<documentid="892BF3F2E1A11A030A54DC7F6FCE06F4"ID-CLB-Dataset="56065"ID-DOI="10.5852/ejt.2022.809.1719"ID-GBIF-Dataset="f76de735-01e3-454c-a7f1-bbafd26686f5"ID-ISSN="2118-9773"ID-Zenodo-Dep="6419425"ID-ZooBank="A754493E-5466-4479-B515-AABEDDE09D93"IM.bibliography_approvedBy="felipe"IM.materialsCitations_approvedBy="felipe"IM.metadata_approvedBy="felipe"IM.taxonomicNames_approvedBy="julia"checkinTime="1649273451180"checkinUser="felipe"docAuthor="Chigray, Svetlana N., Nabozhenko, Maxim V., Chigray, Ivan A. & Abakumov, Evgeny V."docDate="2022"docId="03FC86665576FF9B5079B834FD51F808"docLanguage="en"docName="ejt-809_1-71.pdf"docOrigin="European Journal of Taxonomy 809"docSource="http://dx.doi.org/10.5852/ejt.2022.809.1719"docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1"docStyleId="EF2B578F1D15862ADE45B0C07C620911"docStyleName="EJT.2018-.journal_article.type1"docStyleVersion="14"docTitle="Przewalskia Semenov 1893"docType="treatment"docVersion="10"lastPageNumber="30"masterDocId="FFC5FE1E556AFF865230BB19FFBAFF8B"masterDocTitle="A revision of the Palaearctic Pimeliini (Coleoptera: Tenebrionidae): a comparative analysis and systematic position of Eastern European and Asian taxa with dorso-lateral eyes"masterLastPageNumber="71"masterPageNumber="1"pageNumber="29"updateTime="1728593259121"updateUser="julia"zenodo-license-document="CC-BY-4.0"zenodo-license-figures="CC-BY-4.0"zenodo-license-treatments="UNSPECIFIED">
<mods:titleid="B9E522D6959777B0CE01F124DBAA79D1">A revision of the Palaearctic Pimeliini (Coleoptera: Tenebrionidae): a comparative analysis and systematic position of Eastern European and Asian taxa with dorso-lateral eyes</mods:title>
<mods:affiliationid="BD0B8391644A8F6FF47D86A0FF42AB91">St. Petersburg State University, 16 line of Vasilevskiy Island, 29, St. Petersburg 199178 Russia.</mods:affiliation>
<mods:affiliationid="DF2B18554D18AD2814146BDC9E103A7F">Precaspian Institute of Biological Resources of the Daghestan Federal Research Centre of the Russian Academy of Sciences, M. Gadzhiev str., 45, Makhachkala, Republic of Dagestan 367000 Russia. & Dagestan State University, M. Gadzhiev str., 43 a, Makhachkala, Republic of Dagestan 367000 Russia. & Severtsov Institute of ecology and evolution of the Russian Academy of sciences, Leninsky Prospect, 33, Moscow 119071 Russia.</mods:affiliation>
<mods:affiliationid="CD7B8C9D6D50E445E4E892BEB20621F4">Zoological Institute of the Russian Academy of Sciences, Universitetskaya Emb. 1, St. Petersburg 199034 Russia.</mods:affiliation>
<mods:affiliationid="488D65990A0074B162F685DA72BB48C1">St. Petersburg State University, 16 line of Vasilevskiy Island, 29, St. Petersburg 199178 Russia.</mods:affiliation>
<bibRefCitationid="EFC44A815576FF9A5106B834FC51FCCC"author="Semenov A. P."box="[822,1003,813,839]"pageId="28"pageNumber="29"pagination="1892 - 1893"refId="ref33186"refString="Semenov A. P. 1893. Symbolae ad cognitionem pimeliidarum. I - III. Horae Societatis Entomologicae Rossicae 27 [1892 - 1893]: 249 - 264."type="journal article"year="1893">Semenov, 1893</bibRefCitation>
<figureCitationid="136E2BF55576FF9A5031B849FD3EFCE1"box="[513,644,848,874]"captionStart="Fig"captionStartId="9.[189,232,1892,1918]"captionTargetBox="[233,1353,649,1847]"captionTargetId="figure-90@9.[227,1361,643,1855]"captionTargetPageId="9"captionText="Fig. 4. Species of the genera Przewalskia Semenov, 1893, Mantichorula Reitter, 1889 and Platyope Fischer von Waldheim, 1820, habitus. A. Przewalskia dilatata (Reitter, 1887), ♀ (ZIN), dorsal view. B. Ditto, ventral view. C. M. semenowi Reitter, 1889, ♂ (ZIN), dorsal view. D. Ditto, ventral view. E. Platyope leucogramma (Pallas, 1773), ♂ (ZIN), dorsal view. F. Ditto, ventral view."figureDoi="http://doi.org/10.5281/zenodo.6419435"httpUri="https://zenodo.org/record/6419435/files/figure.png"pageId="28"pageNumber="29">Figs 4A–B</figureCitation>
,
