frisk A. M. 2011. Mass concentration of Hirnantian cephalopods from the Siljan District, Sweden; taxonomy, palaeoecology and palaeobiogeographic relationship. Fossil Record 14: 35 - 53. http: // dx. doi. org / 10.1002 / mmng. 201000014" type="journal article" year="2011">
Kröger
<emphasis id="1790FD50CD746A76F1B540A0FBB5F81E" box="[981,1040,1929,1955]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
2011: 38
</bibRefCitation>
</treatmentCitation>
, fig. 5a-b.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="6DFE72C9CD756A74F2DD4621FB68FEF5" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="diagnosis">
<paragraph id="255B2142CD756A77F2DD4621FE0AFE98" blockId="17.[189,431,267,293]" box="[189,431,267,293]" pageId="17" pageNumber="18">
<heading id="7E13962ECD756A77F2DD4621FE0AFE98" bold="true" box="[189,431,267,293]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="1790FD50CD756A77F2DD4621FE0AFE98" bold="true" box="[189,431,267,293]" pageId="17" pageNumber="18">Emended diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="255B2142CD756A77F2DD4613FAD3FE5D" blockId="17.[189,1398,313,481]" pageId="17" pageNumber="18">
<taxonomicName id="E2E45AC1CD756A77F2DD4613FEF4FEEE" authorityName="Foerste" authorityYear="1924" box="[189,337,313,339]" class="Cephalopoda" family="Oncoceratidae" genus="Beloitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oncocerida" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">
<emphasis id="1790FD50CD756A77F2DD4613FEF4FEEE" box="[189,337,313,339]" italics="true" pageId="17" pageNumber="18">Beloitoceras</emphasis>
</taxonomicName>
with compressed cross section with keeled conch margins at concave and convex side of conch curvature throughout entire length; in longitudinal section antisiphuncular conch margin concave; angle of expansion of phragmocone ca. 1617°; depth of body chamber somewhat less than maximum conch height; conch surface ornamented with irregularly spaced rounded striae, which form a shallow sinus on prosiphuncular side and deep v-shaped sinus on antisiphuncular side; about six chambers
</paragraph>
<caption id="719B71CACD756A77F2DC419FFC0FF87B" ID-DOI="http://doi.org/10.5281/zenodo.3814241" ID-Zenodo-Dep="3814241" httpUri="https://zenodo.org/record/3814241/files/figure.png" pageId="17" pageNumber="18" startId="17.[188,231,1717,1743]" targetBox="[188,1396,541,1671]" targetPageId="17">
<paragraph id="255B2142CD756A77F2DC419FFC0FF87B" blockId="17.[188,1399,1717,1990]" pageId="17" pageNumber="18">
<emphasis id="1790FD50CD756A77F2DC419FFEABF972" bold="true" box="[188,270,1717,1743]" pageId="17" pageNumber="18">Fig. 9.</emphasis>
Oncerida of the Boda Limestone. Note the high variability in conch curvature and angle of expansion in
<taxonomicName id="E2E45AC1CD756A77F30941F2FE58F94F" authorityName="Foerste" authorityYear="1924" box="[361,509,1752,1778]" class="Cephalopoda" family="Oncoceratidae" genus="Beloitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oncocerida" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">
<emphasis id="1790FD50CD756A77F30941F2FE58F94F" box="[361,509,1752,1778]" italics="true" pageId="17" pageNumber="18">Beloitoceras</emphasis>
</taxonomicName>
.
<emphasis id="1790FD50CD756A77F07441F2FDE2F94F" bold="true" box="[532,583,1752,1779]" pageId="17" pageNumber="18">A -B</emphasis>
.
<taxonomicName id="E2E45AC1CD756A77F03D41F2FB3DF94E" authority="(Roemer, 1861)" baseAuthorityName="Roemer" baseAuthorityYear="1861" box="[605,1176,1752,1779]" class="Cephalopoda" family="Oncoceratidae" genus="Beloitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oncocerida" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="sinuososeptatum">
<emphasis id="1790FD50CD756A77F03D41F2FC60F94F" box="[605,965,1752,1778]" italics="true" pageId="17" pageNumber="18">Beloitoceras sinuososeptatum</emphasis>
(
<bibRefCitation id="41755CB3CD756A77F1BB41F2FB2AF94E" author="Roemer C. F." box="[987,1167,1752,1779]" pageId="17" pageNumber="18" refId="ref49074" refString="Roemer C. F. 1861. Die fossile Fauna der silurischen Diluvial-Geschiebe von Sadewitz bei Oels in Nieder-Schlsien. Breslau." type="book" year="1861">Roemer, 1861</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="1790FD50CD756A77F6CB41F2FB65F94F" bold="true" box="[1195,1216,1752,1778]" pageId="17" pageNumber="18">A</emphasis>
. PMU 24769.
<emphasis id="1790FD50CD756A77F2DC41D1FF75F8A8" bold="true" box="[188,208,1787,1813]" pageId="17" pageNumber="18">B</emphasis>
. PMU 26638, Kallholn.
<emphasis id="1790FD50CD756A77F39B41D1FD8AF8A8" bold="true" box="[507,559,1787,1814]" pageId="17" pageNumber="18">C -D</emphasis>
.
<taxonomicName id="E2E45AC1CD756A77F02041D1FC44F8A8" authority="Frye, 1987" authorityName="Frye" authorityYear="1987" box="[576,993,1787,1814]" class="Cephalopoda" family="Oncoceratidae" genus="Beloitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oncocerida" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="siljanense">
<emphasis id="1790FD50CD756A77F02041D1FCF7F8A8" box="[576,850,1787,1813]" italics="true" pageId="17" pageNumber="18">Beloitoceras siljanense</emphasis>
<bibRefCitation id="41755CB3CD756A77F13C41D6FC44F8A8" author="Frye M. W." box="[860,993,1787,1814]" pageId="17" pageNumber="18" pagination="83 - 99" refId="ref47793" refString="Frye M. W. 1987. Upper Ordovocian (Harjuan) oncoceratid nautiloids from Boda Limestone, Siljan District, Sweden. Geologiska Foreningens i Stockholm Forhandlingar 109: 83 - 99. http: // dx. doi. org / 10.1080 / 11035898709454748" type="journal article" year="1987">Frye, 1987</bibRefCitation>
</taxonomicName>
.
<emphasis id="1790FD50CD756A77F18E41D1FBA6F8A8" bold="true" box="[1006,1027,1787,1813]" pageId="17" pageNumber="18">C</emphasis>
. PMU 26644.
<emphasis id="1790FD50CD756A77F6D741D1FB69F8A8" bold="true" box="[1207,1228,1787,1813]" pageId="17" pageNumber="18">D</emphasis>
. PMU 26645, Kallholn.
<emphasis id="1790FD50CD756A77F3554035FEC0F884" bold="true" box="[309,357,1823,1849]" pageId="17" pageNumber="18">E -F</emphasis>
. Apex of
<taxonomicName id="E2E45AC1CD756A77F3834034FDD2F885" authorityName="Foerste" authorityYear="1924" box="[483,631,1822,1848]" class="Cephalopoda" family="Oncoceratidae" genus="Beloitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oncocerida" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">
<emphasis id="1790FD50CD756A77F3834034FDD2F885" box="[483,631,1822,1848]" italics="true" pageId="17" pageNumber="18">Beloitoceras</emphasis>
</taxonomicName>
sp., PMU 26646, Kallholn.
