treatments-xml/data/03/EA/DA/03EADA36FFF6332A03E5F9F85E8AFEA2.xml

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<mods:title id="7B8CC23A0EC6346177F700DEC7880EFC">Revision of the new world species of Hylurgops LeConte, 1876 with the description of a new genus in the Hylastini (Coleoptera: Scolytinae) and comments on some Palearctic species</mods:title>
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<mods:namePart id="A033B663ECFAFC341E1C74C937AFA0B7">Mercado-Vélez, Javier E.</mods:namePart>
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<subSubSection id="C35938ABFFF6332903E5F9F85D36F9B4" pageId="20" pageNumber="321" type="nomenclature">
<paragraph id="8BFC6B20FFF6332903E5F9F85DE7F995" blockId="20.[151,678,1591,1651]" box="[151,678,1591,1618]" pageId="20" pageNumber="321">
<heading id="D0B4DC4CFFF6332903E5F9F85DE7F995" bold="true" box="[151,678,1591,1618]" fontSize="11" level="1" pageId="20" pageNumber="321" reason="1">
<taxonomicName id="4C4310A3FFF6332903E5F9F85DE7F995" ID-CoL="3NH24" authority="Mannerheim, 1843" authorityName="Mannerheim" authorityYear="1843" box="[151,678,1591,1618]" class="Insecta" family="Curculionidae" genus="Hylurgops" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="rugipennis">
<emphasis id="B937B732FFF6332903E5F9F85DE7F995" bold="true" box="[151,678,1591,1618]" pageId="20" pageNumber="321">
<emphasis id="B937B732FFF6332903E5F9F85EDEF996" bold="true" box="[151,415,1591,1617]" italics="true" pageId="20" pageNumber="321">Hylurgops rugipennis</emphasis>
(
<bibRefCitation id="EFD216D1FFF6332902DDF9F75DDFF995" author="Mannerheim" box="[431,670,1592,1618]" pageId="20" pageNumber="321" refString="Mannerheim, C. G. (1843) Beitrag zur kaefer-fauna der Aleutischen Inseln, der Insel Sitkha und Neu-Californiens. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 16, 3 - 142. http: // dx. doi. org / 10.5962 / bhl. title. 37833" type="journal article" year="1843">Mannerheim, 1843</bibRefCitation>
)
</emphasis>
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</paragraph>
<paragraph id="8BFC6B20FFF6332903E5F9955D36F9B4" blockId="20.[151,678,1591,1651]" box="[151,631,1626,1651]" pageId="20" pageNumber="321">
(
<figureCitation id="137877A5FFF6332903ECF9955E48F9B4" box="[158,265,1626,1651]" captionStart="FIGURE 5" captionStartId="6.[151,250,1892,1914]" captionTargetBox="[167,1425,1081,1859]" captionTargetId="figure@6.[158,1428,1078,1870]" captionTargetPageId="6" captionText="FIGURE 5. Depth of the mid-frontal impression of Hylurgops and Hylastes: a) deep and distinct (Hylurgops rugipennis), b) indistinct or absent (Hylastes mexicanus)." httpUri="https://zenodo.org/record/252619/files/figure.png" pageId="20" pageNumber="321">Figures 5</figureCitation>
a, 6d, 7c, 13b, 15c, 16a, 17c, 19)
</paragraph>
</subSubSection>
<subSubSection id="C35938ABFFF6332903E5F96D5D5EF8F4" pageId="20" pageNumber="321" type="reference_group">
<paragraph id="8BFC6B20FFF6332903E5F96D5D0AF932" blockId="20.[151,951,1698,1843]" pageId="20" pageNumber="321">
<treatmentCitationGroup id="AB534C0EFFF6332903E5F96D5D0AF932" pageId="20" pageNumber="321">
<taxonomicName id="4C4310A3FFF6332903E5F96D5D24F97F" ID-CoL="3NH3P" authority="Mannerheim, 1843:297" authorityName="Mannerheim" authorityPageNumber="297" authorityYear="1843" box="[151,613,1698,1720]" class="Insecta" family="Curculionidae" genus="Hylurgus" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="rugipennis">
<emphasis id="B937B732FFF6332903E5F96D5E30F97F" bold="true" box="[151,369,1698,1720]" italics="true" pageId="20" pageNumber="321">Hylurgus rugipennis</emphasis>
<treatmentCitation id="0AE24D31FFF633290204F96D5D24F97F" author="Mannerheim" box="[374,613,1698,1720]" page="297" pageId="20" pageNumber="321" year="1843">
<bibRefCitation id="EFD216D1FFF633290204F96D5D79F97F" author="Mannerheim" box="[374,568,1698,1720]" pageId="20" pageNumber="321" refString="Mannerheim, C. G. (1843) Beitrag zur kaefer-fauna der Aleutischen Inseln, der Insel Sitkha und Neu-Californiens. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 16, 3 - 142. http: // dx. doi. org / 10.5962 / bhl. title. 37833" type="journal article" year="1843">Mannerheim, 1843</bibRefCitation>
:297
</treatmentCitation>
</taxonomicName>
(Sitka Island, Alaska, 
<collectingCountry id="F3542BB0FFF633290038F96D5C3EF97F" box="[842,895,1698,1720]" name="United States of America" pageId="20" pageNumber="321">USA</collectingCountry>
, lost) 
<taxonomicName id="4C4310A3FFF6332903E5F90E5D20F910" authority="Mannerheim, 1852:385" authorityName="Mannerheim" authorityPageNumber="385" authorityYear="1852" box="[151,609,1729,1751]" class="Insecta" family="Curculionidae" genus="Hylastes" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="rugipennis">
<emphasis id="B937B732FFF6332903E5F90E5E27F910" bold="true" box="[151,358,1729,1751]" italics="true" pageId="20" pageNumber="321">Hylastes rugipennis</emphasis>
, 
<treatmentCitation id="0AE24D31FFF633290201F90E5D20F910" author="Mannerheim" box="[371,609,1729,1751]" page="385" pageId="20" pageNumber="321" year="1852">
<bibRefCitation id="EFD216D1FFF633290201F90E5D77F910" author="Mannerheim" box="[371,566,1729,1751]" pageId="20" pageNumber="321" refString="Mannerheim, C. G. (1852) Zweiter Nachtrag zur Kaefer-fauna der Nord-Amerikanischen Laender des Russischen Reiches. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 25, 284 - 387." type="journal article" year="1852">Mannerheim, 1852</bibRefCitation>
:385
</treatmentCitation>
</taxonomicName>
(not previously documented) 
<taxonomicName id="4C4310A3FFF6332903E5F9105D0AF932" ID-CoL="3NH24" authority="LeConte, 1876:390" authorityName="LeConte" authorityPageNumber="390" authorityYear="1876" box="[151,587,1759,1781]" class="Insecta" family="Curculionidae" genus="Hylurgops" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="rugipennis">
<emphasis id="B937B732FFF6332903E5F9105E3AF932" bold="true" box="[151,379,1759,1781]" italics="true" pageId="20" pageNumber="321">Hylurgops rugipennis</emphasis>
, 
<treatmentCitation id="0AE24D31FFF6332902FAF9105D0AF932" author="LeConte" box="[392,587,1759,1781]" page="390" pageId="20" pageNumber="321" year="1876">