<figureCitationid="136E2BF55576FF9A50BFB849FD5DFCE1"box="[655,743,848,874]"captionStart="Fig"captionStartId="20.[189,232,1892,1918]"captionTargetBox="[189,1398,431,1851]"captionTargetId="figure-17@20.[189,1398,431,1853]"captionTargetPageId="20"captionText="Fig. 11. Przewalskia Semenov, 1893 and Mantichorula Reitter, 1889, details of structure. A. P. dilatata (Reitter, 1887), ♂ (ZIN), pronotum. B. Ditto, hemispherical large tubercles on pronotum. C. Ditto, base of left elytron. D. Ditto, pubescence of elytron. E. M. semenowi Reitter, 1889, ♂ (ZIN), pronotum. F. Ditto, tubercles and setation on sides of pronotum."figureDoi="http://doi.org/10.5281/zenodo.6419457"httpUri="https://zenodo.org/record/6419457/files/figure.png"pageId="28"pageNumber="29">11A–D</figureCitation>
,
<figureCitationid="136E2BF55576FF9A50C3B849FCA8FCE1"box="[755,786,848,874]"captionStart="Fig"captionStartId="39.[189,232,1892,1918]"captionTargetBox="[193,1392,716,1852]"captionTargetId="figure-98@39.[189,1398,704,1852]"captionTargetPageId="39"captionText="Fig. 18. Przewalskia dilatata (Reitter, 1887), genitalia (ZIN). A. Tegmen of aedeagus, ventrally.B. Ditto, dorsally. C. Ditto, laterally. D. Median lobe, ventrally. E. Ditto, dorsally. F. Ditto, laterally. G. Spiculum gastrale. H. ♂, inner sternite VIII.I. ♀, inner sternite VIII and spiculum ventrale.J. Ovipositor, ventrally. K. Ditto, dorsally. L. Ditto, laterally. Not to scale. Abbreviations: see Material and methods."figureDoi="http://doi.org/10.5281/zenodo.6419479"httpUri="https://zenodo.org/record/6419479/files/figure.png"pageId="28"pageNumber="29">18</figureCitation>
,
<figureCitationid="136E2BF55576FF9A512FB849FCEFFCE1"box="[799,853,848,874]"captionStart="Fig"captionStartId="46.[189,232,1822,1848]"captionTargetBox="[208,1379,488,1777]"captionTargetId="figure-18@46.[200,1387,478,1783]"captionTargetPageId="46"captionText="Fig. 23. Female genital ducts of Pimeliini. A. Apatopsis grombczewskii Semenov, 1891 (ZIN). B. Habrobates vernalis Semenov, 1903 (ZIN). C. Dietomorpha pardalis Kühnelt, 1957 (ZIN). D. Przewalskia dilatata (Reitter, 1887) (ZIN). E. Mantichorula semenowi Reitter, 1889 (ZIN). F. Platyope leucogramma (Pallas, 1773) (ZIN). G. Platyope grumi (Semenov, 1893), paralectotype (ZIN). H. Earophanta serrata (Semenov, 1893) (ZIN). Not to scale. Abbreviations: see Material and methods."figureDoi="http://doi.org/10.5281/zenodo.6419493"httpUri="https://zenodo.org/record/6419493/files/figure.png"pageId="28"pageNumber="29">23D</figureCitation>
,
<figureCitationid="136E2BF55576FF9A5150B849FC2EFCE1"box="[864,916,848,874]"captionStart="Fig"captionStartId="49.[188,231,1892,1918]"captionTargetBox="[191,1398,531,1851]"captionTargetId="figure-73@49.[189,1398,530,1855]"captionTargetPageId="49"captionText="Fig. 24. Apical antennomeres of different taxa of Pimeliini. A. Habrobates vernalis Semenov, 1903 ♂, paralectotype (ZIN). B. Przewalskia dilatata (Reitter, 1887), ♂. C. Mantichorula semenowi Reitter, 1889, ♂ (ZIN). D. Platyope leucogramma (Pallas, 1773), ♀. E. Earophanta serrata (Semenov, 1893), ♂ (ZIN)."figureDoi="http://doi.org/10.5281/zenodo.6419497"httpUri="https://zenodo.org/record/6419497/files/figure.png"pageId="28"pageNumber="29">24B</figureCitation>
,
<figureCitationid="136E2BF55576FF9A5190B849FC68FCE1"box="[928,978,848,874]"captionStart="Fig"captionStartId="52.[189,232,1822,1848]"captionTargetBox="[290,1286,773,1776]"captionTargetId="figure-71@52.[281,1307,763,1783]"captionTargetPageId="52"captionText="Fig. 26. Protibiae of Pimeliini, optic microscopy. A. Habrochiton vernus Semenov-Tjan-Shansky, 1907, ♂, lectotype (ZIN). B. Habrobates gabrieli (Schuster, 1935) comb. nov., paralectotype (NMB). C. Dietomorpha pardalis Kühnelt, 1957, ♂ (ZIN). D. Mantichorula semenowi Reitter, 1889, ♂ (ZIN). E. Przewalskia dilatata (Reitter, 1887), ♂ (ZIN). F. Apatopsis grombczewskii Semenov, 1891, ♀, paralectotype (ZIN). G. Earophanta serrata (Semenov, 1893), ♂ (ZIN). H. Platyope leucogramma (Pallas, 1773), ♂ (ZIN). Not to scale."figureDoi="http://doi.org/10.5281/zenodo.6419501"httpUri="https://zenodo.org/record/6419501/files/figure.png"pageId="28"pageNumber="29">26E</figureCitation>
,
