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284 lines
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<document id="659AE2C48DF010DEBF97952A9881C845" ID-CLB-Dataset="25414" ID-DOI="10.11646/zootaxa.4674.2.1" ID-GBIF-Dataset="a3c9f2fa-30d7-48b9-9a2e-3f81cc0a3e3e" ID-ISSN="1175-5326" ID-Zenodo-Dep="3458836" ID-ZooBank="B6626FE9-6490-4BC9-BACB-A4437AACDF25" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1569309377345" checkinUser="plazi" docAuthor="Bollino, Maurizio, Uliana, Marco & Sabatinelli, Guido" docDate="2019" docId="03F087D8C154842EFF0CFF31FCFD8C70" docLanguage="en" docName="zootaxa.4674.2.1.pdf.imf" docOrigin="Zootaxa 4674 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pygopleurus Motschulsky 1860" docType="treatment" docVersion="9" lastPageNumber="156" masterDocId="FFC9FFA0C150842BFF9BFFD0FF8D8D17" masterDocTitle="The Pygopleurus Motschulsky (Coleoptera: Scarabaeoidea: Glaphyridae) of mainland Greece: taxonomy, faunistics and ecological notes" masterLastPageNumber="202" masterPageNumber="151" pageNumber="155" updateTime="1698756305895" updateUser="ExternalLinkService">
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<mods:title id="AD31B1DBE1347F50B7F1A83F56A3D258">The Pygopleurus Motschulsky (Coleoptera: Scarabaeoidea: Glaphyridae) of mainland Greece: taxonomy, faunistics and ecological notes</mods:title>
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<mods:name id="0DD65AB8CC8E062F494B209F793818C0" type="personal">
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<mods:namePart id="EE2A8D22BBC7DE0DA2C4121726D8A4AF">Bollino, Maurizio</mods:namePart>
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<mods:namePart id="E3E2AF77E89EF07DD6FE6329FB44D14B">Uliana, Marco</mods:namePart>
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<mods:namePart id="34E49D18910719CF37053EACA3A1C38B">Sabatinelli, Guido</mods:namePart>
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<mods:title id="12F1997C997EBC1894CE56D9264B9AF9">Zootaxa</mods:title>
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<mods:part id="12C3840D53512B059C0FE88FEC001C62">
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<mods:date id="DCB5965B3B7D9674D4E5C69B61AC1978">2019</mods:date>
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<mods:number id="1DD91A1C31128476A851FAD89C4BD793">2019-09-24</mods:number>
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<mods:number id="FB11EEFF44EF35F271E560D0FC0E0238">4674</mods:number>
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<treatment id="03F087D8C154842EFF0CFF31FCFD8C70" ID-GBIF-Taxon="159398097" LSID="urn:lsid:plazi:treatment:03F087D8C154842EFF0CFF31FCFD8C70" httpUri="http://treatment.plazi.org/id/03F087D8C154842EFF0CFF31FCFD8C70" lastPageId="5" lastPageNumber="156" pageId="4" pageNumber="155">
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<subSubSection id="C3436545C154842FFF0CFF31FD758DEB" box="[151,760,225,252]" pageId="4" pageNumber="155" type="nomenclature">
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<paragraph id="8BE636CEC154842FFF0CFF31FD758DEB" blockId="4.[151,1437,225,576]" box="[151,760,225,252]" pageId="4" pageNumber="155">
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<heading id="D0AE81A2C154842FFF0CFF31FD758DEB" bold="true" box="[151,760,225,252]" fontSize="11" level="1" pageId="4" pageNumber="155" reason="1">
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<emphasis id="B92DEADCC154842FFF0CFF31FD758DEB" bold="true" box="[151,760,225,252]" pageId="4" pageNumber="155">
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<taxonomicName id="4C594D4DC154842FFF0CFF31FEA58DEB" ID-CoL="75V3" authority="Motschulsky, 1860" authorityName="Motschulsky" authorityYear="1860" box="[151,296,225,252]" class="Insecta" family="Glaphyridae" genus="Pygopleurus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
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<emphasis id="B92DEADCC154842FFF0CFF31FEA58DEB" bold="true" box="[151,296,225,252]" italics="true" pageId="4" pageNumber="155">Pygopleurus</emphasis>
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</taxonomicName>
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of
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<collectingCountry id="F34E765EC154842FFED6FF31FE288DEB" box="[333,421,225,252]" name="Greece" pageId="4" pageNumber="155">Greece</collectingCountry>
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: brief historical background
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3436545C154842EFF5CFED5FB1E8C08" lastPageId="5" lastPageNumber="156" pageId="4" pageNumber="155" type="discussion">
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<paragraph id="8BE636CEC154842FFF5CFED5FE618CE0" blockId="4.[151,1437,225,576]" pageId="4" pageNumber="155">
