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<document id="B30769CCEE7566B4504B0CE43352420F" ID-DOI="10.5281/zenodo.199397" ID-GBIF-Dataset="1e9c7a43-a511-43dd-99f6-9ffbe683245f" ID-ISSN="1175-5326" ID-Zenodo-Dep="199397" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460335655840" checkinUser="plazi" docAuthor="Zhang, Xu &amp; Jin, Dao-Chao" docDate="2010" docId="E52C87F0527DFFC822F1E72B21DA2436" docLanguage="en" docName="zt02684p024.pdf" docOrigin="Zootaxa 2684" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Sperchon (Hispidosperchon) beijingensis Zhang &amp; Jin, 2010, sp. nov." docType="treatment" docVersion="9" lastPageNumber="17" masterDocId="1915FF88527CFFCC2266E54124462514" masterDocTitle="Three new species and one new record of the subgenus Hispidosperchon Thor, 1901 within the genus Sperchon Kramer, 1877 from China (Acari: Sperchontidae)" masterLastPageNumber="24" masterPageNumber="14" pageNumber="15" updateTime="1698259716482" updateUser="plazi">
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<mods:title id="2C14D9585166535ABBF79E923A634979">Three new species and one new record of the subgenus Hispidosperchon Thor, 1901 within the genus Sperchon Kramer, 1877 from China (Acari: Sperchontidae)</mods:title>
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<mods:name id="8CBC3D4CAC03669905080570054523F4" type="personal">
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<mods:namePart id="720EAF94D17643CFC0EE89A980DDBB17">Zhang, Xu</mods:namePart>
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<mods:namePart id="17C8CBB34D863CC5734681386731ACF4">Jin, Dao-Chao</mods:namePart>
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<mods:date id="ABBD22F429FFB1A08A7743952E79C7D3">2010</mods:date>
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<treatment id="E52C87F0527DFFC822F1E72B21DA2436" ID-DOI="http://doi.org/10.5281/zenodo.6209465" ID-GBIF-Taxon="119405287" ID-Zenodo-Dep="6209465" LSID="urn:lsid:plazi:treatment:E52C87F0527DFFC822F1E72B21DA2436" httpUri="http://treatment.plazi.org/id/E52C87F0527DFFC822F1E72B21DA2436" lastPageId="4" lastPageNumber="17" pageId="1" pageNumber="15">
<subSubSection id="259F656D527DFFCD22F1E72B251A27B3" pageId="1" pageNumber="15" type="nomenclature">
<paragraph id="6D3A36E6527DFFCD22F1E72B26AB2790" blockId="1.[151,749,618,679]" box="[151,749,618,644]" pageId="1" pageNumber="15">
<heading id="3672818A527DFFCD22F1E72B26AB2790" bold="true" box="[151,749,618,644]" fontSize="11" level="1" pageId="1" pageNumber="15" reason="1">
<emphasis id="5FF1EAF4527DFFCD22F1E72B26AB2790" bold="true" box="[151,749,618,644]" pageId="1" pageNumber="15">
<taxonomicName id="AA854D65527DFFCD22F1E72B26CF2790" box="[151,649,618,644]" class="Arachnida" family="Sperchonidae" genus="Sperchon" kingdom="Animalia" order="Trombidiformes" pageId="1" pageNumber="15" phylum="Arthropoda" rank="species" species="beijingensis" status="sp. nov." subGenus="Hispidosperchon">
<emphasis id="5FF1EAF4527DFFCD22F1E72B26CF2790" bold="true" box="[151,649,618,644]" italics="true" pageId="1" pageNumber="15">Sperchon (Hispidosperchon) beijingensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="44C2578F527DFFCD20F6E72B26AB2790" box="[656,749,618,644]" pageId="1" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="6D3A36E6527DFFCD22F8E7CC251A27B3" blockId="1.[151,749,618,679]" box="[158,348,653,679]" pageId="1" pageNumber="15">Figure A (110)</paragraph>
</subSubSection>
<subSubSection id="259F656D527DFFCD22F1E79625432628" pageId="1" pageNumber="15" type="materials_examined">
<paragraph id="6D3A36E6527DFFCD22F1E79625432628" blockId="1.[151,1437,727,940]" pageId="1" pageNumber="15">
<emphasis id="5FF1EAF4527DFFCD22F1E796256E27E5" bold="true" box="[151,296,727,753]" pageId="1" pageNumber="15">
<typeStatus id="B23E8844527DFFCD22F1E796249227E5" box="[151,212,727,753]" pageId="1" pageNumber="15">Type</typeStatus>
series.