<emphasis id="1790FD50CD756A77F1B44035FC4DF884" bold="true" box="[980,1000,1823,1849]" pageId="17" pageNumber="18">E</emphasis>
. Adapical view.
<emphasis id="1790FD50CD756A77F6D64035FB6DF884" bold="true" box="[1206,1224,1823,1849]" pageId="17" pageNumber="18">F</emphasis>
. Lateral view.
<emphasis id="1790FD50CD756A77F2DC406BFF51F8E1" bold="true" box="[188,244,1857,1884]" pageId="17" pageNumber="18">G -H</emphasis>
.
<taxonomicName id="E2E45AC1CD756A77F3614068FD8AF8E1" authority="Kröger, 2013" authorityName="Kröger" authorityYear="2013" box="[257,559,1858,1884]" class="Cephalopoda" family="Oncoceratidae" genus="Cyrtorizoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oncocerida" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="thorslundi" status="sp. nov.">
<emphasis id="1790FD50CD756A77F3614068FD8AF8E1" box="[257,559,1858,1884]" italics="true" pageId="17" pageNumber="18">Cyrtorizoceras thorslundi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0CA3402BCD756A77F0544068FD2AF8E1" box="[564,655,1858,1884]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
, PMU 26657, Osmundsberget, mature body chamber.
<emphasis id="1790FD50CD756A77F760406BFAB2F8E6" bold="true" box="[1280,1303,1857,1883]" pageId="17" pageNumber="18">G</emphasis>
. Lateral view.
<emphasis id="1790FD50CD756A77F362404FFEBCF8C2" bold="true" box="[258,281,1893,1919]" pageId="17" pageNumber="18">H</emphasis>
. Adapical view.
<emphasis id="1790FD50CD756A77F3BB404FFE43F8C2" bold="true" box="[475,486,1893,1919]" pageId="17" pageNumber="18">I</emphasis>
.
<taxonomicName id="E2E45AC1CD756A77F394404FFC29F8C2" authority="Frye, 1987" authorityName="Frye" authorityYear="1987" box="[500,908,1893,1919]" class="Cephalopoda" family="Oncoceratidae" genus="Beloitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oncocerida" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="siljanense">
<emphasis id="1790FD50CD756A77F394404FFCA1F8C2" box="[500,772,1893,1919]" italics="true" pageId="17" pageNumber="18">Beloitoceras siljanense</emphasis>
<bibRefCitation id="41755CB3CD756A77F16B404FFC29F8C2" author="Frye M. W." box="[779,908,1893,1919]" pageId="17" pageNumber="18" pagination="83 - 99" refId="ref47793" refString="Frye M. W. 1987. Upper Ordovocian (Harjuan) oncoceratid nautiloids from Boda Limestone, Siljan District, Sweden. Geologiska Foreningens i Stockholm Forhandlingar 109: 83 - 99. http: // dx. doi. org / 10.1080 / 11035898709454748" type="journal article" year="1987">Frye, 1987</bibRefCitation>
</taxonomicName>
, PMU 26643, Kallholn, with major conch repair at adoral part.
<emphasis id="1790FD50CD756A77F3D740A2FE63F81F" bold="true" box="[439,454,1928,1954]" pageId="17" pageNumber="18">J</emphasis>
.
<taxonomicName id="E2E45AC1CD756A77F3B640A2FCADF81F" authority="Kröger, 2013" authorityName="Kröger" authorityYear="2013" box="[470,776,1928,1954]" class="Cephalopoda" family="Oncoceratidae" genus="Cyrtorizoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oncocerida" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="thorslundi" status="sp. nov.">
<emphasis id="1790FD50CD756A77F3B640A2FCADF81F" box="[470,776,1928,1954]" italics="true" pageId="17" pageNumber="18">Cyrtorizoceras thorslundi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="0CA3402BCD756A77F17040A3FCCBF81E" box="[784,878,1929,1955]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
, PMU 26647, holotype, Kallholn, complete juvenile specimen. Scale bars: A-D, G-J = 10 mm; E-F = 2 mm.
</paragraph>
</caption>
<paragraph id="255B2142CD766A74F2DD4621FB68FEF5" blockId="18.[189,1398,267,329]" pageId="18" pageNumber="19">similar to the height of the conch cross section; sutures with deep lateral lobes; siphuncle situated near ventral margin, segments ca. twice as high as wide, with slightly convex vertical outline.</paragraph>
</subSubSection>
<subSubSection id="6DFE72C9CD766A74F2DD465AFC1BFDF1" pageId="18" pageNumber="19" type="materials_examined">
<paragraph id="255B2142CD766A74F2DD465AFE5EFE37" blockId="18.[189,507,368,394]" box="[189,507,368,394]" pageId="18" pageNumber="19">
<heading id="7E13962ECD766A74F2DD465AFE5EFE37" bold="true" box="[189,507,368,394]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="1790FD50CD766A74F2DD465AFE5EFE37" bold="true" box="[189,507,368,394]" pageId="18" pageNumber="19">
<typeStatus id="FA5F9FE0CD766A74F2DD465AFF5FFE37" box="[189,250,368,394]" pageId="18" pageNumber="19">Type</typeStatus>
locality and horizon
</emphasis>
</heading>
</paragraph>
<paragraph id="255B2142CD766A74F2DD46B4FAA0FE05" blockId="18.[189,1285,414,441]" box="[189,1285,414,441]" pageId="18" pageNumber="19">
Zawidowice near Oleśnica,
<collectingCountry id="5DF361D2CD766A74F06446B4FDFDFE05" box="[516,600,414,440]" name="Poland" pageId="18" pageNumber="19">Poland</collectingCountry>
, erratic boulder of probably latest Katian age, Ordovician.
</paragraph>
<paragraph id="255B2142CD766A74F2DD46CAFE0FFE47" blockId="18.[189,426,480,506]" box="[189,426,480,506]" pageId="18" pageNumber="19">
<heading id="7E13962ECD766A74F2DD46CAFE0FFE47" bold="true" box="[189,426,480,506]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="1790FD50CD766A74F2DD46CAFE0FFE47" bold="true" box="[189,426,480,506]" pageId="18" pageNumber="19">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="255B2142CD766A74F2DD4524FC1BFDF1" blockId="18.[189,1399,526,588]" pageId="18" pageNumber="19">
Thirteen specimens in total:
<collectionCode id="43F5B987CD766A74F0434525FDC6FD94" box="[547,611,527,553]" country="Sweden" httpUri="http://grbio.org/cool/32ji-dbbk" name="Paleontological Museum of Uppsala" pageId="18" pageNumber="19">PMU</collectionCode>
2663326642, 24769, 24772, Kallholn,
<collectingRegion id="E720EFA0CD766A74F63E4524FB1AFD95" box="[1118,1215,526,552]" country="Sweden" name="Dalarna" pageId="18" pageNumber="19">Dalarna</collectingRegion>
; one specimen (
<collectionCode id="43F5B987CD766A74F2A54518FEA2FDF1" box="[197,263,562,588]" country="Sweden" httpUri="http://grbio.org/cool/32ji-dbbk" name="Paleontological Museum of Uppsala" pageId="18" pageNumber="19">PMU</collectionCode>
24770), Skålberget,
<collectingRegion id="E720EFA0CD766A74F39B4518FDFEFDF1" box="[507,603,562,588]" country="Sweden" name="Dalarna" pageId="18" pageNumber="19">Dalarna</collectingRegion>
, Boda Limestone, late Katian.