<bibRefCitation id="EFD216D1FFF6332902FAF9105D61F932" author="LeConte" box="[392,544,1759,1781]" pageId="20" pageNumber="321" refString="LeConte, J. L. (1876) Family IX, Scolytidae. In: LeConte, J. L. &amp; Horn, G. H. (Eds.), The Rhynchophora of America North of Mexico. Proceedings of the American Philosophical Society, 15 (96), pp. 341 - 391." type="journal article" year="1876">LeConte, 1876</bibRefCitation>
:390
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</paragraph>
<paragraph id="8BFC6B20FFF6332903E5F9305D06F8D3" blockId="20.[151,951,1698,1843]" box="[151,583,1790,1812]" pageId="20" pageNumber="321">
<treatmentCitationGroup id="AB534C0EFFF6332903E5F9305D06F8D3" box="[151,583,1790,1812]" pageId="20" pageNumber="321">
<taxonomicName id="4C4310A3FFF6332903E5F9305D06F8D3" ID-CoL="3NH24" authority="Wood, 1982:89" authorityName="Wood" authorityPageNumber="89" authorityYear="1982" box="[151,583,1790,1812]" class="Insecta" family="Curculionidae" genus="Hylurgops" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="subSpecies" species="rugipennis" subSpecies="rugipennis">
<emphasis id="B937B732FFF6332903E5F9305EDFF8D3" bold="true" box="[151,414,1790,1812]" italics="true" pageId="20" pageNumber="321">H. rugipennis rugipennis</emphasis>
, 
<treatmentCitation id="0AE24D31FFF6332902D9F9315D06F8D3" author="Wood" box="[427,583,1790,1812]" page="89" pageId="20" pageNumber="321" year="1982">
<bibRefCitation id="EFD216D1FFF6332902D9F9315D68F8D3" author="Wood" box="[427,553,1790,1812]" pageId="20" pageNumber="321" refString="Wood, S. L. (1982) The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Naturalist Memoirs, 6, 1 - 1359." type="journal article" year="1982">Wood, 1982</bibRefCitation>
:89
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</taxonomicName>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFC6B20FFF6332903E5F8D25D5EF8F4" blockId="20.[151,951,1698,1843]" box="[151,543,1821,1843]" pageId="20" pageNumber="321">
<treatmentCitationGroup id="AB534C0EFFF6332903E5F8D25D5EF8F4" box="[151,543,1821,1843]" pageId="20" pageNumber="321">
<taxonomicName id="4C4310A3FFF6332903E5F8D25D5EF8F4" ID-CoL="8HZSG" authority="Wood, 1982:90" authorityName="Wood" authorityPageNumber="90" authorityYear="1982" box="[151,543,1821,1843]" class="Insecta" family="Curculionidae" genus="Hylurgops" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="subSpecies" species="rugipennis" subSpecies="pinifex">
<emphasis id="B937B732FFF6332903E5F8D25E36F8F4" bold="true" box="[151,375,1821,1843]" italics="true" pageId="20" pageNumber="321">H. rugipennis pinifex</emphasis>
, 
<treatmentCitation id="0AE24D31FFF6332902F1F8D25D5EF8F4" author="Wood" box="[387,543,1821,1843]" page="90" pageId="20" pageNumber="321" year="1982">
<bibRefCitation id="EFD216D1FFF6332902F1F8D25D41F8F4" author="Wood" box="[387,512,1821,1843]" pageId="20" pageNumber="321" refString="Wood, S. L. (1982) The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Naturalist Memoirs, 6, 1 - 1359." type="journal article" year="1982">Wood, 1982</bibRefCitation>
:90
</treatmentCitation>
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35938ABFFF6332903E5F8905A5CF823" pageId="20" pageNumber="321" type="diagnosis">
<paragraph id="8BFC6B20FFF6332903E5F8905A5CF823" blockId="20.[151,1436,1887,2020]" pageId="20" pageNumber="321">
<emphasis id="B937B732FFF6332903E5F8905E50F8BF" bold="true" box="[151,273,1887,1912]" pageId="20" pageNumber="321">Diagnosis.</emphasis>
<taxonomicName id="4C4310A3FFF633290268F8905D4EF8BF" box="[282,527,1887,1912]" class="Insecta" family="Curculionidae" genus="Hylurgops" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="rugipennis">
<emphasis id="B937B732FFF633290268F8905D4EF8BF" box="[282,527,1887,1912]" italics="true" pageId="20" pageNumber="321">Hylurgops rugipennis</emphasis>
</taxonomicName>
is reddish brown above, usually darker ventrally and its pronotum is broad and strongly constricted anteriorly (
<figureCitation id="137877A5FFF63329028BF84C5D0DF85B" box="[505,588,1923,1948]" captionStart="FIGURE 17" captionStartId="18.[151,250,1507,1529]" captionTargetBox="[165,1418,209,1484]" captionTargetId="figure@18.[160,1427,193,1485]" captionTargetPageId="18" captionText="FIGURE 17. Prothoraces of the New World Hylurgops: a) H. palliatus, b) H. planirostris, c) H. rugipennis, d) H. pinifex, e) H. incomptus, f) H. longipennis, g) H. knausi, h) H. reticulatus, and i) H. porosus." httpUri="https://zenodo.org/record/252631/files/figure.png" pageId="20" pageNumber="321">Fig. 17</figureCitation>
c). It is distinguished from 
<taxonomicName id="4C4310A3FFF63329000EF84A5CAAF85B" box="[892,1003,1924,1948]" class="Insecta" family="Curculionidae" genus="Hylurgops" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="321" phylum="Arthropoda" rank="species" species="pinifex">
<emphasis id="B937B732FFF63329000EF84A5CAAF85B" box="[892,1003,1924,1948]" italics="true" pageId="20" pageNumber="321">H. pinifex</emphasis>
</taxonomicName>
at&gt; 50× magnification by the distinct and complete pronotal reticulation (
<figureCitation id="137877A5FFF633290155F8685D39F807" box="[551,632,1959,1984]" captionStart="FIGURE 16" captionStartId="17.[151,250,1920,1942]" captionTargetBox="[151,1436,1224,1897]" captionTargetId="figure@17.[151,1436,1224,1898]" captionTargetPageId="17" captionText="FIGURE 16. Prothoracic discal surfaces of Hylurgops rugipennis and the eastern form of H. pinifex: a) reticulate and dull, sometimes shiny with similarly-sized punctures (H. rugipennis), b) shiny (eastern form), subgranulate (western form) with distinct, differently-sized punctures (H. pinifex)." httpUri="https://zenodo.org/record/252630/files/figure.png" pageId="20" pageNumber="321">Fig. 16</figureCitation>
a), by the regularly-sized pronotal punctures (
<figureCitation id="137877A5FFF633290705F8685B89F807" box="[1143,1224,1959,1984]" captionStart="FIGURE 16" captionStartId="17.[151,250,1920,1942]" captionTargetBox="[151,1436,1224,1897]" captionTargetId="figure@17.[151,1436,1224,1898]" captionTargetPageId="17" captionText="FIGURE 16. Prothoracic discal surfaces of Hylurgops rugipennis and the eastern form of H. pinifex: a) reticulate and dull, sometimes shiny with similarly-sized punctures (H. rugipennis), b) shiny (eastern form), subgranulate (western form) with distinct, differently-sized punctures (H. pinifex)." httpUri="https://zenodo.org/record/252630/files/figure.png" pageId="20" pageNumber="321">Fig. 16</figureCitation>