</emphasis>
<typeStatus id="B23E8844527DFFCD2349E79925E727E6" box="[303,417,728,754]" pageId="1" pageNumber="15" type="holotype">Holotype</typeStatus>
: Male,
<collectingCountry id="15927676527DFFCD2390E799267927E6" box="[502,575,728,754]" name="China" pageId="1" pageNumber="15">China</collectingCountry>
, Beijing City, unnamed stream (
<geoCoordinate id="08B15021527DFFCD21D0E799200427E6" box="[950,1090,728,754]" direction="north" orientation="latitude" pageId="1" pageNumber="15" precision="15" value="35.86139">N35°51'41''</geoCoordinate>
,
<geoCoordinate id="08B15021527DFFCD2628E79920A227E6" box="[1102,1252,728,754]" direction="east" orientation="longitude" pageId="1" pageNumber="15" precision="15" value="104.19528">E104°11'43''</geoCoordinate>
),
<date id="193B1026527DFFCD269FE79921DE27E6" box="[1273,1432,728,754]" pageId="1" pageNumber="15" value="2002-05-12">12 May 2002</date>
, leg. Jian-Jun Guo.
<typeStatus id="B23E8844527DFFCD23E5E7BC25B22603" box="[387,500,765,791]" pageId="1" pageNumber="15" type="paratype">Paratype</typeStatus>
: Two females, the same data as the
<typeStatus id="B23E8844527DFFCD21D3E7BC20602603" box="[949,1062,765,791]" pageId="1" pageNumber="15" type="holotype">holotype</typeStatus>
. All were dissected and slidemounted.
</paragraph>
</subSubSection>
<subSubSection id="259F656D527DFFCD22A3E60625672201" pageId="1" pageNumber="15" type="diagnosis">
<paragraph id="6D3A36E6527DFFCD22A3E606274426B8" blockId="1.[151,1437,727,940]" pageId="1" pageNumber="15">
<emphasis id="5FF1EAF4527DFFCD22A3E60625032675" bold="true" box="[197,325,839,865]" pageId="1" pageNumber="15">Diagnosis.</emphasis>
A1 short and plumose; E4 near to anteromedial margin of CxIII; the second acetabulum near to the third but far away from the first; P-II with a long ventro-distal projection; P-IV with two very small peglike setae on the ventral side; excretory pore smooth.
</paragraph>
<paragraph id="6D3A36E6527DFFCD22F1E39F25672201" blockId="1.[151,1436,1758,1813]" pageId="1" pageNumber="15">
<emphasis id="5FF1EAF4527DFFCD22F1E39F252E23E2" bold="true" box="[151,360,1758,1782]" pageId="1" pageNumber="15">FIGURE A (12).</emphasis>
<taxonomicName id="AA854D65527DFFCD2309E39E277C23E2" box="[367,826,1758,1782]" class="Arachnida" family="Sperchonidae" genus="Sperchon" kingdom="Animalia" order="Trombidiformes" pageId="1" pageNumber="15" phylum="Arthropoda" rank="species" species="beijingensis" status="sp. nov." subGenus="Hispidosperchon">
<emphasis id="5FF1EAF4527DFFCD2309E39E259023E2" box="[367,470,1759,1782]" italics="true" pageId="1" pageNumber="15">Sperchon</emphasis>
(
<emphasis id="5FF1EAF4527DFFCD238EE39E26E223E2" box="[488,676,1759,1782]" italics="true" pageId="1" pageNumber="15">Hispidosperchon</emphasis>
)
<emphasis id="5FF1EAF4527DFFCD20D3E39E277C23E2" box="[693,826,1759,1782]" italics="true" pageId="1" pageNumber="15">beijingensis</emphasis>
</taxonomicName>
<emphasis id="5FF1EAF4527DFFCD2127E39F27DD23E2" bold="true" box="[833,923,1758,1782]" pageId="1" pageNumber="15">
<taxonomicNameLabel id="44C2578F527DFFCD2127E39F27DD23E2" box="[833,923,1758,1782]" pageId="1" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, Male. 1. idiosoma, dorsal view; 2. idiosoma, ventral view.