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The first report of
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<taxonomicName id="4C594D4DC154842FFE0DFED5FDAD8C08" authorityName="MacLeay" authorityYear="1819" box="[406,544,261,287]" class="Insecta" family="Glaphyridae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="family">Glaphyridae</taxonomicName>
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for mainland
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<collectingCountry id="F34E765EC154842FFD24FED5FC838C08" box="[703,782,261,287]" name="Greece" pageId="4" pageNumber="155">Greece</collectingCountry>
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was by
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<bibRefCitation id="EFC84B3FC154842FFCF1FED5FB898C08" author="Brulle, M." box="[874,1028,261,287]" pageId="4" pageNumber="155" pagination="165 - 187" refId="ref29938" refString="Brulle, M. (1832) Expedition scientifique de Moree. Section des Sciences physiques. Ie Partie: Zoologie 2 e section: Des Animaux articules. F. G. Levrault, Paris, 400 pp. [165 - 187]" type="book chapter" year="1832">Brullé (1832)</bibRefCitation>
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, who described six species based on specimens collected during his expedition to the
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<collectingRegion id="499DF82CC154842FFD28FEF9FCC98C54" box="[691,836,297,323]" country="Greece" name="Peloponnisos" pageId="4" pageNumber="155">Peloponnese</collectingRegion>
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. In this work are included some of the current-day
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<emphasis id="B92DEADCC154842FFAEEFEFAFE8A8C70" italics="true" pageId="4" pageNumber="155">Py- gopleurus</emphasis>
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that he referred to the genus
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<taxonomicName id="4C594D4DC154842FFDC9FE9DFD558C70" authorityName="Latreille" authorityYear="1807" box="[594,728,333,359]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
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<emphasis id="B92DEADCC154842FFDC9FE9DFD558C70" box="[594,728,333,359]" italics="true" pageId="4" pageNumber="155">Amphicoma</emphasis>
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</taxonomicName>
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. Subsequently, other authors have mentioned
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<taxonomicName id="4C594D4DC154842FFB44FE9DFAE08C70" box="[1247,1389,333,359]" class="Insecta" family="Glaphyridae" genus="Pygopleurus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
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<emphasis id="B92DEADCC154842FFB44FE9DFAE08C70" box="[1247,1389,333,359]" italics="true" pageId="4" pageNumber="155">Pygopleurus</emphasis>
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</taxonomicName>
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and other
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<taxonomicName id="4C594D4DC154842FFF43FEA1FEEF8C9C" authorityName="MacLeay" authorityYear="1819" box="[216,354,369,395]" class="Insecta" family="Glaphyridae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="family">Glaphyridae</taxonomicName>
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in faunistic lists for
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<collectingCountry id="F34E765EC154842FFDD1FEA1FD148C9C" box="[586,665,369,395]" name="Greece" pageId="4" pageNumber="155">Greece</collectingCountry>
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(
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<bibRefCitation id="EFC84B3FC154842FFD33FEA1FC0D8C9B" author="Kiesenwetter, H. V." box="[680,896,369,396]" pageId="4" pageNumber="155" pagination="231 - 249" refId="ref30208" refString="Kiesenwetter, H. V. (1858) Beitrag zur Kaferfauna Griechelands. Viertes Stuck: Parnidae, Heteroceridae, Lamellicornia, Buprestidae. Berliner Entomologische Zeitschrift, 2, 231 - 249. https: // doi. org / 10.1002 / mmnd. 18580020304" type="journal article" year="1858">Kiesenwetter 1858</bibRefCitation>
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;
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<bibRefCitation id="EFC84B3FC154842FFC17FEA1FBA28C9B" author="Oertzen, E." box="[908,1071,369,396]" pageId="4" pageNumber="155" pagination="189 - 293" refId="ref30722" refString="Oertzen, E. (1886) Verzeichnis der Coleopteren Griechenlands und Cretas. Berliner Entomologische Zeitschrift, 30, 189 - 293." type="journal article" year="1886">Oertzen, 1886</bibRefCitation>
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), or in taxonomic works (