a), by the narrower metatibia, by the ventral lobe of the aedeagus (
<figureCitation id="137877A5FFF6332901D1F8045DA7F823" box="[675,742,1995,2020]" captionStart="FIGURE 7" captionStartId="9.[151,250,1919,1941]" captionTargetBox="[199,1387,194,1893]" captionTargetId="figure@9.[190,1396,193,1897]" captionTargetPageId="9" captionText="FIGURE 7. Aedeagus of the New World Hylurgops and Pachysquamus: a) Pachysquamus subcostulatus, b) Hylurgops planirostris, c) H. rugipennis, d) H. pinifex, e) H. incomptus, f) H. longipennis, g) H. knausi, h) H. reticulatus, and i) H. porosus." httpUri="https://zenodo.org/record/252621/files/figure.png" pageId="20" pageNumber="321">Fig. 7</figureCitation>
c), and by its more restricted distribution (
<figureCitation id="137877A5FFF6332907CFF8045A51F823" box="[1213,1296,1995,2020]" captionStart="FIGURE 19" captionStartId="21.[151,250,999,1021]" captionTargetBox="[153,1436,194,977]" captionTargetId="figure@21.[151,1436,193,978]" captionTargetPageId="21" captionText="FIGURE 19. Species distribution with main hosts: Hylurgops rugipennis (orange) and H. pinifex (black-gray). Arrows indicate sympatric areas where the two species and their respective main hosts co-occur." httpUri="https://zenodo.org/record/252633/files/figure.png" pageId="20" pageNumber="321">Fig. 19</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF3C3BA8FFF7332803E5FC285CFEFBDC" httpUri="https://zenodo.org/record/252633/files/figure.png" pageId="21" pageNumber="322" targetBox="[153,1436,194,977]" targetPageId="21">
<paragraph id="8BFC6B20FFF7332803E5FC285CFEFBDC" blockId="21.[151,1436,999,1051]" pageId="21" pageNumber="322">
<emphasis id="B937B732FFF7332803E5FC285E5EFC3B" bold="true" box="[151,287,999,1021]" pageId="21" pageNumber="322">FIGURE 19.</emphasis>
Species distribution with main hosts: 
<taxonomicName id="4C4310A3FFF7332801EDFC285C3AFC3A" box="[671,891,999,1021]" class="Insecta" family="Curculionidae" genus="Hylurgops" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="rugipennis">
<emphasis id="B937B732FFF7332801EDFC285C3AFC3A" box="[671,891,999,1021]" italics="true" pageId="21" pageNumber="322">Hylurgops rugipennis</emphasis>
</taxonomicName>
(orange) and 
<taxonomicName id="4C4310A3FFF733280775FC275B2BFC3A" box="[1031,1130,999,1021]" class="Insecta" family="Curculionidae" genus="Hylurgops" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="322" phylum="Arthropoda" rank="species" species="pinifex">
<emphasis id="B937B732FFF733280775FC275B2BFC3A" box="[1031,1130,999,1021]" italics="true" pageId="21" pageNumber="322">H. pinifex</emphasis>
</taxonomicName>
(black-gray). Arrows indicate sympatric areas where the two species and their respective main hosts co-occur.
</paragraph>
</caption>
<subSubSection id="C35938ABFFF7332B03B5FB875DD8FEDB" lastPageId="22" lastPageNumber="323" pageId="21" pageNumber="322" type="description">
<paragraph id="8BFC6B20FFF7332803B5FB875EA4F803" blockId="21.[151,1436,1096,2024]" pageId="21" pageNumber="322">
<emphasis id="B937B732FFF7332803B5FB875ED2FBA6" bold="true" box="[199,403,1096,1121]" pageId="21" pageNumber="322">Description. Size</emphasis>
. Length 3.7–4.7 (avg. 4.2 ± 0.3) mm long, 2.4× longer than wide. 
<emphasis id="B937B732FFF7332807C1FB875BB7FBA6" bold="true" box="[1203,1270,1096,1121]" pageId="21" pageNumber="322">Color</emphasis>
. Mature adult reddish to brownish-red with dark spots on anterior margin of pronotum, ventrally dark reddish-brown to nearly black, ventrites light reddish-brown. 
<emphasis id="B937B732FFF733280146FB5F5D38FB6E" bold="true" box="[564,633,1168,1193]" pageId="21" pageNumber="322">Frons</emphasis>
. Transverse impression strong (
<figureCitation id="137877A5FFF733280092FB5F5B62FB6E" box="[992,1059,1168,1193]" captionStart="FIGURE 5" captionStartId="6.[151,250,1892,1914]" captionTargetBox="[167,1425,1081,1859]" captionTargetId="figure@6.[158,1428,1078,1870]" captionTargetPageId="6" captionText="FIGURE 5. Depth of the mid-frontal impression of Hylurgops and Hylastes: a) deep and distinct (Hylurgops rugipennis), b) indistinct or absent (Hylastes mexicanus)." httpUri="https://zenodo.org/record/252619/files/figure.png" pageId="21" pageNumber="322">Fig. 5</figureCitation>
a); inter-puncture areas reticulate, rarely shiny; carina broadly elevated, surface shiny; vestiture hair-like, longer on lower half, 2–7× the length of a puncture’s diameter. 
<emphasis id="B937B732FFF7332802FFFB175D47FB36" bold="true" box="[397,518,1240,1265]" pageId="21" pageNumber="322">Pronotum</emphasis>
. Broad, 0.9–1.0 (0.93 ± 0.02), slightly narrower than elytral base, distinctly constricted on anterior third, widest anterior to middle; lateral margin narrowly rounded on basal fourth, becoming broadly rounded on central two fourths and broadly constricted on anterior fourth; middle line slightly raised, extends from base sometimes across entire pronotal length, discal surface reticulate (
<figureCitation id="137877A5FFF73328073BFA8B5BDBFA9A" box="[1097,1178,1348,1373]" captionStart="FIGURE 16" captionStartId="17.[151,250,1920,1942]" captionTargetBox="[151,1436,1224,1897]" captionTargetId="figure@17.[151,1436,1224,1898]" captionTargetPageId="17" captionText="FIGURE 16. Prothoracic discal surfaces of Hylurgops rugipennis and the eastern form of H. pinifex: a) reticulate and dull, sometimes shiny with similarly-sized punctures (H. rugipennis), b) shiny (eastern form), subgranulate (western form) with distinct, differently-sized punctures (H. pinifex)." httpUri="https://zenodo.org/record/252630/files/figure.png" pageId="21" pageNumber="322">Fig. 16</figureCitation>
a), rarely glossy; discal punctures of two equally abundant sizes, small two-thirds the size of large, inner surface shiny, inter puncture area reticulate; vestiture short, recumbent, hair-like setae indistinct to the length of a large discal punctures on disc, 2× the length near margins. 