</paragraph>
</subSubSection>
<subSubSection id="259F656D527DFFCF22A3E202256723B2" lastPageId="3" lastPageNumber="17" pageId="1" pageNumber="15" type="description">
<paragraph id="6D3A36E6527DFFCE22A3E202267E237D" blockId="1.[151,1436,1859,2035]" lastBlockId="2.[151,1435,1586,1641]" lastPageId="2" lastPageNumber="16" pageId="1" pageNumber="15">
<emphasis id="5FF1EAF4527DFFCD22A3E20225EB2249" bold="true" box="[197,429,1859,1885]" pageId="1" pageNumber="15">Description. Male:</emphasis>
Body oval in outline,
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length,
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width. Cuticle yellow-brown, covered with very fine spinules arranged in hexagonal pattern (Fig. A-3). A1 short and plumose, other dorsal setae thin and long. Chitinous plates and glandular plates on both dorsum and venter were shown in Fig. A-1 and Fig. A-2, one pair of chitinous plates between D1 and D2 on dorsum fused somewhat. Coxae in four groups. ACG
<quantity id="AA7D9B03527DFFCD2716E2F524E822E7" metricMagnitude="0" metricUnit="m" metricValue="5.2832" pageId="1" pageNumber="15" unit="in" value="208.0">208 in</quantity>
length, close to each other but not fused, anterior apodeme well developed. E2 on the lateral interval
<emphasis id="5FF1EAF4527EFFCE22F1E3732522235E" bold="true" box="[151,356,1586,1610]" pageId="2" pageNumber="16">FIGURE A (38).</emphasis>
<taxonomicName id="AA854D65527EFFCE230DE373276F235D" box="[363,809,1586,1609]" class="Arachnida" family="Sperchonidae" genus="Sperchon" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="16" phylum="Arthropoda" rank="species" species="beijingensis" status="sp. nov." subGenus="Hispidosperchon">
<emphasis id="5FF1EAF4527EFFCE230DE3732596235D" box="[363,464,1586,1609]" italics="true" pageId="2" pageNumber="16">Sperchon</emphasis>
(
<emphasis id="5FF1EAF4527EFFCE2386E37326D1235D" box="[480,663,1586,1609]" italics="true" pageId="2" pageNumber="16">Hispidosperchon</emphasis>
)
<emphasis id="5FF1EAF4527EFFCE20CEE373276F235D" box="[680,809,1586,1609]" italics="true" pageId="2" pageNumber="16">beijingensis</emphasis>
</taxonomicName>
<emphasis id="5FF1EAF4527EFFCE2157E37327CE235E" bold="true" box="[817,904,1586,1610]" pageId="2" pageNumber="16">
<taxonomicNameLabel id="44C2578F527EFFCE2157E37327CE235E" box="[817,904,1586,1610]" pageId="2" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, Male. 3. decorations of cuticle; 4. capitulum; 5. chelicera; 6. palp; 7. claw; 8. IV-L-2-6.
</paragraph>
<paragraph id="6D3A36E6527EFFCF22F1E3D9202B24C8" blockId="2.[151,1436,1688,2012]" lastBlockId="3.[151,1437,152,775]" lastPageId="3" lastPageNumber="17" pageId="2" pageNumber="16">
between ACG and PCG. PCG
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length, widely separated. E4 near to anteromedial margin of CxIII. Distance between anterior end of ACG and posterior end of PCG 385. Surface of coxae reticulated. Approximately two-thirds of genital field between PCG. Genital valves not covering the genital acetabula,
<quantity id="AA7D9B03527EFFCE2716E3A324E82236" metricMagnitude="0" metricUnit="m" metricValue="4.572" pageId="2" pageNumber="16" unit="in" value="180.0">180 in</quantity>
length,
<quantity id="AA7D9B03527EFFCE236AE24925112236" box="[268,343,1800,1826]" metricMagnitude="0" metricUnit="m" metricValue="4.0386" pageId="2" pageNumber="16" unit="in" value="159.0">159 in</quantity>
width. One round platelet in front of genital field. Pre- and postgentital sclerites small. Three pairs of small and rounded genital acetabula, the second acetabulum near to the third but far away from the first. A relatively small and rounded platelet behind genital field.