</paragraph>
</subSubSection>
<subSubSection id="6DFE72C9CD766A74F2DD4559FC3EFC44" pageId="18" pageNumber="19" type="description">
<paragraph id="255B2142CD766A74F2DD4559FEEBFD30" blockId="18.[189,334,627,653]" box="[189,334,627,653]" pageId="18" pageNumber="19">
<heading id="7E13962ECD766A74F2DD4559FEEBFD30" bold="true" box="[189,334,627,653]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="1790FD50CD766A74F2DD4559FEEBFD30" bold="true" box="[189,334,627,653]" pageId="18" pageNumber="19">Description</emphasis>
</heading>
</paragraph>
<paragraph id="255B2142CD766A74F2DD4588FC3EFC44" blockId="18.[189,1399,674,1017]" pageId="18" pageNumber="19">
The general features of the conch, the ornamentation and the details of the siphuncle were described by
<bibRefCitation id="41755CB3CD766A74F28745EFFE37FD62" author="Strand T." box="[231,402,709,735]" pageId="18" pageNumber="19" pagination="1 - 117" refId="ref49308" refString="Strand T. 1934. The Upper Ordovician Cephalopods of the Oslo Area. Norsk Geologiske Tidsskrift 14: 1 - 117." type="journal article" year="1934">Strand (1934)</bibRefCitation>
,
<bibRefCitation id="41755CB3CD766A74F3C345EFFD92FD62" author="Dzik J." box="[419,567,709,735]" pageId="18" pageNumber="19" refId="ref47068" refString="Dzik J. 1984. Phylogeny of the Nautiloidea. Palaeontologia Polonica, Vol. 45, 203 pp." type="book" year="1984">Dzik (1984)</bibRefCitation>
and
<bibRefCitation id="41755CB3CD766A74F01C45EFFCB4FD62" author="Frye M. W." box="[636,785,709,735]" pageId="18" pageNumber="19" pagination="83 - 99" refId="ref47793" refString="Frye M. W. 1987. Upper Ordovocian (Harjuan) oncoceratid nautiloids from Boda Limestone, Siljan District, Sweden. Geologiska Foreningens i Stockholm Forhandlingar 109: 83 - 99. http: // dx. doi. org / 10.1080 / 11035898709454748" type="journal article" year="1987">Frye (1987)</bibRefCitation>
. These descriptions were based on relatively few specimens. Here, 15 additional specimens are available for measurements and direct comparison. These specimens reveal a high variability in conch curvature and angle of expansion. The conch curvature varies from strongly curved to relatively widely curved (
<figureCitation id="BDDF3DC7CD766A74F12D4404FC37FCF4" box="[845,914,814,841]" captionStart="Fig" captionStartId="17.[188,231,1717,1743]" captionTargetBox="[188,1396,541,1671]" captionTargetId="figure@17.[188,1399,541,1676]" captionTargetPageId="17" captionText="Fig. 9. Oncerida of the Boda Limestone. Note the high variability in conch curvature and angle of expansion in Beloitoceras. A-B. Beloitoceras sinuososeptatum (Roemer, 1861). A. PMU 24769. B. PMU 26638, Kallholn. C-D. Beloitoceras siljanense Frye, 1987. C. PMU 26644. D. PMU 26645, Kallholn. E-F. Apex of Beloitoceras sp., PMU 26646, Kallholn. E. Adapical view. F. Lateral view. G-H. Cyrtorizoceras thorslundi sp. nov., PMU 26657, Osmundsberget, mature body chamber. G. Lateral view. H. Adapical view. I. Beloitoceras siljanense Frye, 1987, PMU 26643, Kallholn, with major conch repair at adoral part. J. Cyrtorizoceras thorslundi sp. nov., PMU 26647, holotype, Kallholn, complete juvenile specimen. Scale bars: A-D, G-J = 10 mm; E-F = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3814241" httpUri="https://zenodo.org/record/3814241/files/figure.png" pageId="18" pageNumber="19">Fig. 9</figureCitation>
A-B). The angle of expansion varies from 15° (1
<superScript id="D2918C0ACD766A74F368447AFEB6FCE3" attach="left" box="[264,275,848,862]" fontSize="6" pageId="18" pageNumber="19">st</superScript>
quantile) to, 18° (3
<superScript id="D2918C0ACD766A74F062447AFDB5FCE3" attach="left" box="[514,528,848,862]" fontSize="6" pageId="18" pageNumber="19">rd</superScript>
quantile) with a mean of 17° (15 measurements) in the specimens from the Boda Limestone. The conch cross section is always compressed with a typical keel at the anti-, and prosiphuncular side, but the rate of compression varies among the Boda Limestone specimens from 0.85 (1
<superScript id="D2918C0ACD766A74F2B54490FF45FC75" attach="left" box="[213,224,954,968]" fontSize="6" pageId="18" pageNumber="19">st</superScript>
quantile) to 0.92 (3
<superScript id="D2918C0ACD766A74F3A64490FE71FC75" attach="left" box="[454,468,954,968]" fontSize="6" pageId="18" pageNumber="19">rd</superScript>
quantile) with a mean of 0.87 (10 measurements). Within this range of variance no distinct groups of specimens can be distinguished (
<figureCitation id="BDDF3DC7CD766A74F15744F4FC28FC44" box="[823,909,990,1017]" captionStart="Fig" captionStartId="21.[189,232,1841,1867]" captionTargetBox="[201,1359,1426,1795]" captionTargetId="graphics@21.[284,756,1449,1701]" captionTargetPageId="21" captionText="Fig. 11. Morphological variation of Beloitoceras sinuososeptatum (Roemer, 1861). Circles: specimens of the Boda Limestone. Black dot: type specimen of Beloitoceras siljanense Frye, 1987. Black diamonds: types of Beloitoceras heterocurvatum Strand, 1934. Grey dot: type specimen of Beloitoceras landerense Foerste, 1935. A distinction between B. siljanense and B. sinuososeptatum is impossible based on the available material." figureDoi="http://doi.org/10.5281/zenodo.3814247" httpUri="https://zenodo.org/record/3814247/files/figure.png" pageId="18" pageNumber="19">Fig. 11</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="6DFE72C9CD766A75F2DD430AFACBFB2B" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="discussion">
<paragraph id="255B2142CD766A74F2DD430AFE8AFB87" blockId="18.[189,303,1056,1082]" box="[189,303,1056,1082]" pageId="18" pageNumber="19">
<heading id="7E13962ECD766A74F2DD430AFE8AFB87" bold="true" box="[189,303,1056,1082]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
<emphasis id="1790FD50CD766A74F2DD430AFE8AFB87" bold="true" box="[189,303,1056,1082]" pageId="18" pageNumber="19">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="255B2142CD766A74F2DD4365FAD2FB48" blockId="18.[189,1399,1102,1270]" pageId="18" pageNumber="19">
In earlier descriptions the variability of the conch shape of this species was not recognized. Additionally, the adult size of different specimen assigned to this species varies from
<quantity id="E21C8CA7CD766A74F6794358FB16FB31" box="[1049,1203,1138,1164]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" metricValueMax="3.8" metricValueMin="2.8" pageId="18" pageNumber="19" unit="mm" value="33.0" valueMax="38.0" valueMin="28.0">28 to 38 mm</quantity>
maximum body chamber height. Because only three specimens from the Boda Limestone possess an adult body chamber (specimen figured in
<bibRefCitation id="41755CB3CD766A74F3D34393FD8BFB6F" author="Frye M. W." box="[435,558,1208,1235]" pageId="18" pageNumber="19" pagination="83 - 99" refId="ref47793" refString="Frye M. W. 1987. Upper Ordovocian (Harjuan) oncoceratid nautiloids from Boda Limestone, Siljan District, Sweden. Geologiska Foreningens i Stockholm Forhandlingar 109: 83 - 99. http: // dx. doi. org / 10.1080 / 11035898709454748" type="journal article" year="1987">Frye 1987</bibRefCitation>
: fig. 3a, b; PMU 24769, 26634), it is impossible to evaluate whether this difference reflects interspecific variability, sexual dimorphism or the presence of more than one species.