<figureCitationid="136E2BF55576FF9A51EEB849FB88FCE1"box="[990,1074,848,874]"captionStart="Fig"captionStartId="55.[189,232,1892,1918]"captionTargetBox="[189,1398,503,1852]"captionTargetId="figure-12@55.[189,1398,503,1853]"captionTargetPageId="55"captionText="Fig. 28. Protibiae of Pimeliini, SEM. A. Przewalskia dilatata (Reitter, 1887), ♂ (ZIN), dorsal view. B. Ditto, ventral view. C. Earophanta serrata (Semenov, 1893), ♂ (ZIN), dorsal view. D. Platyope leucogramma (Pallas, 1773),♂ (ZIN), dorsal view.E. Platyope grumi(Semenov, 1893), ♂, paralectotype (ZIN), dorsal view."figureDoi="http://doi.org/10.5281/zenodo.6419508"httpUri="https://zenodo.org/record/6419508/files/figure.png"pageId="28"pageNumber="29">28A–B</figureCitation>
<bibRefCitationid="EFC44A815576FF9A5365B88BFDF3FC27"author="Semenov A. P."box="[341,585,914,940]"pageId="28"pageNumber="29"pagination="1892 - 1893"refId="ref33186"refString="Semenov A. P. 1893. Symbolae ad cognitionem pimeliidarum. I - III. Horae Societatis Entomologicae Rossicae 27 [1892 - 1893]: 249 - 264."type="journal article"year="1893">Semenov, 1893: 262</bibRefCitation>
<bibRefCitationid="EFC44A815576FF9A535AB8CAFD80FC66"author="Reitter E."box="[362,570,979,1005]"pageId="28"pageNumber="29"pagination="201 - 250"refId="ref32741"refString="Reitter E. 1893. Bestimmungs-Tabelle der unechten Pimeliden aus der palaearctischen Fauna. Verhandlungen des Naturforschenden Vereines in Brunn 31 (1892): 201 - 250."type="journal article"year="1893">Reitter 1893: 204</bibRefCitation>
</treatmentCitation>
, 249. —
<treatmentCitationid="0AF411615576FF9A5093B8CAFC36FC66"author="Semenov A. P."box="[675,908,979,1005]"page="11"pageId="28"pageNumber="29"year="1903">
<bibRefCitationid="EFC44A815576FF9A5093B8CAFC36FC66"author="Semenov A. P."box="[675,908,979,1005]"pageId="28"pageNumber="29"pagination="8 - 13"refId="ref33217"refString="Semenov A. P. 1903 a. Coleoptera novae faunae turanicae. Revue russe dEntomologie 3: 8 - 13."type="journal article"year="1903">Semenov 1903a: 11</bibRefCitation>
</treatmentCitation>
, nota 4. —
<treatmentCitationid="0AF411615576FF9A5623B8CAFEA3FB9B"author="Semenov-Tjan-Shansky A. P."page="177"pageId="28"pageNumber="29"year="1907">
<bibRefCitationid="EFC44A815576FF9A5623B8CAFEA3FB9B"author="Semenov-Tjan-Shansky A. P."pageId="28"pageNumber="29"pagination="1907 - 1908"refId="ref33261"refString="Semenov-Tjan-Shansky A. P. 1907. Synopsis generum tribus Platyopinorum (Coleoptera, Tenebrionidae Pimeliini). Horae Societatis Entomologicae Rossicae 38 [1907 - 1908]: 175 - 184."type="journal article"year="1907">Semenov-Tjan-Shansky 1907: 177</bibRefCitation>
<bibRefCitationid="EFC44A815576FF9A5379B8EFFDBAFB9B"author="Gebien H."box="[329,512,1014,1040]"pageId="28"pageNumber="29"pagination="1 - 3"refId="ref30595"refString="Gebien H. 1940. Die von Dr. Trinkler in Tibet gesammelten Tenebrioniden (Col. Tenebr.). Mitteilungen der Munchener Entomologischen Gesellschaft 30: 1 - 3."type="journal article"year="1940">Gebien 1940: 3</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitationid="0AF411615576FF9A5001B8EEFCF9FB9B"author="Ren G. - D. & Yu Y."box="[561,835,1014,1041]"page="67"pageId="28"pageNumber="29"year="1999">
<bibRefCitationid="EFC44A815576FF9A5001B8EEFCF9FB9B"author="Ren G. - D. & Yu Y."box="[561,835,1014,1041]"pageId="28"pageNumber="29"refId="ref32918"refString="Ren G. - D. & Yu Y. 1999. The Darkling Beetles from Deserts and Semideserts of China (Coleoptera: Tenebrionidae). Hebei University Publishing House, Baoding. [In Chinese.]"type="book"year="1999">Ren & Yu 1999: 67–68</bibRefCitation>
</treatmentCitation>
, fig. 24. —
<treatmentCitationid="0AF411615576FF9A51FDB8EEFB5FFB9B"author="Ren G. - D. & Dong S."box="[973,1253,1014,1041]"page="296"pageId="28"pageNumber="29"year="2001">
<bibRefCitationid="EFC44A815576FF9A51FDB8EEFB5FFB9B"author="Ren G. - D. & Dong S."box="[973,1253,1014,1041]"pageId="28"pageNumber="29"pagination="296 - 302"refId="ref32868"refString="Ren G. - D. & Dong S. 2001. The beetles of the tribe Platyopini in China and description of two new species (Coleoptera: Tenebrionidae). Journal of Hebei University (Natural Science Edition) 21 (3): 296 - 302."type="journal article"year="2001">Ren & Dong 2001: 296</bibRefCitation>