<date id="193B1026527EFFCE21A6E21327A32278" box="[960,997,1874,1900]" pageId="2" pageNumber="16">V1</date>
on very small sclerites and without accompanying glandularia. Excretory pore smooth. Gnathosoma with long rostrum, length 220. Chelicera total length 232, basal segment length 164, claw length 68, ratio of basal segment/claw length 2.4. Dorsal lengths of the palpal segments: P-I, 32; P-II, 140; P-III, 173; P-IV, 201; P-V, 41. P-I without seta. P-II with long ventro-distal projection bearing two setae, of which one shorter than another one. About twelve setae on the lateral and dorsal side of P-II, and six of them plumose. The ventral side of P-III straight and without seta, ten smooth setae on the lateral and the dorsal. The venter of P-IV with two small peg-like setae and two normal thin setae, the proximal peg-like seta on the middle and the distal one near to terminal end of the segment. The dorsal of P-IV with three thin seta, two of them located distally. Dorsal lengths of leg I: I-L-1, 60; I-L-2, 75; I-L-3, 100; I-L-4, 150; I-L-5, 155; I-L-6, 140. Dorsal lengths of leg IV (the first segment missing): IV-L-2, 163; IV-L-3, 149; IV-L-4, 269; IV-L-5, 265; IV-L-6, 225. Leg IV with a few short setae, plumose two of them on distal end of IV-L-3 and IV-L-4 separately. Ambulacrum with two claws. Each claw with claw blade well protruding, bearing a long dorsal and a shorter ventral clawlet.
</paragraph>
<paragraph id="6D3A36E6527FFFCF22A3E4A620E22613" blockId="3.[151,1437,152,775]" pageId="3" pageNumber="17">
<emphasis id="5FF1EAF4527FFFCF22A3E4A6256D2715" bold="true" box="[197,299,487,513]" pageId="3" pageNumber="17">Female:</emphasis>
Similar to male except the characteristics of genital field and the size of dorsalia and ventralia (Fig. A-9 and Fig. A-10). Body
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length,
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width. ACG
<quantity id="AA7D9B03527FFFCF21CEE74C27B42733" box="[936,1010,525,551]" metricMagnitude="0" metricUnit="m" metricValue="6.4008" pageId="3" pageNumber="17" unit="in" value="252.0">252 in</quantity>
length, with well-develop apodeme. PCG
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length. Distance between anterior end of ACG and posterior end of PCG 483. Gential field
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length,
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width. Pregentital sclerite well developed. The platelet behind genital field small and oblong. Gnathosoma length 270. Chelicera total length 300, basal segment length 210, claw length 210, ratio of basal segment/claw length 2.3. Dorsal lengths of the palpal segments: P-I, 32; P-II, 174; P-III, 154; P-IV, 267; P-V, 47. Dorsal lengths of leg I: I-L-1, 59; I-L-2, 67; I-L-3, 103; I-L-4, 184; I-L-5, 109; I-L-6, 132. Dorsal lengths of leg IV: IV-L-1, 121; IV-L-2, 124; IV-L-3, 149; IV-L-4, 317; IV-L-5, 278; IV-L-6, 194.