</paragraph>
<paragraph id="255B2142CD766A74F2DD4237FBBCF92A" blockId="18.[189,1399,1309,1687]" pageId="18" pageNumber="19">
The type specimen from Vormsi/Pirgu Regional Stage limestones of the erratic boulders of northern
<collectingCountry id="5DF361D2CD766A74F2DD426AFEB4FAE7" box="[189,273,1344,1370]" name="Poland" pageId="18" pageNumber="19">Poland</collectingCountry>
, figured by
<bibRefCitation id="41755CB3CD766A74F3CA426AFD90FAE7" author="Dzik J." box="[426,565,1344,1370]" pageId="18" pageNumber="19" refId="ref47068" refString="Dzik J. 1984. Phylogeny of the Nautiloidea. Palaeontologia Polonica, Vol. 45, 203 pp." type="book" year="1984">Dzik (1984</bibRefCitation>
, pl. 8, fig. 1a-d), differs from the Boda Limestone specimens in its exceptionally low angle of expansion of only 10°. Moreover, the
<typeStatus id="FA5F9FE0CD766A74F1CD424EFBB7FAC3" box="[941,1042,1380,1406]" pageId="18" pageNumber="19" type="paratype">paratype</typeStatus>
figured by
<bibRefCitation id="41755CB3CD766A74F6F54249FA80FAC0" author="Dzik J." box="[1173,1317,1379,1405]" pageId="18" pageNumber="19" refId="ref47068" refString="Dzik J. 1984. Phylogeny of the Nautiloidea. Palaeontologia Polonica, Vol. 45, 203 pp." type="book" year="1984">Dzik (1984)</bibRefCitation>
, with a width/height ratio of 0.72, and the specimen from the Oslo area figured by
<bibRefCitation id="41755CB3CD766A74F67642ACFB14FA1D" author="Strand T." box="[1046,1201,1414,1440]" pageId="18" pageNumber="19" pagination="1 - 117" refId="ref49308" refString="Strand T. 1934. The Upper Ordovician Cephalopods of the Oslo Area. Norsk Geologiske Tidsskrift 14: 1 - 117." type="journal article" year="1934">Strand (1934</bibRefCitation>
: pl. 10, fig. 8) are considerably more compressed than the specimens from the Boda Limestone. Also, these two specimens represent the lower range of the angle of expansion for all the specimens measured (
<figureCitation id="BDDF3DC7CD766A74F6F942E7FB4AFA5A" box="[1177,1263,1485,1511]" captionStart="Fig" captionStartId="21.[189,232,1841,1867]" captionTargetBox="[201,1359,1426,1795]" captionTargetId="graphics@21.[284,756,1449,1701]" captionTargetPageId="21" captionText="Fig. 11. Morphological variation of Beloitoceras sinuososeptatum (Roemer, 1861). Circles: specimens of the Boda Limestone. Black dot: type specimen of Beloitoceras siljanense Frye, 1987. Black diamonds: types of Beloitoceras heterocurvatum Strand, 1934. Grey dot: type specimen of Beloitoceras landerense Foerste, 1935. A distinction between B. siljanense and B. sinuososeptatum is impossible based on the available material." figureDoi="http://doi.org/10.5281/zenodo.3814247" httpUri="https://zenodo.org/record/3814247/files/figure.png" pageId="18" pageNumber="19">Fig. 11</figureCitation>
). However, based on the available information it is impossible to distinguish different species of
<taxonomicName id="E2E45AC1CD766A74F6ED42DAFA84F9B7" authorityName="Foerste" authorityYear="1924" box="[1165,1313,1520,1546]" class="Cephalopoda" family="Oncoceratidae" genus="Beloitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oncocerida" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus">
<emphasis id="1790FD50CD766A74F6ED42DAFA84F9B7" box="[1165,1313,1520,1546]" italics="true" pageId="18" pageNumber="19">Beloitoceras</emphasis>
</taxonomicName>
among the specimens figured and described by
<bibRefCitation id="41755CB3CD766A74F0F24139FCB9F990" author="Dzik J." box="[658,796,1555,1581]" pageId="18" pageNumber="19" refId="ref47068" refString="Dzik J. 1984. Phylogeny of the Nautiloidea. Palaeontologia Polonica, Vol. 45, 203 pp." type="book" year="1984">Dzik (1984</bibRefCitation>
: pl. 8, fig. 1a-d),
<bibRefCitation id="41755CB3CD766A74F1844139FB2CF990" author="Strand T." box="[996,1161,1555,1581]" pageId="18" pageNumber="19" pagination="1 - 117" refId="ref49308" refString="Strand T. 1934. The Upper Ordovician Cephalopods of the Oslo Area. Norsk Geologiske Tidsskrift 14: 1 - 117." type="journal article" year="1934">Strand (1934)</bibRefCitation>
,
<bibRefCitation id="41755CB3CD766A74F6F5413EFABEF990" author="Frye M. W." box="[1173,1307,1555,1582]" pageId="18" pageNumber="19" pagination="83 - 99" refId="ref47793" refString="Frye M. W. 1987. Upper Ordovocian (Harjuan) oncoceratid nautiloids from Boda Limestone, Siljan District, Sweden. Geologiska Foreningens i Stockholm Forhandlingar 109: 83 - 99. http: // dx. doi. org / 10.1080 / 11035898709454748" type="journal article" year="1987">Frye (1987</bibRefCitation>
: fig. 3a, b) and the material described herein. Based on the available material,
<taxonomicName id="E2E45AC1CD766A74F191411DFB70F9ED" baseAuthorityName="Roemer" baseAuthorityYear="1861" box="[1009,1237,1591,1617]" class="Cephalopoda" family="Oncoceratidae" genus="Beloitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oncocerida" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="sinuososeptatum">
<emphasis id="1790FD50CD766A74F191411DFBA6F9ED" box="[1009,1027,1591,1616]" italics="true" pageId="18" pageNumber="19">B</emphasis>
.