<emphasis id="B937B732FFF7332802BBFA7F5D54FA0E" bold="true" box="[457,533,1456,1481]" pageId="21" pageNumber="322">Elytra</emphasis>
. Anterior margin moderately procurved, slightly raised dark margin; striae shallowly concave with round to keyhole-shaped, smooth, dark punctures, averaging half their diameter apart at disc; interstriae wide as first striae, wider than rest, smooth, shiny, minutely punctate (seen at&gt; 50× magnification), each with a short, recumbent hair-like setae becoming scale-like on posterior half to declivity and a central punctate granule with a long, sub-erect hair-like setae as long as a strial puncture appearing as rugosities on disc, separated by length of 1.5 strial puncture diameter. 
<emphasis id="B937B732FFF73328011FF9AB5D94F9BA" bold="true" box="[621,725,1636,1661]" pageId="21" pageNumber="322">Declivity</emphasis>
. Striae slightly impressed, less than half as wide as interstriae, punctures oval and deep, some keyhole-shaped, smaller than at disc, separated by distance equal to their diameter, 2nd interstriae slightly impressed, wider than 1st, 3rd widest to equal than 2nd, usually intersecting 6th (
<figureCitation id="137877A5FFF73328067FF9605A1EF90F" box="[1293,1375,1711,1736]" captionStart="FIGURE 13" captionStartId="15.[151,250,1472,1494]" captionTargetBox="[161,1428,990,1447]" captionTargetId="figure@15.[153,1433,988,1451]" captionTargetPageId="15" captionText="FIGURE 13. Declivity strial pattern of Hylurgops: a) third striae not touching the sixth (H. pinifex eastern form), b) third striae sometimes touching the sixth (H. rugipennis)." httpUri="https://zenodo.org/record/252627/files/figure.png" pageId="21" pageNumber="322">Fig. 13</figureCitation>
b) all with regularly spaced pointed granules, 2 punctures width apart; vestiture scale-like, short, recumbent, confused and a central reddish, erect, hair-like setae as long as strial width. 
<emphasis id="B937B732FFF733280005F9385B76F8D7" bold="true" box="[887,1079,1783,1808]" pageId="21" pageNumber="322">Ventral sclerites</emphasis>
. Surface punctured, interspaces reticulate; recumbent setae hair-like, as long as size of three punctures or more; precoxal ridge acutely elevated. 
<emphasis id="B937B732FFF7332803E5F8F05F8EF89F" bold="true" box="[151,207,1855,1880]" pageId="21" pageNumber="322">Legs</emphasis>
. Tibia reddish, narrower than in other species; protibiae with 1 or 2 mid-sized socketed teeth before apical angle, mesotibiae with 2 mid-sized socketed teeth before apical angle, and metatibiae broad with 3 medium-sized socketed teeth before apical angle; third tarsal segments bilobed, broader than second. 
<emphasis id="B937B732FFF7332807E8F8485A4CF867" bold="true" box="[1178,1293,1927,1952]" pageId="21" pageNumber="322">Aedeagus</emphasis>
. Showing a distinct ventral lobe (
<figureCitation id="137877A5FFF7332802F7F8645E89F803" box="[389,456,1963,1988]" captionStart="FIGURE 7" captionStartId="9.[151,250,1919,1941]" captionTargetBox="[199,1387,194,1893]" captionTargetId="figure@9.[190,1396,193,1897]" captionTargetPageId="9" captionText="FIGURE 7. Aedeagus of the New World Hylurgops and Pachysquamus: a) Pachysquamus subcostulatus, b) Hylurgops planirostris, c) H. rugipennis, d) H. pinifex, e) H. incomptus, f) H. longipennis, g) H. knausi, h) H. reticulatus, and i) H. porosus." httpUri="https://zenodo.org/record/252621/files/figure.png" pageId="21" pageNumber="322">Fig. 7</figureCitation>
d).
</paragraph>
<paragraph id="8BFC6B20FFF7332803B5F8005D38F82F" blockId="21.[151,1436,1096,2024]" box="[199,633,1999,2024]" pageId="21" pageNumber="322">
<emphasis id="B937B732FFF7332803B5F8005E45F82F" bold="true" box="[199,260,1999,2024]" pageId="21" pageNumber="322">Male</emphasis>
. Declivital hair-like setae longer.
</paragraph>
<paragraph id="8BFC6B20FFF4332B03B5FF585DD8FEDB" blockId="22.[151,1436,151,2016]" pageId="22" pageNumber="323">
<emphasis id="B937B732FFF4332B03B5FF585E60FF77" bold="true" box="[199,289,151,176]" pageId="22" pageNumber="323">Gallery</emphasis>
: Maternal gallery longitudinal, slightly sinuate, extending up and down from entrance hole (
<bibRefCitation id="EFD216D1FFF4332B0647FF585E52FF13" author="Bright" pageId="22" pageNumber="323" refString="Bright, D. E. &amp; Stark, R. W. (1973) The Bark and Ambrosia Beetles of California: Coleoptera: Scolytidae and Platypodidae. University of California Press, Berkeley, California, 169 pp." type="book" year="1973">Bright &amp; Stark 1973</bibRefCitation>
), usually just over the root collar of a dead tree or stump. Larval galleries extend perpendicular to brood gallery becoming confused at later stages. These galleries can extend into the roots, often at considerable distance from the brood gallery (
<bibRefCitation id="EFD216D1FFF4332B02D0FECB5DCCFEDB" author="Bright" box="[418,653,260,285]" pageId="22" pageNumber="323" refString="Bright, D. E. &amp; Stark, R. W. (1973) The Bark and Ambrosia Beetles of California: Coleoptera: Scolytidae and Platypodidae. University of California Press, Berkeley, California, 169 pp." type="book" year="1973">Bright &amp; Stark 1973</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35938ABFFF4332B03B5FEE85A21FBD5" pageId="22" pageNumber="323" type="materials_examined">
<paragraph id="8BFC6B20FFF4332B03B5FEE85DF6FC63" blockId="22.[151,1436,151,2016]" pageId="22" pageNumber="323">
<emphasis id="B937B732FFF4332B03B5FEE85EE9FE87" bold="true" box="[199,424,295,320]" pageId="22" pageNumber="323">Material examined</emphasis>
. 
<specimenCount id="9D45A0A9FFF4332B02C4FEE85D1FFE87" box="[438,606,295,320]" pageId="22" pageNumber="323" type="generic">250 specimens</specimenCount>
. 
<collectingCountry id="F3542BB0FFF4332B011AFEE75DA2FE87" box="[616,739,296,320]" name="Canada" pageId="22" pageNumber="323">CANADA</collectingCountry>
. 
<emphasis id="B937B732FFF4332B019FFEE85C08FE87" bold="true" box="[749,841,295,320]" pageId="22" pageNumber="323">Alberta</emphasis>
: Red Rock Cny. Waterton Lakes NP (
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). 