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<bibRefCitation id="EFC84B3FC154842FFAD5FEA1FF5E8CB8" author="Reitter, E." pageId="4" pageNumber="155" pagination="53 - 64" refId="ref31091" refString="Reitter, E. (1890) Analytische revision der Coleopteren-Gattung Amphicoma, subgenus Pygopleurus Motsch. Deutsche Entomologische Zeitschrift, 1890, 53 - 64. https: // doi. org / 10.1002 / mmnd. 48018900307" type="journal article" year="1890">Reitter 1890</bibRefCitation>
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), but over seventy years passed before a new taxon was proposed for this country (
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<bibRefCitation id="EFC84B3FC154842FFB0FFE45FAA58CB8" author="Reitter, E." box="[1172,1320,405,431]" pageId="4" pageNumber="155" pagination="28 - 158" refId="ref31130" refString="Reitter, E. (1903) Bestimmungs-Tabelle der Melolonthidae. IV. Rutelini, Hopliini und Glaphyrini. Verhandlungen des Naturforschenden Vereins, Brunn, 51, 28 - 158. [1902]" type="journal article" year="1903">Reitter 1903</bibRefCitation>
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). After 50 years,
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<bibRefCitation id="EFC84B3FC154842FFF44FE69FEF08CC3" author="Miksic, R." box="[223,381,441,468]" pageId="4" pageNumber="155" pagination="39 - 44" refId="ref30446" refString="Miksic, R. (1959) Beitrage zur Kenntnis der Scarabaeidae. Fauna der Mediterranen Gebietes und der Balkan-Halbinsel. Bollettino della Societa Entomologica Italiana, 89, 39 - 44." type="journal article" year="1959">Mikšić (1959)</bibRefCitation>
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and
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<bibRefCitation id="EFC84B3FC154842FFE2AFE69FDD28CC3" author="Petrovitz, R." box="[433,607,441,468]" pageId="4" pageNumber="155" pagination="38 - 68" refId="ref30828" refString="Petrovitz, R. (1958) Das Subgenus Pygopleurus Motsch. der Gattung Amphicoma Latr. (Col. Scarabaeidae, Glaphyrinae). Entomologisk Tidskrift, 78 (Supplement), 38 - 68." type="journal article" year="1958">Petrovitz (1958</bibRefCitation>
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,
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<bibRefCitation id="EFC84B3FC154842FFDF2FE69FD298CC4" author="Petrovitz, R." box="[617,676,441,467]" pageId="4" pageNumber="155" pagination="99 - 105" refId="ref30927" refString="Petrovitz, R. (1964) Amphicoma - Studien. Zur Kenntnis einiger Amphicoma - Arten. Reichenbachia, 3 (5), 99 - 105." type="journal article" year="1964">1964</bibRefCitation>
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,
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<bibRefCitation id="EFC84B3FC154842FFD35FE69FD6A8CC4" author="Petrovitz, R." box="[686,743,441,467]" pageId="4" pageNumber="155" pagination="21 - 30" refId="ref31015" refString="Petrovitz, R. (1972) Palaarktische und Orientalische Glaphyrinae-Formen (Coleoptera, Scarabaeidae). Mitteilungen aus dem Zoologischen Museum in Berlin, 48, 21 - 30. https: // doi. org / 10.1002 / mmnz. 19720480103" type="journal article" year="1972">1972</bibRefCitation>
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) described a few species and subspecies and introduced names for “forms” and “aberrations”.
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</paragraph>
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<paragraph id="8BE636CEC154842FFF5CFDD1FA968F28" blockId="4.[151,1437,225,576]" pageId="4" pageNumber="155">
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Finally,
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<bibRefCitation id="EFC84B3FC154842FFEB9FDD1FE498F0C" author="Baraud, J." box="[290,452,513,539]" pageId="4" pageNumber="155" pagination="331 - 375" refId="ref29740" refString="Baraud, J. (1989) Revision du genre Pygopleurus Motschulsky (Coleoptera, Scarabaoidea: Glaphyridae). Annales de la Societe Entomologique de France, 25 (3), 331 - 375, 89 figs." type="book chapter" year="1989">Baraud (1989)</bibRefCitation>
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consolidated the taxonomy of the Greek
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<taxonomicName id="4C594D4DC154842FFC14FDD1FB908F0C" box="[911,1053,513,539]" class="Insecta" family="Glaphyridae" genus="Pygopleurus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
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<emphasis id="B92DEADCC154842FFC14FDD1FB908F0C" box="[911,1053,513,539]" italics="true" pageId="4" pageNumber="155">Pygopleurus</emphasis>
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</taxonomicName>
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in his revision of the genus recognizing 7 species for the mainland. To date, no further taxonomic treatments nor studies were published.