</paragraph>
<paragraph id="6D3A36E6527FFFCF22F1E32E256723B2" blockId="3.[151,1436,1647,1702]" pageId="3" pageNumber="17">
<emphasis id="5FF1EAF4527FFFCF22F1E32E25282393" bold="true" box="[151,366,1647,1671]" pageId="3" pageNumber="17">FIGURE A (910).</emphasis>
<taxonomicName id="AA854D65527FFFCF2313E32E276B2392" box="[373,813,1647,1670]" class="Arachnida" family="Sperchonidae" genus="Sperchon" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="17" phylum="Arthropoda" rank="species" species="beijingensis" status="sp. nov." subGenus="Hispidosperchon">
<emphasis id="5FF1EAF4527FFFCF2313E32E259F2392" box="[373,473,1647,1670]" italics="true" pageId="3" pageNumber="17">Sperchon</emphasis>
(
<emphasis id="5FF1EAF4527FFFCF238FE32E26D82392" box="[489,670,1647,1670]" italics="true" pageId="3" pageNumber="17">Hispidosperchon</emphasis>
)
<emphasis id="5FF1EAF4527FFFCF20C8E32E276B2392" box="[686,813,1647,1670]" italics="true" pageId="3" pageNumber="17">beijingensis</emphasis>
</taxonomicName>
<emphasis id="5FF1EAF4527FFFCF2153E32E27CD2393" bold="true" box="[821,907,1647,1671]" pageId="3" pageNumber="17">
<taxonomicNameLabel id="44C2578F527FFFCF2153E32E27CD2393" box="[821,907,1647,1671]" pageId="3" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, Female. 9. idiosoma, dorsal view; 10. idiosoma, ventral view.
</paragraph>
</subSubSection>
<subSubSection id="259F656D527FFFCF22A3E395204423FA" box="[197,1026,1748,1774]" pageId="3" pageNumber="17" type="etymology">
<paragraph id="6D3A36E6527FFFCF22A3E395204423FA" blockId="3.[151,1436,1748,2036]" box="[197,1026,1748,1774]" pageId="3" pageNumber="17">
<emphasis id="5FF1EAF4527FFFCF22A3E395251723FA" bold="true" box="[197,337,1748,1774]" pageId="3" pageNumber="17">Etymology.</emphasis>
The species is named after the city where it was collected.
</paragraph>
</subSubSection>
<subSubSection id="259F656D527FFFC822A3E3B821DA2436" lastPageId="4" lastPageNumber="18" pageId="3" pageNumber="17" type="discussion">
<paragraph id="6D3A36E6527FFFC822A3E3B821DA2436" blockId="3.[151,1436,1748,2036]" lastBlockId="4.[151,1437,152,290]" lastPageId="4" lastPageNumber="18" pageId="3" pageNumber="17">
<emphasis id="5FF1EAF4527FFFCF22A3E3B825782207" bold="true" box="[197,318,1785,1811]" pageId="3" pageNumber="17">Remarks.</emphasis>
The new species is closely related to
<taxonomicName id="AA854D65527FFFCF209FE3BB21112200" authority="Koenike, 1895 (Tuzovskij 2010)" authorityName="Koenike, 1895 (Tuzovskij" authorityYear="2010" box="[761,1367,1786,1812]" class="Arachnida" family="Sperchonidae" genus="Sperchon" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="17" phylum="Arthropoda" rank="species" species="hispidus">
<emphasis id="5FF1EAF4527FFFCF209FE3BB27892207" box="[761,975,1786,1811]" italics="true" pageId="3" pageNumber="17">Sperchon hispidus</emphasis>
Koenike, 1895 (
<bibRefCitation id="09144B17527FFFCF26F4E3BB21092200" author="Tuzovskij" box="[1170,1359,1786,1812]" pageId="3" pageNumber="17" refString="Tuzovskij, P. V. (2010) On diagnostics of the water mite Sperchon hispidus (Acariformes, Sperchontidae). Entomological Review, 90, 473 - 480." type="journal article" year="2010">Tuzovskij 2010</bibRefCitation>
)
</taxonomicName>
, from which it can be easily distinguished by the arrangement and the shape of the three acetabula. In
<taxonomicName id="AA854D65527FFFCF2775E26121D3222D" box="[1299,1429,1824,1849]" class="Arachnida" family="Sperchonidae" genus="Sperchon" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="17" phylum="Arthropoda" rank="species" species="hispidus">
<emphasis id="5FF1EAF4527FFFCF2775E26121D3222D" box="[1299,1429,1824,1849]" italics="true" pageId="3" pageNumber="17">S. hispidus</emphasis>