<bibRefCitationid="EFC44A815576FF9A53A2BF7FFD96FB0B"author="Reitter E."box="[402,556,1126,1152]"pageId="28"pageNumber="29"pagination="355 - 389"refId="ref32671"refString="Reitter E. 1887. Insecta in itinere Cl. N. Przewalskii in Asia Centrali novissime lecta. IX. Tenebrionidae. Horae Societatis Entomologicae Rossicae 21: 355 - 389."type="journal article"year="1887">Reitter, 1887</bibRefCitation>
<bibRefCitationid="EFC44A815576FF9A53AABFCFFD8DFB7B"author="Reitter E."box="[410,567,1238,1264]"pageId="28"pageNumber="29"pagination="355 - 389"refId="ref32671"refString="Reitter E. 1887. Insecta in itinere Cl. N. Przewalskii in Asia Centrali novissime lecta. IX. Tenebrionidae. Horae Societatis Entomologicae Rossicae 21: 355 - 389."type="journal article"year="1887">Reitter, 1887</bibRefCitation>
<bibRefCitationid="EFC44A815576FF9A50FBBFCFFCD2FB7B"author="Reitter E."box="[715,872,1238,1264]"pageId="28"pageNumber="29"pagination="355 - 389"refId="ref32671"refString="Reitter E. 1887. Insecta in itinere Cl. N. Przewalskii in Asia Centrali novissime lecta. IX. Tenebrionidae. Horae Societatis Entomologicae Rossicae 21: 355 - 389."type="journal article"year="1887">Reitter, 1887</bibRefCitation>
<bibRefCitationid="EFC44A815576FF9A5633BFCFFB1EFB7B"author="Gebien H."box="[1027,1188,1238,1264]"pageId="28"pageNumber="29"pagination="1 - 3"refId="ref30595"refString="Gebien H. 1940. Die von Dr. Trinkler in Tibet gesammelten Tenebrioniden (Col. Tenebr.). Mitteilungen der Munchener Entomologischen Gesellschaft 30: 1 - 3."type="journal article"year="1940">Gebien, 1940</bibRefCitation>
<figureCitationid="136E2BF55576FF9A533EBE5FFE35FAEB"box="[270,399,1350,1376]"captionStart="Fig"captionStartId="9.[189,232,1892,1918]"captionTargetBox="[233,1353,649,1847]"captionTargetId="figure-90@9.[227,1361,643,1855]"captionTargetPageId="9"captionText="Fig. 4. Species of the genera Przewalskia Semenov, 1893, Mantichorula Reitter, 1889 and Platyope Fischer von Waldheim, 1820, habitus. A. Przewalskia dilatata (Reitter, 1887), ♀ (ZIN), dorsal view. B. Ditto, ventral view. C. M. semenowi Reitter, 1889, ♂ (ZIN), dorsal view. D. Ditto, ventral view. E. Platyope leucogramma (Pallas, 1773), ♂ (ZIN), dorsal view. F. Ditto, ventral view."figureDoi="http://doi.org/10.5281/zenodo.6419435"httpUri="https://zenodo.org/record/6419435/files/figure.png"pageId="28"pageNumber="29">Fig. 4A–B</figureCitation>
) from pale brown to dark brown, robust, elytra rounded (semispherical), sometimes slightly flattened dorsally, completely covered with recumbent, white or yellowish and brown setae, with longitudinally striped pattern on elytra. Body length
Pronotum almost square (width subequal to length) (
<figureCitationid="136E2BF55576FF9A5102BEE8FC28F980"box="[818,914,1521,1547]"captionStart="Fig"captionStartId="8.[189,232,1892,1918]"captionTargetBox="[249,1340,656,1847]"captionTargetId="figure-122@8.[241,1346,643,1855]"captionTargetPageId="8"captionText="Fig. 3. Species of the genus Dietomorpha Kühnelt, 1957, habitus. A. D. pardalis Kühnelt, 1957, ♂ (ZIN), dorsal view. B. Ditto, ventral view. C. D. gonzalesi S. Chigray & Nabozhenko sp. nov., ♀, holotype (MNHP), dorsal view. D. Ditto, ventral view. E. Ditto, anterior view. F. Ditto, posterior view. G. Ditto, dorso-antero-lateral view. C–G: photographs by David Gonzales."figureDoi="http://doi.org/10.5281/zenodo.6419433"httpUri="https://zenodo.org/record/6419433/files/figure.png"pageId="28"pageNumber="29">Fig. 3A</figureCitation>
), with lateral margins straight or weakly emarginated basally. Anterolateral angles strongly projecting, acutely angulated (but rounded at apex), visibly raised above level of eyes. Disc of pronotum moderately convex, with rounded, deep depression in middle and two triangular, deep depressions at base, surface around depressions densely covered with very large semispherical tubercles with seta (