<emphasis id="1790FD50CD766A74F671411DFB70F9ED" box="[1041,1237,1591,1616]" italics="true" pageId="18" pageNumber="19">sinuososeptatum</emphasis>
</taxonomicName>
is considered a species of
<taxonomicName id="E2E45AC1CD766A74F32A4173FE7BF9CE" authorityName="Foerste" authorityYear="1924" box="[330,478,1625,1651]" class="Cephalopoda" family="Oncoceratidae" genus="Beloitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oncocerida" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus">
<emphasis id="1790FD50CD766A74F32A4173FE7BF9CE" box="[330,478,1625,1651]" italics="true" pageId="18" pageNumber="19">Beloitoceras</emphasis>
</taxonomicName>
with a highly variable conch curvature, high variance in the angle of expansion (
<figureCitation id="BDDF3DC7CD766A74F2A64157FEBEF92A" box="[198,283,1661,1687]" captionStart="Fig" captionStartId="21.[189,232,1841,1867]" captionTargetBox="[201,1359,1426,1795]" captionTargetId="graphics@21.[284,756,1449,1701]" captionTargetPageId="21" captionText="Fig. 11. Morphological variation of Beloitoceras sinuososeptatum (Roemer, 1861). Circles: specimens of the Boda Limestone. Black dot: type specimen of Beloitoceras siljanense Frye, 1987. Black diamonds: types of Beloitoceras heterocurvatum Strand, 1934. Grey dot: type specimen of Beloitoceras landerense Foerste, 1935. A distinction between B. siljanense and B. sinuososeptatum is impossible based on the available material." figureDoi="http://doi.org/10.5281/zenodo.3814247" httpUri="https://zenodo.org/record/3814247/files/figure.png" pageId="18" pageNumber="19">Fig. 11</figureCitation>
), and potentially encompasses macro-, and microconch varieties.
</paragraph>
<paragraph id="255B2142CD766A74F2DD4194FC69F8D8" blockId="18.[189,1398,1726,1893]" pageId="18" pageNumber="19">
An apical fragment, PMU 26646, is interpreted as
<taxonomicName id="E2E45AC1CD766A74F16B4194FC3AF965" authorityName="Foerste" authorityYear="1924" box="[779,927,1726,1752]" class="Cephalopoda" family="Oncoceratidae" genus="Beloitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oncocerida" pageId="18" pageNumber="19" phylum="Mollusca" rank="genus">
<emphasis id="1790FD50CD766A74F16B4194FC3AF965" box="[779,927,1726,1752]" italics="true" pageId="18" pageNumber="19">Beloitoceras</emphasis>
</taxonomicName>
sp. (
<figureCitation id="BDDF3DC7CD766A74F1B74194FBBEF965" box="[983,1051,1726,1752]" captionStart="Fig" captionStartId="17.[188,231,1717,1743]" captionTargetBox="[188,1396,541,1671]" captionTargetId="figure@17.[188,1399,541,1676]" captionTargetPageId="17" captionText="Fig. 9. Oncerida of the Boda Limestone. Note the high variability in conch curvature and angle of expansion in Beloitoceras. A-B. Beloitoceras sinuososeptatum (Roemer, 1861). A. PMU 24769. B. PMU 26638, Kallholn. C-D. Beloitoceras siljanense Frye, 1987. C. PMU 26644. D. PMU 26645, Kallholn. E-F. Apex of Beloitoceras sp., PMU 26646, Kallholn. E. Adapical view. F. Lateral view. G-H. Cyrtorizoceras thorslundi sp. nov., PMU 26657, Osmundsberget, mature body chamber. G. Lateral view. H. Adapical view. I. Beloitoceras siljanense Frye, 1987, PMU 26643, Kallholn, with major conch repair at adoral part. J. Cyrtorizoceras thorslundi sp. nov., PMU 26647, holotype, Kallholn, complete juvenile specimen. Scale bars: A-D, G-J = 10 mm; E-F = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3814241" httpUri="https://zenodo.org/record/3814241/files/figure.png" pageId="18" pageNumber="19">Fig. 9</figureCitation>
E-F), because of its similarity in general conch shape and ornamentation with the larger fragments. The apex is cup-shaped and reaches a conch height of
<quantity id="E21C8CA7CD766A74F3F7402FFE51F8A2" box="[407,500,1797,1823]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="18" pageNumber="19" unit="mm" value="4.3">4.3 mm</quantity>
and a width of
<quantity id="E21C8CA7CD766A74F0D5402FFD5EF8A2" box="[693,763,1797,1823]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="18" pageNumber="19" unit="mm" value="4.0">4 mm</quantity>
at a distance of
<quantity id="E21C8CA7CD766A74F1A4402FFBAEF8A2" box="[964,1035,1797,1823]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="19" unit="mm" value="2.0">2 mm</quantity>
from the tip. The shell of this specimen is ornamented with fine rounded transverse annuli (ca. 810 per millimetre), which form a shallow hyponomic sinus on the concave side of the curved conch.
</paragraph>
<paragraph id="255B2142CD776A75F2DD4621FEFDFE98" blockId="19.[189,344,267,293]" box="[189,344,267,293]" pageId="19" pageNumber="20">
<heading id="7E13962ECD776A75F2DD4621FEFDFE98" bold="true" box="[189,344,267,293]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="1790FD50CD776A75F2DD4621FEFDFE98" bold="true" box="[189,344,267,293]" pageId="19" pageNumber="20">Comparison</emphasis>
</heading>
</paragraph>
<paragraph id="255B2142CD776A75F2DD4610FB94FD09" blockId="19.[189,1399,313,692]" pageId="19" pageNumber="20">
Two groups of
<taxonomicName id="E2E45AC1CD776A75F3114613FDA0FEEE" authorityName="Foerste" authorityYear="1924" box="[369,517,313,339]" class="Cephalopoda" family="Oncoceratidae" genus="Beloitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oncocerida" pageId="19" pageNumber="20" phylum="Mollusca" rank="genus">
<emphasis id="1790FD50CD776A75F3114613FDA0FEEE" box="[369,517,313,339]" italics="true" pageId="19" pageNumber="20">Beloitoceras</emphasis>
</taxonomicName>
are distinguishable within the Boda Limestone. According to
<bibRefCitation id="41755CB3CD776A75F6834610FAD7FEE9" author="Frye M. W." box="[1251,1394,314,340]" pageId="19" pageNumber="20" pagination="83 - 99" refId="ref47793" refString="Frye M. W. 1987. Upper Ordovocian (Harjuan) oncoceratid nautiloids from Boda Limestone, Siljan District, Sweden. Geologiska Foreningens i Stockholm Forhandlingar 109: 83 - 99. http: // dx. doi. org / 10.1080 / 11035898709454748" type="journal article" year="1987">Frye (1987)</bibRefCitation>
,
<taxonomicName id="E2E45AC1CD776A75F2DD4677FE47FECA" authority="Frye, 1987" authorityName="Frye" authorityYear="1987" box="[189,482,349,375]" class="Cephalopoda" family="Oncoceratidae" genus="Beloitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oncocerida" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="siljanense">
<emphasis id="1790FD50CD776A75F2DD4677FF6AFECB" box="[189,207,349,374]" italics="true" pageId="19" pageNumber="20">B</emphasis>
.