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</paragraph>
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<paragraph id="8BE636CEC154842FFF0CFDBEFBBE8F9F" blockId="4.[151,1437,621,1044]" box="[151,1075,621,648]" pageId="4" pageNumber="155">
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<heading id="D0AE81A2C154842FFF0CFDBEFBBE8F9F" bold="true" box="[151,1075,621,648]" fontSize="11" level="1" pageId="4" pageNumber="155" reason="1">
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<emphasis id="B92DEADCC154842FFF0CFDBEFBBE8F9F" bold="true" box="[151,1075,621,648]" pageId="4" pageNumber="155">
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The taxa described by Brullé: restriction of
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<typeStatus id="54E2886CC154842FFD39FDBEFD588F9F" box="[674,725,622,648]" pageId="4" pageNumber="155">type</typeStatus>
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localities and
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<typeStatus id="54E2886CC154842FFC1BFDBEFC3E8F9F" box="[896,947,622,648]" pageId="4" pageNumber="155">type</typeStatus>
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specimens
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BE636CEC154842FFF5CFD41FAE08EB0" blockId="4.[151,1437,621,1044]" pageId="4" pageNumber="155">
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All taxa described by
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<bibRefCitation id="EFC84B3FC154842FFE59FD41FDD68FBC" author="Brulle, M." box="[450,603,657,683]" pageId="4" pageNumber="155" pagination="165 - 187" refId="ref29938" refString="Brulle, M. (1832) Expedition scientifique de Moree. Section des Sciences physiques. Ie Partie: Zoologie 2 e section: Des Animaux articules. F. G. Levrault, Paris, 400 pp. [165 - 187]" type="book chapter" year="1832">Brullé (1832)</bibRefCitation>
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were collected during his scientific expedition to Morea (currently, Peloponnese) in 1829, but most of them had no
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<typeStatus id="54E2886CC154842FFDE8FD66FD2E8FC7" box="[627,675,694,720]" pageId="4" pageNumber="155">type</typeStatus>
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locality indicated. By crosschecking information provided by
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<bibRefCitation id="EFC84B3FC154842FFACDFD65FF6F8FE4" author="Brulle, M." pageId="4" pageNumber="155" pagination="165 - 187" refId="ref29938" refString="Brulle, M. (1832) Expedition scientifique de Moree. Section des Sciences physiques. Ie Partie: Zoologie 2 e section: Des Animaux articules. F. G. Levrault, Paris, 400 pp. [165 - 187]" type="book chapter" year="1832">Brullé (1832)</bibRefCitation>
|
|||
|
|
and those found in the report by
|
|||
|
|
<bibRefCitation id="EFC84B3FC154842FFDC5FD0AFC388FE3" author="Bory de Saint-Vincent, J." box="[606,949,729,756]" pageId="4" pageNumber="155" refId="ref29907" refString="Bory de Saint-Vincent, J. (1836) Expedition Scientifique de Moree. Tome Premier. Relation. F. G. Levrault, Paris iv + 472 pp." type="book" year="1836">
|
|||
|
|
Bory De
|
|||
|
|
<collectingCountry id="F34E765EC154842FFD5DFD09FCED8FE4" box="[710,864,729,755]" name="Saint Vincent and the Grenadines" pageId="4" pageNumber="155">Saint-Vincent</collectingCountry>
|
|||
|
|
(1836)
|
|||
|
|
</bibRefCitation>
|
|||
|
|
, we determined that Brullé carried out two collecting trips in the spring 1829:
|
|||
|
|
<date id="FFE7100EC154842FFD80FD2DFC9F8E00" box="[539,786,764,792]" pageId="4" pageNumber="155" value="2015-03" valueMax="2015-04-10" valueMin="2015-03-30">
|
|||
|
|
30
|
|||
|
|
<superScript id="7C2C9B86C154842FFDACFD2CFDC98E1D" attach="left" box="[567,580,764,778]" fontSize="6" pageId="4" pageNumber="155">th</superScript>
|
|||
|
|
March – 10
|
|||
|
|
<superScript id="7C2C9B86C154842FFD5CFD2CFD598E1D" attach="left" box="[711,724,764,778]" fontSize="6" pageId="4" pageNumber="155">th</superScript>
|
|||
|
|
April
|
|||
|
|
</date>
|
|||
|
|
and
|
|||
|
|
<date id="FFE7100EC154842FFCDCFD2DFBA38E0F" box="[839,1070,764,792]" pageId="4" pageNumber="155">
|
|||
|
|
16
|
|||
|
|
<superScript id="7C2C9B86C154842FFCF8FD2CFCFD8E1D" attach="right" box="[867,880,764,778]" fontSize="6" pageId="4" pageNumber="155">th</superScript>
|
|||
|
|
April – 15
|
|||
|
|
<superScript id="7C2C9B86C154842FFC7FFD2CFC7C8E1D" attach="left" box="[996,1009,764,778]" fontSize="6" pageId="4" pageNumber="155">th</superScript>
|
|||
|
|
May.