<emphasis id="B937B732FFF4332B063BFEE85E4CFEA2" bold="true" pageId="22" pageNumber="323">
British 
<collectingCountry id="F3542BB0FFF4332B03E5FE835E4CFEA2" box="[151,269,332,357]" name="United States of America" pageId="22" pageNumber="323">Columbia</collectingCountry>
</emphasis>
: Creston (
<collectionCode id="ED52F3E5FFF4332B02F4FE835E8FFEA3" box="[390,462,332,356]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
), 
<quantity id="4CBBC6C5FFF4332B0291FE835D5AFEA2" box="[483,539,332,357]" metricMagnitude="4" metricUnit="m" metricValue="1.2874752" pageId="22" pageNumber="323" unit="mi" value="8.0">8 mi</quantity>
. W Creston (
<collectionCode id="ED52F3E5FFF4332B01C1FE835DBAFEA3" box="[691,763,332,356]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
), 
<quantity id="4CBBC6C5FFF4332B0062FE835C17FEA2" box="[784,854,332,357]" metricMagnitude="4" metricUnit="m" metricValue="3.3796224" pageId="22" pageNumber="323" unit="mi" value="21.0">21 mi</quantity>
. W Creston (
<collectionCode id="ED52F3E5FFF4332B009FFE835B74FEA3" box="[1005,1077,332,356]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
), 
<quantity id="4CBBC6C5FFF4332B0739FE835BC2FEA2" box="[1099,1155,332,357]" metricMagnitude="3" metricUnit="m" metricValue="3.218688" pageId="22" pageNumber="323" unit="mi" value="2.0">2 mi</quantity>
. S Salmo River, Creston (
<collectionCode id="ED52F3E5FFF4332B03EDFEBF5FA6FE4F" box="[159,231,368,392]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
), Glacier (
<collectionCode id="ED52F3E5FFF4332B0214FEBF5EEFFE4F" box="[358,430,368,392]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
), Inverness (
<collectionCode id="ED52F3E5FFF4332B0137FEBF5DCFFE4F" box="[581,654,368,392]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
), 
<quantity id="4CBBC6C5FFF4332B01D4FEBF5DAEFE4F" box="[678,751,367,392]" metricMagnitude="4" metricUnit="m" metricValue="1.9312128" pageId="22" pageNumber="323" unit="mi" value="12.0">12 mi</quantity>
. S Long Beach, Tofino (
<collectionCode id="ED52F3E5FFF4332B076BFEBF5B20FE4F" box="[1049,1121,368,392]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
), Lorna (
<collectionCode id="ED52F3E5FFF4332B07A2FEBF5A59FE4F" box="[1232,1304,368,392]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
), Mainland (
<collectionCode id="ED52F3E5FFF4332B03EDFE5B5FA6FE6B" box="[159,231,404,428]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
), Massett, Graham Is. QCI (
<collectionCode id="ED52F3E5FFF4332B015AFE5B5D31FE6B" box="[552,624,404,428]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
), 
<quantity id="4CBBC6C5FFF4332B01F7FE5B5D95FE6A" box="[645,724,404,429]" metricMagnitude="3" metricUnit="m" metricValue="4.7" pageId="22" pageNumber="323" unit="km" value="4.7">4.7 km</quantity>
N Renell Sound Rd. Ghost Cr., Graham Is. (
<collectionCode id="ED52F3E5FFF4332B07A0FE5B5A5BFE6B" box="[1234,1306,404,428]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
), Sicamous (
<collectionCode id="ED52F3E5FFF4332B03EDFE775FA6FE17" box="[159,231,440,464]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
), Skidegate, Graham Is. (
<collectionCode id="ED52F3E5FFF4332B0164FE775D1EFE17" box="[534,607,440,464]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
), Stanley Park, Vancouver (
<collectionCode id="ED52F3E5FFF4332B00DBFE775CB0FE17" box="[937,1009,440,464]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
), 
<quantity id="4CBBC6C5FFF4332B0778FE775B04FE17" box="[1034,1093,439,464]" metricMagnitude="3" metricUnit="m" metricValue="1.609344" pageId="22" pageNumber="323" unit="mi" value="1.0">1 mi</quantity>
. NW Tlell, Graham Is., QCI (
<collectionCode id="ED52F3E5FFF4332B03EDFE135FA6FE33" box="[159,231,476,500]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
), Tow Hill, Graham Is. (
<collectionCode id="ED52F3E5FFF4332B0179FE135D12FE33" box="[523,595,476,500]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
), Trinity Valley (
<collectionCode id="ED52F3E5FFF4332B006EFE135C22FE33" box="[796,867,476,500]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
, 
<collectionCode id="ED52F3E5FFF4332B0000FE135C80FE33" box="[882,961,476,500]" pageId="22" pageNumber="323">DEBC</collectionCode>
), Vancouver (
<collectionCode id="ED52F3E5FFF4332B0711FE135BEAFE33" box="[1123,1195,476,500]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
), Laskeek Bay, Reef Island, QCI (
<collectionCode id="ED52F3E5FFF4332B0258FDCF5E32FDDF" box="[298,371,512,536]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
). 
<collectingCountry id="F3542BB0FFF4332B02F7FDCF5E80FDDF" box="[389,449,512,536]" name="United States of America" pageId="22" pageNumber="323">USA</collectingCountry>
. 
<emphasis id="B937B732FFF4332B02BEFE305D5FFDDF" bold="true" box="[460,542,511,536]" pageId="22" pageNumber="323">Alaska</emphasis>
: 
<quantity id="4CBBC6C5FFF4332B015CFE305D34FDDF" box="[558,629,511,536]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="22" pageNumber="323" unit="km" value="15.0">15 km</quantity>
N Juneau, Chichagof Is. (
<collectionCode id="ED52F3E5FFF4332B00EEFDCF5CADFDDF" box="[924,1004,512,536]" pageId="22" pageNumber="323">DEBC</collectionCode>
), Douglas Is. (
<collectionCode id="ED52F3E5FFF4332B07FDFDCF5B9EFDDF" box="[1167,1247,512,536]" pageId="22" pageNumber="323">DEBC</collectionCode>
), N end Douglas Is (
<collectionCode id="ED52F3E5FFF4332B03CFFDEB5E47FDFB" box="[189,262,548,572]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
); 
<quantity id="4CBBC6C5FFF4332B026FFDEB5E24FDFA" box="[285,357,548,573]" metricMagnitude="4" metricUnit="m" metricValue="1.9312128" pageId="22" pageNumber="323" unit="mi" value="12.0">12 mi</quantity>
. N Juneau (
<collectionCode id="ED52F3E5FFF4332B0281FDEB5D02FDFB" box="[499,579,548,572]" pageId="22" pageNumber="323">DEBC</collectionCode>
), 
<quantity id="4CBBC6C5FFF4332B012BFDEB5DF8FDFA" box="[601,697,548,573]" metricMagnitude="4" metricUnit="m" metricValue="4.15" pageId="22" pageNumber="323" unit="km" value="41.5">41.5 km</quantity>
N Juneau (
<collectionCode id="ED52F3E5FFF4332B0031FDEB5CCDFDFB" box="[835,908,548,572]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
); Juneau (
<collectionCode id="ED52F3E5FFF4332B0771FDEB5B15FDFB" box="[1027,1108,548,572]" pageId="22" pageNumber="323">DEBC</collectionCode>
). 