<emphasis id="1790FD50CD776A75F2804677FEF0FECA" box="[224,341,349,375]" italics="true" pageId="19" pageNumber="20">siljanense</emphasis>
<bibRefCitation id="41755CB3CD776A75F3004677FE47FECA" author="Frye M. W." box="[352,482,349,375]" pageId="19" pageNumber="20" pagination="83 - 99" refId="ref47793" refString="Frye M. W. 1987. Upper Ordovocian (Harjuan) oncoceratid nautiloids from Boda Limestone, Siljan District, Sweden. Geologiska Foreningens i Stockholm Forhandlingar 109: 83 - 99. http: // dx. doi. org / 10.1080 / 11035898709454748" type="journal article" year="1987">Frye, 1987</bibRefCitation>
</taxonomicName>
can be distinguished from
<taxonomicName id="E2E45AC1CD776A75F14F4677FBB2FECB" baseAuthorityName="Roemer" baseAuthorityYear="1861" box="[815,1047,349,375]" class="Cephalopoda" family="Oncoceratidae" genus="Beloitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oncocerida" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="sinuososeptatum">
<emphasis id="1790FD50CD776A75F14F4677FCE4FECB" box="[815,833,349,374]" italics="true" pageId="19" pageNumber="20">B</emphasis>
.
<emphasis id="1790FD50CD776A75F1334677FBB2FECB" box="[851,1047,349,374]" italics="true" pageId="19" pageNumber="20">sinuososeptatum</emphasis>
</taxonomicName>
in having an antisiphuncular conch margin that straightens relatively early during growth and remains straight in the mature body chamber. By contrast, in
<taxonomicName id="E2E45AC1CD776A75F384468EFD6CFE00" baseAuthorityName="Roemer" baseAuthorityYear="1861" box="[484,713,420,446]" class="Cephalopoda" family="Oncoceratidae" genus="Beloitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oncocerida" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="sinuososeptatum">
<emphasis id="1790FD50CD776A75F384468EFE53FE00" box="[484,502,420,445]" italics="true" pageId="19" pageNumber="20">B</emphasis>
.
<emphasis id="1790FD50CD776A75F065468EFD6CFE00" box="[517,713,420,445]" italics="true" pageId="19" pageNumber="20">sinuososeptatum</emphasis>
</taxonomicName>
the antisiphuncular conch margin is entirely curved along the entire length of the concave side of the conch. It was emphasized above that the conch curvature and angle of expansion vary strongly between specimens of
<taxonomicName id="E2E45AC1CD776A75F13046C3FC41FDBE" authorityName="Foerste" authorityYear="1924" box="[848,996,489,515]" class="Cephalopoda" family="Oncoceratidae" genus="Beloitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oncocerida" pageId="19" pageNumber="20" phylum="Mollusca" rank="genus">
<emphasis id="1790FD50CD776A75F13046C3FC41FDBE" box="[848,996,489,515]" italics="true" pageId="19" pageNumber="20">Beloitoceras</emphasis>
</taxonomicName>
from the Boda Limestone. Within this continuous spectrum of variation, the
<typeStatus id="FA5F9FE0CD776A75F0C54527FD46FD9A" box="[677,739,525,551]" pageId="19" pageNumber="20">types</typeStatus>
of both species represent two extremes with a maximum conch expansion in
<taxonomicName id="E2E45AC1CD776A75F3C9451BFD6DFDF7" authority="Frye, 1987" authorityName="Frye" authorityYear="1987" box="[425,712,560,586]" class="Cephalopoda" family="Oncoceratidae" genus="Beloitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oncocerida" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="siljanense">
<emphasis id="1790FD50CD776A75F3C9451BFE1EFDF7" box="[425,443,561,586]" italics="true" pageId="19" pageNumber="20">B</emphasis>
.
<emphasis id="1790FD50CD776A75F3AB451AFDE5FDF7" box="[459,576,560,586]" italics="true" pageId="19" pageNumber="20">siljanense</emphasis>
<bibRefCitation id="41755CB3CD776A75F028451AFD6DFDF7" author="Frye M. W." box="[584,712,560,586]" pageId="19" pageNumber="20" pagination="83 - 99" refId="ref47793" refString="Frye M. W. 1987. Upper Ordovocian (Harjuan) oncoceratid nautiloids from Boda Limestone, Siljan District, Sweden. Geologiska Foreningens i Stockholm Forhandlingar 109: 83 - 99. http: // dx. doi. org / 10.1080 / 11035898709454748" type="journal article" year="1987">Frye, 1987</bibRefCitation>
</taxonomicName>
(23°) and a minimum in
<taxonomicName id="E2E45AC1CD776A75F196451BFB7EFDF7" baseAuthorityName="Roemer" baseAuthorityYear="1861" box="[1014,1243,560,586]" class="Cephalopoda" family="Oncoceratidae" genus="Beloitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oncocerida" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="sinuososeptatum">
<emphasis id="1790FD50CD776A75F196451BFBADFDF7" box="[1014,1032,561,586]" italics="true" pageId="19" pageNumber="20">B</emphasis>
.
<emphasis id="1790FD50CD776A75F677451BFB7EFDF7" box="[1047,1243,561,586]" italics="true" pageId="19" pageNumber="20">sinuososeptatum</emphasis>
</taxonomicName>
(10°). Based on the angle of expansion alone, it is not possible to distinguish between the two species (
<figureCitation id="BDDF3DC7CD776A75F76F4579FACDFDD0" box="[1295,1384,595,621]" captionStart="Fig" captionStartId="21.[189,232,1841,1867]" captionTargetBox="[201,1359,1426,1795]" captionTargetId="graphics@21.[284,756,1449,1701]" captionTargetPageId="21" captionText="Fig. 11. Morphological variation of Beloitoceras sinuososeptatum (Roemer, 1861). Circles: specimens of the Boda Limestone. Black dot: type specimen of Beloitoceras siljanense Frye, 1987. Black diamonds: types of Beloitoceras heterocurvatum Strand, 1934. Grey dot: type specimen of Beloitoceras landerense Foerste, 1935. A distinction between B. siljanense and B. sinuososeptatum is impossible based on the available material." figureDoi="http://doi.org/10.5281/zenodo.3814247" httpUri="https://zenodo.org/record/3814247/files/figure.png" pageId="19" pageNumber="20">Fig. 11</figureCitation>
).