|
|||
|
|
</date>
|
|||
|
|
He undertook additional collecting trips during the late spring and the summer of the same year, however after the period of activity of
|
|||
|
|
<taxonomicName id="4C594D4DC154842FFA94FCF1FA108E2C" box="[1295,1437,801,827]" class="Insecta" family="Glaphyridae" genus="Pygopleurus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B92DEADCC154842FFA94FCF1FA108E2C" box="[1295,1437,801,827]" italics="true" pageId="4" pageNumber="155">Pygopleurus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
that, in the area, ends by late May. During both spring trips Brullé collected in an area roughly comprised between Filiatra (presently Trifyllia), Kalamata, and Methoni (presently Pylos-Nestoras), within the current Messenia regional unit. Consequently, we restrict the
|
|||
|
|
<typeStatus id="54E2886CC154842FFDF1FC5EFD178EBF" box="[618,666,910,936]" pageId="4" pageNumber="155">type</typeStatus>
|
|||
|
|
locality of
|
|||
|
|
<taxonomicName id="4C594D4DC154842FFC81FC5DFC258EB0" box="[794,936,909,935]" class="Insecta" family="Glaphyridae" genus="Pygopleurus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B92DEADCC154842FFC81FC5DFC258EB0" box="[794,936,909,935]" italics="true" pageId="4" pageNumber="155">Pygopleurus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
taxa described by Brullé to “Messenia”.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BE636CEC154842FFF5CFC61FCF68904" blockId="4.[151,1437,621,1044]" pageId="4" pageNumber="155">
|
|||
|
|
The material collected by Brullé, according to Horn & Kale (1937), should be housed at the MNHN. Unfortunately, most of the original specimens are untraceable or wrongly identified as such.
|
|||
|
|
<bibRefCitation id="EFC84B3FC154842FFBDBFC05FB6E8EF8" author="Baraud, J." box="[1088,1251,981,1007]" pageId="4" pageNumber="155" pagination="331 - 375" refId="ref29740" refString="Baraud, J. (1989) Revision du genre Pygopleurus Motschulsky (Coleoptera, Scarabaoidea: Glaphyridae). Annales de la Societe Entomologique de France, 25 (3), 331 - 375, 89 figs." type="book chapter" year="1989">Baraud (1989)</bibRefCitation>
|
|||
|
|
, who designated some
|
|||
|
|
<typeStatus id="54E2886CC154842FFF42FC2AFEB28903" box="[217,319,1018,1044]" pageId="4" pageNumber="155" type="neotype">neotypes</typeStatus>
|
|||
|
|
, already acknowledged the loss of type specimens.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BE636CEC154842FFF0CFB91FCC5894B" blockId="4.[151,1437,1089,1835]" box="[151,840,1089,1116]" pageId="4" pageNumber="155">
|
|||
|
|
<heading id="D0AE81A2C154842FFF0CFB91FCC5894B" bold="true" box="[151,840,1089,1116]" fontSize="11" level="1" pageId="4" pageNumber="155" reason="1">
|
|||
|
|
<emphasis id="B92DEADCC154842FFF0CFB91FCC5894B" bold="true" box="[151,840,1089,1116]" pageId="4" pageNumber="155">Infrasubspecific value of the forms described by Petrovitz</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BE636CEC154842FFF5CFBB5FE2289DF" blockId="4.[151,1437,1089,1835]" pageId="4" pageNumber="155">
|
|||
|
|
<bibRefCitation id="EFC84B3FC154842FFF5CFBB5FE0C8997" author="Petrovitz, R." box="[199,385,1125,1152]" pageId="4" pageNumber="155" pagination="38 - 68" refId="ref30828" refString="Petrovitz, R. (1958) Das Subgenus Pygopleurus Motsch. der Gattung Amphicoma Latr. (Col. Scarabaeidae, Glaphyrinae). Entomologisk Tidskrift, 78 (Supplement), 38 - 68." type="journal article" year="1958">Petrovitz (1958)</bibRefCitation>
|
|||
|
|
, in his work on
|
|||
|
|