<figureCitationid="136E2BF55576FF9A5123BD67FC19F913"box="[787,931,1662,1688]"captionStart="Fig"captionStartId="20.[189,232,1892,1918]"captionTargetBox="[189,1398,431,1851]"captionTargetId="figure-17@20.[189,1398,431,1853]"captionTargetPageId="20"captionText="Fig. 11. Przewalskia Semenov, 1893 and Mantichorula Reitter, 1889, details of structure. A. P. dilatata (Reitter, 1887), ♂ (ZIN), pronotum. B. Ditto, hemispherical large tubercles on pronotum. C. Ditto, base of left elytron. D. Ditto, pubescence of elytron. E. M. semenowi Reitter, 1889, ♂ (ZIN), pronotum. F. Ditto, tubercles and setation on sides of pronotum."figureDoi="http://doi.org/10.5281/zenodo.6419457"httpUri="https://zenodo.org/record/6419457/files/figure.png"pageId="28"pageNumber="29">Fig. 11A–B</figureCitation>
), with setae between them. Prosternum long, 1.8× as short as longitudinal diameter of one procoxa (
<figureCitationid="136E2BF55576FF9A514EBDB8FC60F930"box="[894,986,1697,1723]"captionStart="Fig"captionStartId="9.[189,232,1892,1918]"captionTargetBox="[233,1353,649,1847]"captionTargetId="figure-90@9.[227,1361,643,1855]"captionTargetPageId="9"captionText="Fig. 4. Species of the genera Przewalskia Semenov, 1893, Mantichorula Reitter, 1889 and Platyope Fischer von Waldheim, 1820, habitus. A. Przewalskia dilatata (Reitter, 1887), ♀ (ZIN), dorsal view. B. Ditto, ventral view. C. M. semenowi Reitter, 1889, ♂ (ZIN), dorsal view. D. Ditto, ventral view. E. Platyope leucogramma (Pallas, 1773), ♂ (ZIN), dorsal view. F. Ditto, ventral view."figureDoi="http://doi.org/10.5281/zenodo.6419435"httpUri="https://zenodo.org/record/6419435/files/figure.png"pageId="28"pageNumber="29">Fig. 4B</figureCitation>
), without depression along anterior margin. Prosternal process narrower between middle level of coxae and strongly widened to apex (1.5 × as long as wide), not raised between procoxae and not protruding beyond procoxae.
Elytra semispherical (2 or more × as wide as pronotum), humeral rib weakly expressed, not strongly elevated, with row of sparse tubercles. Elytral pubescence has longitudinally striped pattern (from
<figureCitationid="136E2BF55576FF9A51B0BC76FC60F801"box="[896,986,1903,1930]"captionStart="Fig"captionStartId="9.[189,232,1892,1918]"captionTargetBox="[233,1353,649,1847]"captionTargetId="figure-90@9.[227,1361,643,1855]"captionTargetPageId="9"captionText="Fig. 4. Species of the genera Przewalskia Semenov, 1893, Mantichorula Reitter, 1889 and Platyope Fischer von Waldheim, 1820, habitus. A. Przewalskia dilatata (Reitter, 1887), ♀ (ZIN), dorsal view. B. Ditto, ventral view. C. M. semenowi Reitter, 1889, ♂ (ZIN), dorsal view. D. Ditto, ventral view. E. Platyope leucogramma (Pallas, 1773), ♂ (ZIN), dorsal view. F. Ditto, ventral view."figureDoi="http://doi.org/10.5281/zenodo.6419435"httpUri="https://zenodo.org/record/6419435/files/figure.png"pageId="28"pageNumber="29">Fig. 4A</figureCitation>
) as a result of presence of denser or sparser setae (
<figureCitationid="136E2BF55576FF9A536EBC8AFE56F826"box="[350,492,1939,1965]"captionStart="Fig"captionStartId="20.[189,232,1892,1918]"captionTargetBox="[189,1398,431,1851]"captionTargetId="figure-17@20.[189,1398,431,1853]"captionTargetPageId="20"captionText="Fig. 11. Przewalskia Semenov, 1893 and Mantichorula Reitter, 1889, details of structure. A. P. dilatata (Reitter, 1887), ♂ (ZIN), pronotum. B. Ditto, hemispherical large tubercles on pronotum. C. Ditto, base of left elytron. D. Ditto, pubescence of elytron. E. M. semenowi Reitter, 1889, ♂ (ZIN), pronotum. F. Ditto, tubercles and setation on sides of pronotum."figureDoi="http://doi.org/10.5281/zenodo.6419457"httpUri="https://zenodo.org/record/6419457/files/figure.png"pageId="28"pageNumber="29">Fig. 11C–D</figureCitation>
) to 4–6 longitudinal rows of very small granules with long seta in denser pubescent (light) striae. Scutellar shield not hidden under base of pronotum, widened to apex; surface around scutellar shield without depression. Transverse length of one metacoxa subequal to intercoxal process of abdominal ventrite 1.