<taxonomicName id="E2E45AC1CD776A75F2DD455DFEF6FD2D" authorityName="Frye" authorityYear="1987" box="[189,339,630,657]" class="Cephalopoda" family="Oncoceratidae" genus="Beloitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oncocerida" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="siljanense">
<emphasis id="1790FD50CD776A75F2DD455DFF6AFD2D" box="[189,207,631,656]" italics="true" pageId="19" pageNumber="20">B</emphasis>
.
<emphasis id="1790FD50CD776A75F2BE455CFEF6FD2D" box="[222,339,630,656]" italics="true" pageId="19" pageNumber="20">siljanense</emphasis>
</taxonomicName>
may be distinguished from
<taxonomicName id="E2E45AC1CD776A75F0FC455DFC25FD2D" baseAuthorityName="Roemer" baseAuthorityYear="1861" box="[668,896,631,657]" class="Cephalopoda" family="Oncoceratidae" genus="Beloitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oncocerida" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="sinuososeptatum">
<emphasis id="1790FD50CD776A75F0FC455DFD0BFD2D" box="[668,686,631,656]" italics="true" pageId="19" pageNumber="20">B</emphasis>
.
<emphasis id="1790FD50CD776A75F0DC455DFC25FD2D" box="[700,896,631,656]" italics="true" pageId="19" pageNumber="20">sinuososeptatum</emphasis>
</taxonomicName>
in having a finer transverse ornamentation and a less curved antisiphuncular conch margin (compare
<figureCitation id="BDDF3DC7CD776A75F10745B3FC0EFD09" box="[871,939,665,692]" captionStart="Fig" captionStartId="17.[188,231,1717,1743]" captionTargetBox="[188,1396,541,1671]" captionTargetId="figure@17.[188,1399,541,1676]" captionTargetPageId="17" captionText="Fig. 9. Oncerida of the Boda Limestone. Note the high variability in conch curvature and angle of expansion in Beloitoceras. A-B. Beloitoceras sinuososeptatum (Roemer, 1861). A. PMU 24769. B. PMU 26638, Kallholn. C-D. Beloitoceras siljanense Frye, 1987. C. PMU 26644. D. PMU 26645, Kallholn. E-F. Apex of Beloitoceras sp., PMU 26646, Kallholn. E. Adapical view. F. Lateral view. G-H. Cyrtorizoceras thorslundi sp. nov., PMU 26657, Osmundsberget, mature body chamber. G. Lateral view. H. Adapical view. I. Beloitoceras siljanense Frye, 1987, PMU 26643, Kallholn, with major conch repair at adoral part. J. Cyrtorizoceras thorslundi sp. nov., PMU 26647, holotype, Kallholn, complete juvenile specimen. Scale bars: A-D, G-J = 10 mm; E-F = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.3814241" httpUri="https://zenodo.org/record/3814241/files/figure.png" pageId="19" pageNumber="20">Fig. 9</figureCitation>
A-B, C-D).
</paragraph>
<paragraph id="255B2142CD776A75F2DD45F1FD15FCE2" blockId="19.[189,1399,731,863]" pageId="19" pageNumber="20">
<taxonomicName id="E2E45AC1CD776A75F2DD45F1FD2CFD4B" authority="Foerste, 1935" authorityName="Foerste" authorityYear="1935" box="[189,649,731,758]" class="Cephalopoda" family="Oncoceratidae" genus="Beloitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oncocerida" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="landerense">
<emphasis id="1790FD50CD776A75F2DD45F1FE79FD48" box="[189,476,731,757]" italics="true" pageId="19" pageNumber="20">Beloitoceras landerense</emphasis>
Foerste, 1935
</taxonomicName>
from the mid Katian Lander Sandstone of
<collectingRegion id="E720EFA0CD776A75F6FA45F1FAB4FD48" box="[1178,1297,731,757]" country="United States of America" name="Wyoming" pageId="19" pageNumber="20">Wyoming</collectingRegion>
,
<collectingCountry id="5DF361D2CD776A75F77F45F1FAFDFD48" box="[1311,1368,731,757]" name="United States of America" pageId="19" pageNumber="20">USA</collectingCountry>
is very similar to
<taxonomicName id="E2E45AC1CD776A75F31A45D5FDC6FCA5" baseAuthorityName="Roemer" baseAuthorityYear="1861" box="[378,611,767,793]" class="Cephalopoda" family="Oncoceratidae" genus="Beloitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oncocerida" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="sinuososeptatum">
<emphasis id="1790FD50CD776A75F31A45D5FE29FCA5" box="[378,396,767,792]" italics="true" pageId="19" pageNumber="20">B</emphasis>
.
<emphasis id="1790FD50CD776A75F3FF45D5FDC6FCA5" box="[415,611,767,792]" italics="true" pageId="19" pageNumber="20">sinuososeptatum</emphasis>
</taxonomicName>
in general conch shape and dimensions (
<figureCitation id="BDDF3DC7CD776A75F63E45D4FB1DFCA5" box="[1118,1208,766,792]" captionStart="Fig" captionStartId="21.[189,232,1841,1867]" captionTargetBox="[201,1359,1426,1795]" captionTargetId="graphics@21.[284,756,1449,1701]" captionTargetPageId="21" captionText="Fig. 11. Morphological variation of Beloitoceras sinuososeptatum (Roemer, 1861). Circles: specimens of the Boda Limestone. Black dot: type specimen of Beloitoceras siljanense Frye, 1987. Black diamonds: types of Beloitoceras heterocurvatum Strand, 1934. Grey dot: type specimen of Beloitoceras landerense Foerste, 1935. A distinction between B. siljanense and B. sinuososeptatum is impossible based on the available material." figureDoi="http://doi.org/10.5281/zenodo.3814247" httpUri="https://zenodo.org/record/3814247/files/figure.png" pageId="19" pageNumber="20">Fig. 11</figureCitation>
). As the details of the ornamentation of this species are not known, direct comparison and synonymisation with
<taxonomicName id="E2E45AC1CD776A75F73D4408FE24FCE3" baseAuthorityName="Roemer" baseAuthorityYear="1861" class="Cephalopoda" family="Oncoceratidae" genus="Beloitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oncocerida" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="sinuososeptatum">
<emphasis id="1790FD50CD776A75F73D4408FACAFC86" box="[1373,1391,802,827]" italics="true" pageId="19" pageNumber="20">B</emphasis>
.
<emphasis id="1790FD50CD776A75F2DD446FFE24FCE3" box="[189,385,837,862]" italics="true" pageId="19" pageNumber="20">sinuososeptatum</emphasis>
</taxonomicName>
is not possible at present.