<taxonomicName id="4C594D4DC154842FFDA1FBB5FD458968" box="[570,712,1125,1151]" class="Insecta" family="Glaphyridae" genus="Pygopleurus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B92DEADCC154842FFDA1FBB5FD458968" box="[570,712,1125,1151]" italics="true" pageId="4" pageNumber="155">Pygopleurus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, proposed many new names as f.[orm]. Following, we maintain that these names should be considered infrasubsecific according to article 45.6.4 of ICZN, and hence unavailable according to article 45.5.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BE636CEC154842FFF5CFB02FF7D8A3C" blockId="4.[151,1437,1089,1835]" pageId="4" pageNumber="155">
|
|||
|
|
In the introduction, Petrovitz clearly states “
|
|||
|
|
<emphasis id="B92DEADCC154842FFD5EFB02FAEC8824" italics="true" pageId="4" pageNumber="155">
|
|||
|
|
Eine endgültige Klärung der noch offenen Probleme, vor allem bezüglich der wirklichen Wertigkeit
|
|||
|
|
<emphasis id="B92DEADCC154842FFDB6FB25FBDF8807" bold="true" box="[557,1106,1269,1296]" italics="true" pageId="4" pageNumber="155">der hier als Aberrationen bezeichneten Formen</emphasis>
|
|||
|
|
, wird vielleicht nicht in allen Fällen am Schreibtisch, sondern in der eingangs erwähnten Art biologischer Beobachtungen zu suchen sein
|
|||
|
|
</emphasis>
|
|||
|
|
[= A final elucidation of still open issues, primarily on the true value of
|
|||
|
|
<emphasis id="B92DEADCC154842FFC03FAEEFAF4884F" bold="true" box="[920,1401,1341,1368]" pageId="4" pageNumber="155">the forms described here as aberrations</emphasis>
|
|||
|
|
, is perhaps not to be sought in all cases at desk, but by biological observations of the species listed in the introduction]. Hence, the rank of the f.[orm]
|
|||
|
|
<emphasis id="B92DEADCC154842FFE70FA56FDA48888" box="[491,553,1414,1439]" italics="true" pageId="4" pageNumber="155">sensu</emphasis>
|
|||
|
|
Petrovitz, is clearly introduced as infrasubspecific. In addition, in the same paper, Petrovitz explicitly used the term of subspecies (cited as “ssp.”) for some taxa, applying it at a rank higher than forms (“f.”): this is clear from the
|
|||
|
|
<typeStatus id="54E2886CC154842FFD81FA1EFDC788FF" box="[538,586,1486,1512]" pageId="4" pageNumber="155">type</typeStatus>
|
|||
|
|
setting of the paper itself and by the subordination of some “forms” (hence quadrinomial) to “
|
|||
|
|
<taxonomicName id="4C594D4DC154842FFEF2FA21FC188B1C" authority="Petrovitz 1958: 42" authorityName="Miksic" authorityPageNumber="42" authorityYear="1955" baseAuthorityName="Petrovitz" baseAuthorityPageNumber="42" baseAuthorityYear="1958" box="[361,917,1521,1548]" class="Insecta" family="Glaphyridae" genus="Amphicoma" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="subSpecies" species="vulpes" subGenus="Pygopleurus" subSpecies="bogoevskii">
|
|||
|
|
<emphasis id="B92DEADCC154842FFEF2FA21FE628B1C" box="[361,495,1521,1547]" italics="true" pageId="4" pageNumber="155">Amphicoma</emphasis>
|
|||
|
|
(
|
|||
|
|
<emphasis id="B92DEADCC154842FFE64FA21FD008B1C" box="[511,653,1521,1547]" italics="true" pageId="4" pageNumber="155">Pygopleurus</emphasis>
|
|||
|
|
)
|
|||
|
|
<emphasis id="B92DEADCC154842FFD06FA21FD688B1C" box="[669,741,1521,1547]" italics="true" pageId="4" pageNumber="155">vulpes</emphasis>
|
|||
|
|
ssp.
|
|||
|
|
<emphasis id="B92DEADCC154842FFC86FA21FC188B1C" box="[797,917,1521,1547]" italics="true" pageId="4" pageNumber="155">bogoevskii</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
” (
|
|||
|
|
<bibRefCitation id="EFC84B3FC154842FFC2AFA21FB048B1C" author="Petrovitz, R." box="[945,1161,1521,1548]" pageId="4" pageNumber="155" pagination="38 - 68" refId="ref30828" refString="Petrovitz, R. (1958) Das Subgenus Pygopleurus Motsch. der Gattung Amphicoma Latr. (Col. Scarabaeidae, Glaphyrinae). Entomologisk Tidskrift, 78 (Supplement), 38 - 68." type="journal article" year="1958">Petrovitz 1958: 42</bibRefCitation>
|
|||
|
|
). To further substantiate our conclusions, we can add that many “forms” shared a same descriptive color name which was applied in combination to several species at the same time (see, for example, f.