<emphasis id="B937B732FFF4332B0718FDEB5BA4FDFA" bold="true" box="[1130,1253,548,573]" pageId="22" pageNumber="323">California</emphasis>
: Del Norte Co. (
<collectionCode id="ED52F3E5FFF4332B03EDFD875FA9FDA7" box="[159,232,584,608]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
); El Dorado Co.: Blodgett Forest 
<collectingCountry id="F3542BB0FFF4332B011EFD875DD4FDA7" box="[620,661,584,608]" name="Australia" pageId="22" pageNumber="323">UC</collectingCountry>
, 
<quantity id="4CBBC6C5FFF4332B01D0FD875DA9FDA7" box="[674,744,583,608]" metricMagnitude="4" metricUnit="m" metricValue="1.609344" pageId="22" pageNumber="323" unit="mi" value="10.0">10 mi</quantity>
. E Georgetown (
<collectionCode id="ED52F3E5FFF4332B00DBFD875CB3FDA7" box="[937,1010,584,608]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
); Fresno Co. (
<collectionCode id="ED52F3E5FFF4332B07E6FD875B9CFDA7" box="[1172,1245,584,608]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
); Humboldt Co.: Eureka (
<collectionCode id="ED52F3E5FFF4332B038BFDA35E03FD43" box="[249,322,620,644]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
); Marin Co.: 
<quantity id="4CBBC6C5FFF4332B0291FDA35D5CFD42" box="[483,541,620,645]" metricMagnitude="3" metricUnit="m" metricValue="1.609344" pageId="22" pageNumber="323" unit="mi" value="1.0">1 mi</quantity>
. SE Inverness (
<collectionCode id="ED52F3E5FFF4332B01A7FDA35C5FFD43" box="[725,798,620,644]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
); Mendocino Co.: Mendocino (
<collectionCode id="ED52F3E5FFF4332B07FFFDA35B9FFD43" box="[1165,1246,620,644]" pageId="22" pageNumber="323">DEBC</collectionCode>
); Monterey Co.: Carmel (
<collectionCode id="ED52F3E5FFF4332B0388FD5F5E03FD6F" box="[250,322,656,680]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
), Monterey (
<collectionCode id="ED52F3E5FFF4332B02A6FD5F5D5CFD6F" box="[468,541,656,680]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
); San Mateo Co.: Año Nuevo, 
<quantity id="4CBBC6C5FFF4332B00F2FD5F5C88FD6F" box="[896,969,655,680]" metricMagnitude="4" metricUnit="m" metricValue="2.8" pageId="22" pageNumber="323" unit="km" value="28.0">28 km</quantity>
NNE of Santa Cruz (
<collectionCode id="ED52F3E5FFF4332B07B3FD5F5A4BFD6F" box="[1217,1290,656,680]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
); Santa Cruz Co.: Swanton (
<collectionCode id="ED52F3E5FFF4332B024FFD7B5EC7FD0B" box="[317,390,692,716]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
); Sonoma Co.: Salt Point St. Park (
<collectionCode id="ED52F3E5FFF4332B0078FD7B5C1AFD0B" box="[778,859,692,716]" pageId="22" pageNumber="323">DEBC</collectionCode>
); Tulare Co.: Kaweah R., Mid Fork (
<collectionCode id="ED52F3E5FFF4332B0781FD7B5A7DFD0B" box="[1267,1340,692,716]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
). 
<emphasis id="B937B732FFF4332B063CFD7B5AD5FD0A" bold="true" box="[1358,1428,692,717]" pageId="22" pageNumber="323">Idaho</emphasis>
: Kootenai Co.: Coeur d'Alene NF (
<collectionCode id="ED52F3E5FFF4332B0156FD175D32FD37" box="[548,627,728,752]" httpUri="http://grbio.org/cool/je3k-bvrg" name="C.P. Gillette Museum of Arthropod Diversity" pageId="22" pageNumber="323">CSUC</collectionCode>
). 
<emphasis id="B937B732FFF4332B01FAFD185DA3FD37" bold="true" box="[648,738,727,752]" pageId="22" pageNumber="323">Oregon</emphasis>
: Clatsop Co.: app. 
<quantity id="4CBBC6C5FFF4332B00B7FD175CBFFD37" box="[965,1022,727,752]" metricMagnitude="3" metricUnit="m" metricValue="4.828032" pageId="22" pageNumber="323" unit="mi" value="3.0">3 mi</quantity>
. SE of Olney (
<collectionCode id="ED52F3E5FFF4332B07DEFD175BB4FD37" box="[1196,1269,728,752]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
); Lincoln Co.: Otis, Cascade Head Exp. For. (
<collectionCode id="ED52F3E5FFF4332B029FFD335D77FCD3" box="[493,566,764,788]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
); Linn Co.: Santiam Pass (
<collectionCode id="ED52F3E5FFF4332B002EFD335CE2FCD3" box="[860,931,764,788]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
, 
<collectionCode id="ED52F3E5FFF4332B00DCFD335CBCFCD3" box="[942,1021,764,788]" pageId="22" pageNumber="323">DEBC</collectionCode>
); Tillamook Co.: Kiwanda viewpoint on Cape Lookout, 
<quantity id="4CBBC6C5FFF4332B0201FCD05E83FCFF" box="[371,450,799,824]" metricMagnitude="3" metricUnit="m" metricValue="4.02336" pageId="22" pageNumber="323" unit="mi" value="2.5">2.5 mi</quantity>
. 
<geoCoordinate id="EE770DE7FFF4332B02A2FCEF5D53FCFF" box="[464,530,799,824]" direction="north" orientation="latitude" pageId="22" pageNumber="323" precision="5555" value="1.5">
N 
<quantity id="4CBBC6C5FFF4332B029DFCD05D53FCFF" box="[495,530,799,824]" metricMagnitude="3" metricUnit="m" metricValue="2.414016" pageId="22" pageNumber="323" unit="mi" value="1.5">1.5</quantity>
</geoCoordinate>
mi. W Sand Lake (
<collectionCode id="ED52F3E5FFF4332B018DFCEF5C09FCFF" box="[767,840,800,824]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
); Wasco Co.: Wapinitia (
<collectionCode id="ED52F3E5FFF4332B071CFCEF5BFEFCFF" box="[1134,1215,800,824]" pageId="22" pageNumber="323">DEBC</collectionCode>
). 
<emphasis id="B937B732FFF4332B07A7FCD05A25FCFF" bold="true" box="[1237,1380,799,824]" pageId="22" pageNumber="323">Washington</emphasis>
: (no county) Mount Rainier NP (
<collectionCode id="ED52F3E5FFF4332B02A4FC8B5D5CFC9B" box="[470,541,836,860]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
, 
<collectionCode id="ED52F3E5FFF4332B0158FC8B5D36FC9B" box="[554,631,836,860]" httpUri="http://grbio.org/cool/je3k-bvrg" name="C.P. Gillette Museum of Arthropod Diversity" pageId="22" pageNumber="323">CSUC</collectionCode>
), (no county) Nisqually R. Mt. Rainier NP (
<collectionCode id="ED52F3E5FFF4332B071EFC8B5BF4FC9B" box="[1132,1205,836,860]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
); Grays Harbor Co.: Westport (
<collectionCode id="ED52F3E5FFF4332B027FFCA75E1DFC47" box="[269,348,872,896]" httpUri="http://grbio.org/cool/je3k-bvrg" name="C.P. Gillette Museum of Arthropod Diversity" pageId="22" pageNumber="323">CSUC</collectionCode>
); Jefferson Co.: Hoh Ranger Station, Olympic NP (
<collectionCode id="ED52F3E5FFF4332B00C5FCA75B41FC47" box="[951,1024,872,896]" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="22" pageNumber="323">CNCI</collectionCode>
); Pierce Co.: West Side Mt Rainier (
<collectionCode id="ED52F3E5FFF4332B03EDFC435FAFFC63" box="[159,238,908,932]" httpUri="http://grbio.org/cool/je3k-bvrg" name="C.P. Gillette Museum of Arthropod Diversity" pageId="22" pageNumber="323">CSUC</collectionCode>
); West Side Rd. 