</paragraph>
<paragraph id="255B2142CD776A75F2DD44ACFACBFB2B" blockId="19.[189,1399,902,1175]" pageId="19" pageNumber="20">
<taxonomicName id="E2E45AC1CD776A75F2DD44ACFDC5FC1C" authority="(Hall, 1861)" baseAuthorityName="Hall" baseAuthorityYear="1861" box="[189,608,902,929]" class="Cephalopoda" family="Oncoceratidae" genus="Beloitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oncocerida" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="whitneyi">
<emphasis id="1790FD50CD776A75F2DD44ACFE65FC1D" box="[189,448,902,928]" italics="true" pageId="19" pageNumber="20">Beloitoceras whitneyi</emphasis>
(Hall, 1861)
</taxonomicName>
from the Elgin Formation, late Katian,
<collectingRegion id="E720EFA0CD776A75F63044ACFB28FC1D" box="[1104,1165,902,928]" country="United States of America" name="Iowa" pageId="19" pageNumber="20">Iowa</collectingRegion>
,
<collectingCountry id="5DF361D2CD776A75F6FE44ACFB72FC1D" box="[1182,1239,902,928]" name="United States of America" pageId="19" pageNumber="20">USA</collectingCountry>
is similar to
<taxonomicName id="E2E45AC1CD776A75F2DD4480FE02FC7E" baseAuthorityName="Roemer" baseAuthorityYear="1861" box="[189,423,938,964]" class="Cephalopoda" family="Oncoceratidae" genus="Beloitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oncocerida" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="sinuososeptatum">
<emphasis id="1790FD50CD776A75F2DD4480FF6AFC7E" box="[189,207,938,963]" italics="true" pageId="19" pageNumber="20">B</emphasis>
.
<emphasis id="1790FD50CD776A75F2834480FE02FC7E" box="[227,423,938,963]" italics="true" pageId="19" pageNumber="20">sinuososeptatum</emphasis>
</taxonomicName>
in general conch shape, its transversely ornamented shell, and in possessing a relatively large adult conch height of ca.
<quantity id="E21C8CA7CD776A75F0C944E6FCA5FC5A" box="[681,768,972,999]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="19" pageNumber="20" unit="mm" value="35.0">35 mm</quantity>
. It may be conspecific with the Boda specimens, but with ca. 910 chambers in a length similar to the corresponding conch height, the chamber distance (cameral depth) is significantly lower in
<taxonomicName id="E2E45AC1CD776A75F0D04339FC16FB90" baseAuthorityName="Hall" baseAuthorityYear="1861" box="[688,947,1043,1069]" class="Cephalopoda" family="Oncoceratidae" genus="Beloitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oncocerida" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="whitneyi">
<emphasis id="1790FD50CD776A75F0D04339FC16FB90" box="[688,947,1043,1069]" italics="true" pageId="19" pageNumber="20">Beloitoceras whitneyi</emphasis>
</taxonomicName>
than that of
<taxonomicName id="E2E45AC1CD776A75F637433EFAE5FB90" baseAuthorityName="Roemer" baseAuthorityYear="1861" box="[1111,1344,1043,1069]" class="Cephalopoda" family="Oncoceratidae" genus="Beloitoceras" higherTaxonomySource="GBIF" kingdom="Animalia" order="Oncocerida" pageId="19" pageNumber="20" phylum="Mollusca" rank="species" species="sinuososeptatum">
<emphasis id="1790FD50CD776A75F637433EFBCCFB90" box="[1111,1129,1044,1069]" italics="true" pageId="19" pageNumber="20">B</emphasis>
.
<emphasis id="1790FD50CD776A75F61C433EFAE5FB90" box="[1148,1344,1044,1069]" italics="true" pageId="19" pageNumber="20">sinuososeptatum</emphasis>
</taxonomicName>
(six chambers per length similar to corresponding conch height) and nothing is known about the variability of this character in either species. A synonymisation of this species is very likely, but may only be possible to demonstrate after comparison of the variability of the chamber height in both assemblages.
</paragraph>
</subSubSection>
<subSubSection id="6DFE72C9CD776A75F2DD4397FD69FACD" pageId="19" pageNumber="20" type="multiple">
<paragraph id="255B2142CD776A75F2DD4397FDD8FB65" blockId="19.[189,637,1213,1240]" box="[189,637,1213,1240]" pageId="19" pageNumber="20">
<heading id="7E13962ECD776A75F2DD4397FDD8FB65" bold="true" box="[189,637,1213,1240]" fontSize="11" level="3" pageId="19" pageNumber="20" reason="3">
<emphasis id="1790FD50CD776A75F2DD4397FDD8FB65" bold="true" box="[189,637,1213,1240]" pageId="19" pageNumber="20">Stratigraphic and geographic range</emphasis>
</heading>
</paragraph>
<paragraph id="255B2142CD776A75F2DD43C7FD69FACD" blockId="19.[189,1398,1260,1392]" pageId="19" pageNumber="20">
Bønsnes Formation, Ringerike district, Skogerholm Formation, Oslo-Asker district,
<collectingCountry id="5DF361D2CD776A75F6D743C7FAB0FABA" box="[1207,1301,1261,1287]" name="Norway" pageId="19" pageNumber="20">Norway</collectingCountry>
(
<bibRefCitation id="41755CB3CD776A75F74843C6FF59FA97" author="Strand T." pageId="19" pageNumber="20" pagination="1 - 117" refId="ref49308" refString="Strand T. 1934. The Upper Ordovician Cephalopods of the Oslo Area. Norsk Geologiske Tidsskrift 14: 1 - 117." type="journal article" year="1934">Strand 1934</bibRefCitation>
); erratic limestone boulders from Vormsi/Pirgu Regional Stage (“Lyckholm stage”), Zawidowice,
<collectingCountry id="5DF361D2CD776A75F2DD4219FEABFAF0" box="[189,270,1331,1357]" name="Poland" pageId="19" pageNumber="20">Poland</collectingCountry>
(
<bibRefCitation id="41755CB3CD776A75F37F4219FE04FAF0" author="Dzik J." box="[287,417,1331,1357]" pageId="19" pageNumber="20" refId="ref47068" refString="Dzik J. 1984. Phylogeny of the Nautiloidea. Palaeontologia Polonica, Vol. 45, 203 pp." type="book" year="1984">Dzik 1984</bibRefCitation>
); Vormsi/Pirgu Regional Stage (“Lyckholm stage”),
<collectingCountry id="5DF361D2CD776A75F6714219FBCCFAF0" box="[1041,1129,1331,1357]" name="Estonia" pageId="19" pageNumber="20">Estonia</collectingCountry>
(
<bibRefCitation id="41755CB3CD776A75F6194219FA87FAF0" author="Teichert C." box="[1145,1314,1331,1357]" pageId="19" pageNumber="20" pagination="264 - 312" refId="ref49750" refString="Teichert C. 1930. Die Cephalopoden-Fauna der Lyckholm-Stufe des Ostbaltikums. Palaontologische Zeitschrift 12: 264 - 312." type="journal article" year="1930">Teichert 1930</bibRefCitation>
); Boda Limestone,
<collectingRegion id="E720EFA0CD776A75F327427CFE02FACD" box="[327,423,1366,1392]" country="Sweden" name="Dalarna" pageId="19" pageNumber="20">Dalarna</collectingRegion>
,
<collectingCountry id="5DF361D2CD776A75F3D3427CFDB1FACD" box="[435,532,1366,1392]" name="Sweden" pageId="19" pageNumber="20">Sweden</collectingCountry>
; all late Katian.
</paragraph>
</subSubSection>
</treatment>
</document>