|
|||
|
|
<emphasis id="B92DEADCC154842FFCE9F9E9FC5C8B44" box="[882,977,1593,1619]" italics="true" pageId="4" pageNumber="155">violacea</emphasis>
|
|||
|
|
and f.
|
|||
|
|
<emphasis id="B92DEADCC154842FFBBAF9EAFBE38B44" box="[1057,1134,1594,1619]" italics="true" pageId="4" pageNumber="155">cuprea</emphasis>
|
|||
|
|
, each used in combination with six species, or f.
|
|||
|
|
<emphasis id="B92DEADCC154842FFE10F98DFE5E8B60" box="[395,467,1629,1655]" italics="true" pageId="4" pageNumber="155">viridis</emphasis>
|
|||
|
|
, used in combination with eleven species) then with clear sense of individual form not species-specific. For example, treating
|
|||
|
|
<taxonomicName id="4C594D4DC154842FFDC2F951FB4C8B8C" authority="(Petrovitz 1958: 49) Petrovitz" authorityName="Petrovitz" authorityPageNumber="49" baseAuthorityName="Petrovitz" baseAuthorityPageNumber="49" baseAuthorityYear="1958" box="[601,1217,1665,1692]" class="Insecta" family="Glaphyridae" genus="Pygopleurus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="humeralis">
|
|||
|
|
<emphasis id="B92DEADCC154842FFDC2F951FCEC8B8C" box="[601,865,1665,1691]" italics="true" pageId="4" pageNumber="155">Pygopleurus humeralis</emphasis>
|
|||
|
|
(
|
|||
|
|
<bibRefCitation id="EFC84B3FC154842FFCE9F951FBC78B8C" author="Petrovitz, R." box="[882,1098,1665,1692]" pageId="4" pageNumber="155" pagination="38 - 68" refId="ref30828" refString="Petrovitz, R. (1958) Das Subgenus Pygopleurus Motsch. der Gattung Amphicoma Latr. (Col. Scarabaeidae, Glaphyrinae). Entomologisk Tidskrift, 78 (Supplement), 38 - 68." type="journal article" year="1958">Petrovitz 1958: 49</bibRefCitation>
|
|||
|
|
) Petrovitz
|
|||
|
|
</taxonomicName>
|
|||
|
|
stated “
|
|||
|
|
<emphasis id="B92DEADCC154842FFAB8F951FCA08BA8" italics="true" pageId="4" pageNumber="155">Vorderkörper leuchtend grün (f. viridis), mitunter mit Goldschimmer</emphasis>
|
|||
|
|
” [= Pronotum bright green (f.
|
|||
|
|
<emphasis id="B92DEADCC154842FFB10F975FB5E8BA8" box="[1163,1235,1701,1727]" italics="true" pageId="4" pageNumber="155">viridis</emphasis>
|
|||
|
|
), sometimes with golden shimmer] which clarify that Petrovitz considered his f.[orm] as individual, and suggests that he was prone to apply the name “
|
|||
|
|
<emphasis id="B92DEADCC154842FFEEAF93DFE348A10" box="[369,441,1773,1799]" italics="true" pageId="4" pageNumber="155">viridis</emphasis>
|
|||
|
|
” to any green individual of
|
|||
|
|
<taxonomicName id="4C594D4DC154842FFD69F93DFC0D8A10" box="[754,896,1773,1799]" class="Insecta" family="Glaphyridae" genus="Pygopleurus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B92DEADCC154842FFD69F93DFC0D8A10" box="[754,896,1773,1799]" italics="true" pageId="4" pageNumber="155">Pygopleurus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
not matching the color of the
|
|||
|
|
<typeStatus id="54E2886CC154842FFB54F93EFA8F8A1F" box="[1231,1282,1774,1800]" pageId="4" pageNumber="155">type</typeStatus>
|
|||
|
|
, regardless of species.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BE636CEC154842FFF0CF889FEF38A63" blockId="4.[151,1436,1881,2016]" box="[151,382,1881,1908]" pageId="4" pageNumber="155">
|
|||
|
|
<heading id="D0AE81A2C154842FFF0CF889FEF38A63" bold="true" box="[151,382,1881,1908]" fontSize="11" level="1" pageId="4" pageNumber="155" reason="1">
|
|||
|
|
<emphasis id="B92DEADCC154842FFF0CF889FEF38A63" bold="true" box="[151,382,1881,1908]" pageId="4" pageNumber="155">Taxonomic account</emphasis>
|
|||
|
|
</heading>
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BE636CEC154842FFF5CF8ADFF638AAC" blockId="4.[151,1436,1881,2016]" pageId="4" pageNumber="155">
|
|||
|
|
Our study led to the identification of ten species and one subspecies of
|
|||
|
|