<geoCoordinate id="EE770DE7FFF4332B02D5FC435E9FFC62" box="[423,478,908,933]" direction="north" orientation="latitude" pageId="22" pageNumber="323" precision="5555" value="1.6">1.6N</geoCoordinate>
Hwy 706 (
<collectionCode id="ED52F3E5FFF4332B012EFC435DEAFC63" box="[604,683,908,932]" httpUri="http://grbio.org/cool/je3k-bvrg" name="C.P. Gillette Museum of Arthropod Diversity" pageId="22" pageNumber="323">CSUC</collectionCode>
).
</paragraph>
<paragraph id="8BFC6B20FFF4332B03B5FC7D5A21FBD5" blockId="22.[151,1436,151,2016]" pageId="22" pageNumber="323">
<emphasis id="B937B732FFF4332B03B5FC7D5E4BFC0C" bold="true" box="[199,266,946,971]" pageId="22" pageNumber="323">Hosts</emphasis>
: 
<taxonomicName id="4C4310A3FFF4332B0250FC7D5E92FC0C" box="[290,467,946,971]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="22" pageNumber="323" phylum="Tracheophyta" rank="species" species="concolor">
<emphasis id="B937B732FFF4332B0250FC7D5E92FC0C" box="[290,467,946,971]" italics="true" pageId="22" pageNumber="323">Abies concolor</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4310A3FFF4332B0298FC7C5DD6FC0C" box="[490,663,946,971]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="22" pageNumber="323" phylum="Tracheophyta" rank="species" species="lasciocarpa">
<emphasis id="B937B732FFF4332B0298FC7C5DD6FC0C" box="[490,663,946,971]" italics="true" pageId="22" pageNumber="323">A. lasciocarpa</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4310A3FFF4332B01DCFC7D5C8FFC0C" box="[686,974,946,971]" class="Pinopsida" family="Cupressaceae" genus="Callitropsis" higherTaxonomySource="CoL" kingdom="Plantae" order="Pinales" pageId="22" pageNumber="323" phylum="Tracheophyta" rank="species" species="nootkatensis">
<emphasis id="B937B732FFF4332B01DCFC7D5C8FFC0C" box="[686,974,946,971]" italics="true" pageId="22" pageNumber="323">Callitropsis nootkatensis</emphasis>
</taxonomicName>
(see 
<bibRefCitation id="EFD216D1FFF4332B0766FC7C5A00FC0C" author="Patterson" box="[1044,1345,946,971]" pageId="22" pageNumber="323" refString="Patterson, G. K. &amp; Hatch, M. H. (1945) An annotated list of the Scolytoidea of Washington. University of Washington Publications in Biology, 10 (4), 145 - 154." type="journal article" year="1945">Patterson &amp; Hatch 1945</bibRefCitation>
), 
<taxonomicName id="4C4310A3FFF4332B062DFC7B5E5FFC28" class="Pinopsida" family="Pinaceae" genus="Larix" kingdom="Plantae" order="Pinales" pageId="22" pageNumber="323" phylum="Tracheophyta" rank="species" species="occidentalis">
<emphasis id="B937B732FFF4332B062DFC7B5E5FFC28" italics="true" pageId="22" pageNumber="323">Larix occidentalis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4310A3FFF4332B0241FC175D4BFC28" box="[307,522,982,1007]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="22" pageNumber="323" phylum="Tracheophyta" rank="species" species="engelmannii">
<emphasis id="B937B732FFF4332B0241FC175D4BFC28" box="[307,522,982,1007]" italics="true" pageId="22" pageNumber="323">Picea engelmannii</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4310A3FFF4332B016DFC175DEFFC28" box="[543,686,982,1007]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="22" pageNumber="323" phylum="Tracheophyta" rank="species" species="sitchensis">
<emphasis id="B937B732FFF4332B016DFC175DEFFC28" box="[543,686,982,1007]" italics="true" pageId="22" pageNumber="323">P. sitchensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4310A3FFF4332B01B0FC175C38FC28" box="[706,889,984,1007]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="22" pageNumber="323" phylum="Tracheophyta" rank="species" species="attenuata">
<emphasis id="B937B732FFF4332B01B0FC175C38FC28" box="[706,889,984,1007]" italics="true" pageId="22" pageNumber="323">Pinus attenuata</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4310A3FFF4332B00FFFC175B72FC28" box="[909,1075,982,1007]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="22" pageNumber="323" phylum="Tracheophyta" rank="species" species="balfouriana">
<emphasis id="B937B732FFF4332B00FFFC175B72FC28" box="[909,1075,982,1007]" italics="true" pageId="22" pageNumber="323">P. balfouriana</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4310A3FFF4332B073AFC175B9FFC28" box="[1096,1246,982,1007]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="22" pageNumber="323" phylum="Tracheophyta" rank="species" species="banksiana">
<emphasis id="B937B732FFF4332B073AFC175B9FFC28" box="[1096,1246,982,1007]" italics="true" pageId="22" pageNumber="323">P. banksiana</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4310A3FFF4332B0780FC175A32FC28" box="[1266,1395,984,1007]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="22" pageNumber="323" phylum="Tracheophyta" rank="species" species="contorta">
<emphasis id="B937B732FFF4332B0780FC175A32FC28" box="[1266,1395,984,1007]" italics="true" pageId="22" pageNumber="323">P. contorta</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4310A3FFF4332B06F5FC175E61FBD4" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="22" pageNumber="323" phylum="Tracheophyta" rank="species" species="lambertiana">
<emphasis id="B937B732FFF4332B06F5FC175E61FBD4" italics="true" pageId="22" pageNumber="323">P. lambertiana</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4310A3FFF4332B025CFC335EF8FBD4" box="[302,441,1018,1043]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="22" pageNumber="323" phylum="Tracheophyta" rank="species" species="monticola">
<emphasis id="B937B732FFF4332B025CFC335EF8FBD4" box="[302,441,1018,1043]" italics="true" pageId="22" pageNumber="323">P. monticola</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4310A3FFF4332B02BCFC335D0EFBD4" box="[462,591,1020,1043]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="22" pageNumber="323" phylum="Tracheophyta" rank="species" species="muricata">
<emphasis id="B937B732FFF4332B02BCFC335D0EFBD4" box="[462,591,1020,1043]" italics="true" pageId="22" pageNumber="323">P. muricata</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4310A3FFF4332B012FFC335DACFBD4" box="[605,749,1018,1043]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="22" pageNumber="323" phylum="Tracheophyta" rank="species" species="ponderosa">