</taxonomicName>
, three pairs of acetabula arranged equidistantly, the anterior two pairs elliptic, and the posterior pair more or less rounded, whereas in the new species, the second pair near to the third but far away from the first, and all the three rounded and much smaller. Due to the form of genital field especially the arrangement of the three acetabula, the new species is also similar to
<taxonomicName id="AA854D65527FFFCF20CEE2F4206122DA" authority="Tuzovskij, 1990" authorityName="Tuzovskij" authorityYear="1990" box="[680,1063,1972,1998]" class="Arachnida" family="Sperchonidae" genus="Sperchon" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="17" phylum="Arthropoda" rank="species" species="prosperoides">
<emphasis id="5FF1EAF4527FFFCF20CEE2F4271922DA" box="[680,863,1973,1998]" italics="true" pageId="3" pageNumber="17">S. prosperoides</emphasis>
Tuzovskij, 1990
</taxonomicName>
from
<collectingCountry id="15927676527FFFCF2617E2F5208622DA" box="[1137,1216,1972,1998]" name="Russia" pageId="3" pageNumber="17">Russia</collectingCountry>
(
<bibRefCitation id="09144B17527FFFCF26B6E2F521C922DA" author="Tuzovskij" box="[1232,1423,1972,1998]" pageId="3" pageNumber="17" refString="Tuzovskij, P. V. (2008) The morphology of three species of adult water mites of the genus Sperchon (S. kuluensis, S. prosperoides, and S. orientalis, Acariformes, Sperchontidae). Entomological Review, 88, 139 - 149." type="journal article" year="2008">Tuzovskij 2008</bibRefCitation>
). However, in
<taxonomicName id="AA854D65527FFFCF234BE29B25A722E7" box="[301,481,2010,2035]" class="Arachnida" family="Sperchonidae" genus="Sperchon" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="17" phylum="Arthropoda" rank="species" species="prosperoides">
<emphasis id="5FF1EAF4527FFFCF234BE29B25A722E7" box="[301,481,2010,2035]" italics="true" pageId="3" pageNumber="17">S. prosperoides</emphasis>
</taxonomicName>
cuticle is covered with papillae and small ribs in various shapes, while in the new species cuticle is covered with very fine spinules arranged in hexagonal pattern which is typical of the subgenus
<taxonomicName id="AA854D655278FFC8236AE5FC259525C2" box="[268,467,189,214]" class="Arachnida" family="Sperchonidae" genus="Hispidosperchon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="18" phylum="Arthropoda" rank="genus">
<emphasis id="5FF1EAF45278FFC8236AE5FC259525C2" box="[268,467,189,214]" italics="true" pageId="4" pageNumber="18">Hispidosperchon</emphasis>
</taxonomicName>
. Further, P-III with two thick setae on the ventral side in
<taxonomicName id="AA854D655278FFC82611E5FC216D25C2" box="[1143,1323,189,214]" class="Arachnida" family="Sperchonidae" genus="Sperchon" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="18" phylum="Arthropoda" rank="species" species="prosperoides">
<emphasis id="5FF1EAF45278FFC82611E5FC216D25C2" box="[1143,1323,189,214]" italics="true" pageId="4" pageNumber="18">S. prosperoides</emphasis>
</taxonomicName>
, but with no setae in the new species. The locations of the two peg-like setae on ventral of P-IV are also different: distal one is almost at distal 1/3 of P- IV in
<taxonomicName id="AA854D655278FFC8202BE44927472435" box="[589,769,264,289]" class="Arachnida" family="Sperchonidae" genus="Sperchon" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="18" phylum="Arthropoda" rank="species" species="prosperoides">
<emphasis id="5FF1EAF45278FFC8202BE44927472435" box="[589,769,264,289]" italics="true" pageId="4" pageNumber="18">S. prosperoides</emphasis>
</taxonomicName>
, but near to distal end of the segment in the new species.
</paragraph>
</subSubSection>
</treatment>
</document>