<taxonomicName id="4C594D4DC154842FFB85F8ADFB218A80" box="[1054,1196,1917,1943]" class="Insecta" family="Glaphyridae" genus="Pygopleurus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B92DEADCC154842FFB85F8ADFB218A80" box="[1054,1196,1917,1943]" italics="true" pageId="4" pageNumber="155">Pygopleurus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
inhabiting mainland
|
|||
|
|
<collectingCountry id="F34E765EC154842FFF0CF871FF678AAC" box="[151,234,1953,1979]" name="Greece" pageId="4" pageNumber="155">Greece</collectingCountry>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8BE636CEC154842EFF5CF815FB1E8C08" blockId="4.[151,1436,1881,2016]" lastBlockId="5.[151,1437,153,359]" lastPageId="5" lastPageNumber="156" pageId="4" pageNumber="155">
|
|||
|
|
The arrangement of species we follow does not conform to the informal groups I–IV introduced by
|
|||
|
|
<bibRefCitation id="EFC84B3FC154842EFAD0F815FF698DA4" author="Baraud, J." lastPageId="5" lastPageNumber="156" pageId="4" pageNumber="155" pagination="331 - 375" refId="ref29740" refString="Baraud, J. (1989) Revision du genre Pygopleurus Motschulsky (Coleoptera, Scarabaoidea: Glaphyridae). Annales de la Societe Entomologique de France, 25 (3), 331 - 375, 89 figs." type="book chapter" year="1989">Baraud (1989)</bibRefCitation>
|
|||
|
|
, as we consider them non-natural. Based on the morphological similarity of external morphology, paramera and everted endophallus, we formally introduce two species-groups that seem well supported with respect to the whole genus
|
|||
|
|
<taxonomicName id="4C594D4DC155842EFEB7FF31FE378DEC" box="[300,442,225,251]" class="Insecta" family="Glaphyridae" genus="Pygopleurus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B92DEADCC155842EFEB7FF31FE378DEC" box="[300,442,225,251]" italics="true" pageId="5" pageNumber="156">Pygopleurus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. The other species here considered are not assigned to groups, and appear to loosely related, with the only exception of
|
|||
|
|
<taxonomicName id="4C594D4DC155842EFD92FED6FD268C08" baseAuthorityName="Brulle" baseAuthorityYear="1832" box="[521,683,261,287]" class="Insecta" family="Glaphyridae" genus="Pygopleurus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="chrysonotus">
|
|||
|
|
<emphasis id="B92DEADCC155842EFD92FED6FD268C08" box="[521,683,261,287]" italics="true" pageId="5" pageNumber="156">P. chrysonotus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C594D4DC155842EFD79FED6FCF88C08" baseAuthorityName="Brulle" baseAuthorityYear="1832" box="[738,885,261,287]" class="Insecta" family="Glaphyridae" genus="Pygopleurus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="scutellatus">
|
|||
|
|
<emphasis id="B92DEADCC155842EFD79FED6FCF88C08" box="[738,885,261,287]" italics="true" pageId="5" pageNumber="156">P. scutellatus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
, very close to each other.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3436545C155842EFF5CFEF9FCFD8C70" pageId="5" pageNumber="156" type="synonymic_list">
|
|||
|
|
<paragraph id="8BE636CEC155842EFF5CFEF9FCFD8C70" blockId="5.[151,1437,153,359]" pageId="5" pageNumber="156">
|
|||
|
|
A checklist of species and species-groups, with corresponding old names, is presented in
|
|||
|
|
<tableCitation id="C6DB0375C155842EFB5EFEF9FAB68C54" box="[1221,1339,297,323]" captionStart="TABLE 1" captionStartId="5.[151,244,404,431]" captionTargetBox="[186,1304,482,627]" captionTargetPageId="5" captionText="TABLE 1. Checklist of Pygopleurus species and subspecies here recognised for mainland Greece, and previous treatment for each taxon." httpUri="http://table.plazi.org/id/DF266646C155842EFF0CFE44FE3D8CC4" pageId="5" pageNumber="156" tableUuid="DF266646C155842EFF0CFE44FE3D8CC4">Table 1. A</tableCitation>
|
|||
|
|
detailed taxonomic account follows, with species arranged as in the table.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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