At least pro- and mesotrochanters with very long setae. Meso- and metafemora curved along elytral lateral vertical side. Protibiae widened from proximal to distal part, weakly curved outward, with strong, short spines (on outer margin) and very dense, long setae (
<figureCitationid="136E2BF55577FF9B51D5BACEFBECFE7A"box="[997,1110,471,497]"captionStart="Fig"captionStartId="52.[189,232,1822,1848]"captionTargetBox="[290,1286,773,1776]"captionTargetId="figure-71@52.[281,1307,763,1783]"captionTargetPageId="52"captionText="Fig. 26. Protibiae of Pimeliini, optic microscopy. A. Habrochiton vernus Semenov-Tjan-Shansky, 1907, ♂, lectotype (ZIN). B. Habrobates gabrieli (Schuster, 1935) comb. nov., paralectotype (NMB). C. Dietomorpha pardalis Kühnelt, 1957, ♂ (ZIN). D. Mantichorula semenowi Reitter, 1889, ♂ (ZIN). E. Przewalskia dilatata (Reitter, 1887), ♂ (ZIN). F. Apatopsis grombczewskii Semenov, 1891, ♀, paralectotype (ZIN). G. Earophanta serrata (Semenov, 1893), ♂ (ZIN). H. Platyope leucogramma (Pallas, 1773), ♂ (ZIN). Not to scale."figureDoi="http://doi.org/10.5281/zenodo.6419501"httpUri="https://zenodo.org/record/6419501/files/figure.png"pageId="29"pageNumber="30">Figs 26E</figureCitation>
,
<figureCitationid="136E2BF55577FF9B5655BACEFB04FE7A"box="[1125,1214,471,497]"captionStart="Fig"captionStartId="55.[189,232,1892,1918]"captionTargetBox="[189,1398,503,1852]"captionTargetId="figure-12@55.[189,1398,503,1853]"captionTargetPageId="55"captionText="Fig. 28. Protibiae of Pimeliini, SEM. A. Przewalskia dilatata (Reitter, 1887), ♂ (ZIN), dorsal view. B. Ditto, ventral view. C. Earophanta serrata (Semenov, 1893), ♂ (ZIN), dorsal view. D. Platyope leucogramma (Pallas, 1773),♂ (ZIN), dorsal view.E. Platyope grumi(Semenov, 1893), ♂, paralectotype (ZIN), dorsal view."figureDoi="http://doi.org/10.5281/zenodo.6419508"httpUri="https://zenodo.org/record/6419508/files/figure.png"pageId="29"pageNumber="30">28A–B</figureCitation>
); spines denser at apex. Mesotibiae curved outward, with smaller and sparser spines and dense pubescence on inner margin. Metatibiae straight, with small, sparse spines and dense pubescence at least in apical part. Protibial terminal spurs usually lanceolate (at least outer spur), large, widened and flattened, slightly curved inward, subequal in length, extending to base of protarsomere 3; meso- and metatibial terminal spurs different in length, narrowed from base to apex, mesotibial terminal spurs extending to base of mesotarsomere 2, the latter ones extending to apex of metatarsomere 1. Pro- and mesotarsi flattened from lateral sides, covered with long fine setae dorsally and shorter, stronger setae ventrally; metatarsi also flattened from lateral sides, with long setae dorsally and very short ones ventrally; setae form dense, flattened brushes. Tarsal claws elongate, narrow, weakly curved outward.
<figureCitationid="136E2BF55577FF9B539FB84CFD84FCFB"box="[431,574,853,880]"captionStart="Fig"captionStartId="39.[189,232,1892,1918]"captionTargetBox="[193,1392,716,1852]"captionTargetId="figure-98@39.[189,1398,704,1852]"captionTargetPageId="39"captionText="Fig. 18. Przewalskia dilatata (Reitter, 1887), genitalia (ZIN). A. Tegmen of aedeagus, ventrally.B. Ditto, dorsally. C. Ditto, laterally. D. Median lobe, ventrally. E. Ditto, dorsally. F. Ditto, laterally. G. Spiculum gastrale. H. ♂, inner sternite VIII.I. ♀, inner sternite VIII and spiculum ventrale.J. Ovipositor, ventrally. K. Ditto, dorsally. L. Ditto, laterally. Not to scale. Abbreviations: see Material and methods."figureDoi="http://doi.org/10.5281/zenodo.6419479"httpUri="https://zenodo.org/record/6419479/files/figure.png"pageId="29"pageNumber="30">Fig. 18A–H</figureCitation>
<figureCitationid="136E2BF55577FF9B5398B860FDAEFC18"box="[424,532,889,915]"captionStart="Fig"captionStartId="39.[189,232,1892,1918]"captionTargetBox="[193,1392,716,1852]"captionTargetId="figure-98@39.[189,1398,704,1852]"captionTargetPageId="39"captionText="Fig. 18. Przewalskia dilatata (Reitter, 1887), genitalia (ZIN). A. Tegmen of aedeagus, ventrally.B. Ditto, dorsally. C. Ditto, laterally. D. Median lobe, ventrally. E. Ditto, dorsally. F. Ditto, laterally. G. Spiculum gastrale. H. ♂, inner sternite VIII.I. ♀, inner sternite VIII and spiculum ventrale.J. Ovipositor, ventrally. K. Ditto, dorsally. L. Ditto, laterally. Not to scale. Abbreviations: see Material and methods."figureDoi="http://doi.org/10.5281/zenodo.6419479"httpUri="https://zenodo.org/record/6419479/files/figure.png"pageId="29"pageNumber="30">Fig. 18H</figureCitation>
) weakly, widely sclerotized, except for narrow, memranous median part in middle, densely covered with long setae (setae much longer and denser along posterior margin); posterior margin with deep V-shaped median emargination; gland absent (or at least, not observed). Spiculum gastrale (
<figureCitationid="136E2BF55577FF9B5392B8FBFDB7FC77"box="[418,525,994,1020]"captionStart="Fig"captionStartId="39.[189,232,1892,1918]"captionTargetBox="[193,1392,716,1852]"captionTargetId="figure-98@39.[189,1398,704,1852]"captionTargetPageId="39"captionText="Fig. 18. Przewalskia dilatata (Reitter, 1887), genitalia (ZIN). A. Tegmen of aedeagus, ventrally.B. Ditto, dorsally. C. Ditto, laterally. D. Median lobe, ventrally. E. Ditto, dorsally. F. Ditto, laterally. G. Spiculum gastrale. H. ♂, inner sternite VIII.I. ♀, inner sternite VIII and spiculum ventrale.J. Ovipositor, ventrally. K. Ditto, dorsally. L. Ditto, laterally. Not to scale. Abbreviations: see Material and methods."figureDoi="http://doi.org/10.5281/zenodo.6419479"httpUri="https://zenodo.org/record/6419479/files/figure.png"pageId="29"pageNumber="30">Fig. 18G</figureCitation>
) without common stem, with weakly sclerotized membrane, rods roundly connected; derivatives of inner sternite IX elongate-oval, evenly sclerotized, with long setae apically.
<figureCitationid="136E2BF55577FF9B538BBF5EFDF2FBEA"box="[443,584,1095,1121]"captionStart="Fig"captionStartId="39.[189,232,1892,1918]"captionTargetBox="[193,1392,716,1852]"captionTargetId="figure-98@39.[189,1398,704,1852]"captionTargetPageId="39"captionText="Fig. 18. Przewalskia dilatata (Reitter, 1887), genitalia (ZIN). A. Tegmen of aedeagus, ventrally.B. Ditto, dorsally. C. Ditto, laterally. D. Median lobe, ventrally. E. Ditto, dorsally. F. Ditto, laterally. G. Spiculum gastrale. H. ♂, inner sternite VIII.I. ♀, inner sternite VIII and spiculum ventrale.J. Ovipositor, ventrally. K. Ditto, dorsally. L. Ditto, laterally. Not to scale. Abbreviations: see Material and methods."figureDoi="http://doi.org/10.5281/zenodo.6419479"httpUri="https://zenodo.org/record/6419479/files/figure.png"pageId="29"pageNumber="30">Fig. 18A–C</figureCitation>
) long, slender, with basal piece slightly shorter than apical one; apical piece bare, slightly curved, acutely angulate at apex ventral apophyses not expressed, dorsal apophyses long, merged in triangular plate; basal piece with narrowly rounded apex; median lobe with wide sclerotization in apical and basal parts and small membranous area in middle, base narrowly rounded, not bifurcate, apex acutely angulate (
<figureCitationid="136E2BF55577FF9B5040BFCDFD47FB65"box="[624,765,1236,1262]"captionStart="Fig"captionStartId="39.[189,232,1892,1918]"captionTargetBox="[193,1392,716,1852]"captionTargetId="figure-98@39.[189,1398,704,1852]"captionTargetPageId="39"captionText="Fig. 18. Przewalskia dilatata (Reitter, 1887), genitalia (ZIN). A. Tegmen of aedeagus, ventrally.B. Ditto, dorsally. C. Ditto, laterally. D. Median lobe, ventrally. E. Ditto, dorsally. F. Ditto, laterally. G. Spiculum gastrale. H. ♂, inner sternite VIII.I. ♀, inner sternite VIII and spiculum ventrale.J. Ovipositor, ventrally. K. Ditto, dorsally. L. Ditto, laterally. Not to scale. Abbreviations: see Material and methods."figureDoi="http://doi.org/10.5281/zenodo.6419479"httpUri="https://zenodo.org/record/6419479/files/figure.png"pageId="29"pageNumber="30">Fig. 18D–F</figureCitation>
<figureCitationid="136E2BF55577FF9B539BBE21FDB7FAD9"box="[427,525,1336,1362]"captionStart="Fig"captionStartId="39.[189,232,1892,1918]"captionTargetBox="[193,1392,716,1852]"captionTargetId="figure-98@39.[189,1398,704,1852]"captionTargetPageId="39"captionText="Fig. 18. Przewalskia dilatata (Reitter, 1887), genitalia (ZIN). A. Tegmen of aedeagus, ventrally.B. Ditto, dorsally. C. Ditto, laterally. D. Median lobe, ventrally. E. Ditto, dorsally. F. Ditto, laterally. G. Spiculum gastrale. H. ♂, inner sternite VIII.I. ♀, inner sternite VIII and spiculum ventrale.J. Ovipositor, ventrally. K. Ditto, dorsally. L. Ditto, laterally. Not to scale. Abbreviations: see Material and methods."figureDoi="http://doi.org/10.5281/zenodo.6419479"httpUri="https://zenodo.org/record/6419479/files/figure.png"pageId="29"pageNumber="30">Fig. 18I</figureCitation>
) with long common stem, rods narrow, anterior margin of sternite VIII nearly straight, with long, dense setae, apophyses of sternite VIII short, narrow, narrowly rounded at
<figureCitationid="136E2BF55577FF9B537CBED9FE75FA51"box="[332,463,1472,1498]"captionStart="Fig"captionStartId="39.[189,232,1892,1918]"captionTargetBox="[193,1392,716,1852]"captionTargetId="figure-98@39.[189,1398,704,1852]"captionTargetPageId="39"captionText="Fig. 18. Przewalskia dilatata (Reitter, 1887), genitalia (ZIN). A. Tegmen of aedeagus, ventrally.B. Ditto, dorsally. C. Ditto, laterally. D. Median lobe, ventrally. E. Ditto, dorsally. F. Ditto, laterally. G. Spiculum gastrale. H. ♂, inner sternite VIII.I. ♀, inner sternite VIII and spiculum ventrale.J. Ovipositor, ventrally. K. Ditto, dorsally. L. Ditto, laterally. Not to scale. Abbreviations: see Material and methods."figureDoi="http://doi.org/10.5281/zenodo.6419479"httpUri="https://zenodo.org/record/6419479/files/figure.png"pageId="29"pageNumber="30">Fig. 18J–L</figureCitation>
) moderately sclerotized, elongate. Coxite with four distinct, not merged lobes. Baculi of coxite lobe I well expressed, but much shorter than paraproct baculi; lobe II short, transverse; lobe III elongate; lobe IV strongly sclerotized, transformed into curved spatulate structure. Paraproct elongate, with narrow, long baculi. Proctiger also elongate and with narrow baculi, anterior margin rounded. Vulva with pair of sclerotized ‘spicula’ in the middle on ventral side. Paraproct in base, coxite and proctiger with very dense and long pubescence.
<figureCitationid="136E2BF55577FF9B53CFBDABFDD0F947"box="[511,618,1714,1740]"captionStart="Fig"captionStartId="46.[189,232,1822,1848]"captionTargetBox="[208,1379,488,1777]"captionTargetId="figure-18@46.[200,1387,478,1783]"captionTargetPageId="46"captionText="Fig. 23. Female genital ducts of Pimeliini. A. Apatopsis grombczewskii Semenov, 1891 (ZIN). B. Habrobates vernalis Semenov, 1903 (ZIN). C. Dietomorpha pardalis Kühnelt, 1957 (ZIN). D. Przewalskia dilatata (Reitter, 1887) (ZIN). E. Mantichorula semenowi Reitter, 1889 (ZIN). F. Platyope leucogramma (Pallas, 1773) (ZIN). G. Platyope grumi (Semenov, 1893), paralectotype (ZIN). H. Earophanta serrata (Semenov, 1893) (ZIN). Not to scale. Abbreviations: see Material and methods."figureDoi="http://doi.org/10.5281/zenodo.6419493"httpUri="https://zenodo.org/record/6419493/files/figure.png"pageId="29"pageNumber="30">Fig. 23D</figureCitation>
)
</paragraph>
<paragraphid="8BEA37705577FF9B528DBDCCFB9DF899"blockId="29.[189,1398,1713,1811]"pageId="29"pageNumber="30">Vagina elongated, with longitudinal wrinkles and separated anterior part. Spermatheca as short single tube. Accessory gland long, sacciform, with one thick constriction at base.</paragraph>