<emphasis id="B937B732FFF4332B012FFC335DACFBD4" box="[605,749,1018,1043]" italics="true" pageId="22" pageNumber="323">P. ponderosa</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4310A3FFF4332B0189FC335C28FBD4" box="[763,873,1018,1043]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="22" pageNumber="323" phylum="Tracheophyta" rank="species" species="radiata">
<emphasis id="B937B732FFF4332B0189FC335C28FBD4" box="[763,873,1018,1043]" italics="true" pageId="22" pageNumber="323">P. radiata</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4310A3FFF4332B0005FC355B34FBD4" box="[887,1141,1018,1043]" class="Pinopsida" family="Pinaceae" genus="Pseudotsuga" kingdom="Plantae" order="Pinales" pageId="22" pageNumber="323" phylum="Tracheophyta" rank="species" species="menziesii">
<emphasis id="B937B732FFF4332B0005FC355B34FBD4" box="[887,1141,1018,1043]" italics="true" pageId="22" pageNumber="323">Pseudotsuga menziesii</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4310A3FFF4332B07F6FC335A18FBD4" box="[1156,1369,1018,1043]" class="Pinopsida" family="Pinaceae" genus="Tsuga" kingdom="Plantae" order="Pinales" pageId="22" pageNumber="323" phylum="Tracheophyta" rank="species" species="heterophylla">
<emphasis id="B937B732FFF4332B07F6FC335A18FBD4" box="[1156,1369,1018,1043]" italics="true" pageId="22" pageNumber="323">Tsuga heterophylla</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C35938ABFFF4332B03B5FBD15CC6FB9C" pageId="22" pageNumber="323" type="distribution">
<paragraph id="8BFC6B20FFF4332B03B5FBD15CC6FB9C" blockId="22.[151,1436,151,2016]" pageId="22" pageNumber="323">
<emphasis id="B937B732FFF4332B03B5FBD15E18FBF0" bold="true" box="[199,345,1054,1079]" pageId="22" pageNumber="323">Distribution</emphasis>
(
<figureCitation id="137877A5FFF4332B021EFBD15E83FBF0" box="[364,450,1054,1079]" captionStart="FIGURE 19" captionStartId="21.[151,250,999,1021]" captionTargetBox="[153,1436,194,977]" captionTargetId="figure@21.[151,1436,193,978]" captionTargetPageId="21" captionText="FIGURE 19. Species distribution with main hosts: Hylurgops rugipennis (orange) and H. pinifex (black-gray). Arrows indicate sympatric areas where the two species and their respective main hosts co-occur." httpUri="https://zenodo.org/record/252633/files/figure.png" pageId="22" pageNumber="323">Fig. 19</figureCitation>
). NORTH 
<collectingCountry id="F3542BB0FFF4332B0131FBD05D89FBF0" box="[579,712,1055,1079]" name="United States of America" pageId="22" pageNumber="323">AMERICA</collectingCountry>
. 
<collectingCountry id="F3542BB0FFF4332B01A7FBD05C0CFBF0" box="[725,845,1055,1079]" name="Canada" pageId="22" pageNumber="323">CANADA</collectingCountry>
and 
<collectingCountry id="F3542BB0FFF4332B00FBFBD05C84FBF0" box="[905,965,1055,1079]" name="United States of America" pageId="22" pageNumber="323">USA</collectingCountry>
. Kodiak Island, Alaska to Monterey and Tulare in southern California; mostly east of the Rocky Mountains.
</paragraph>
</subSubSection>
<subSubSection id="C35938ABFFF4332A03B5FBA95E8AFEA2" lastPageId="23" lastPageNumber="324" pageId="22" pageNumber="323" type="discussion">
<paragraph id="8BFC6B20FFF4332B03B5FBA95E52FB2D" blockId="22.[151,1436,151,2016]" pageId="22" pageNumber="323">
<emphasis id="B937B732FFF4332B03B5FBA95E0DFBB8" bold="true" box="[199,332,1126,1151]" pageId="22" pageNumber="323">Discussion.</emphasis>
<bibRefCitation id="EFD216D1FFF4332B022BFBA95D7EFBB8" author="Mannerheim" box="[345,575,1126,1151]" pageId="22" pageNumber="323" refString="Mannerheim, C. G. (1843) Beitrag zur kaefer-fauna der Aleutischen Inseln, der Insel Sitkha und Neu-Californiens. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 16, 3 - 142. http: // dx. doi. org / 10.5962 / bhl. title. 37833" type="journal article" year="1843">Mannerheim (1843)</bibRefCitation>
described 
<taxonomicName id="4C4310A3FFF4332B01B6FBA95CF1FBB8" box="[708,944,1126,1151]" class="Insecta" family="Curculionidae" genus="Hylurgus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="rugipennis">
<emphasis id="B937B732FFF4332B01B6FBA95CF1FBB8" box="[708,944,1126,1151]" italics="true" pageId="22" pageNumber="323">Hylurgus rugipennis</emphasis>
</taxonomicName>
from the Sitka Peninsula in Alaska, and placed it in the genus 
<taxonomicName id="4C4310A3FFF4332B02FDFB455EB1FB64" box="[399,496,1162,1187]" class="Insecta" family="Curculionidae" genus="Hylastes" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="genus">
<emphasis id="B937B732FFF4332B02FDFB455EB1FB64" box="[399,496,1162,1187]" italics="true" pageId="22" pageNumber="323">Hylastes</emphasis>
</taxonomicName>
in 1852. 
<bibRefCitation id="EFD216D1FFF4332B0129FB445C4CFB64" author="LeConte" box="[603,781,1162,1187]" pageId="22" pageNumber="323" refString="LeConte, J. L. (1868) Synopsis of the Scolytidae of America North of Mexico, Appendix. Transactions of the American Entomological Society, 2 (1), pp. 150 - 178." type="journal article" year="1868">LeConte (1868)</bibRefCitation>
and 
<bibRefCitation id="EFD216D1FFF4332B0037FB455CACFB64" author="Chapuis" box="[837,1005,1162,1187]" pageId="22" pageNumber="323" refString="Chapuis, F. (1869) Synopsis des Scolytides (Prodrome d'un travail monographique). Societe Royale des Sciences de Liege, Liege, Belgium, 61 pp." type="book" year="1869">Chapuis (1869</bibRefCitation>
, 
<bibRefCitation id="EFD216D1FFF4332B008AFB445B70FB64" author="Chapuis" box="[1016,1073,1163,1187]" pageId="22" pageNumber="323" refString="Chapuis, F. (1873) Synopsis des Scolytides. Memoires de la Societe Royale des Sciences de Liege, 2 (3), 213 - 269." type="journal article" year="1873">1873</bibRefCitation>
) supported the placement under Erichson’s (1836) second division of the genus 
<taxonomicName id="4C4310A3FFF4332B01A0FB615C73FB00" box="[722,818,1198,1223]" class="Insecta" family="Curculionidae" genus="Hylastes" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="genus">
<emphasis id="B937B732FFF4332B01A0FB615C73FB00" box="[722,818,1198,1223]" italics="true" pageId="22" pageNumber="323">Hylastes</emphasis>
</taxonomicName>
. 
<bibRefCitation id="EFD216D1FFF4332B0034FB605CBCFB00" author="LeConte" box="[838,1021,1198,1223]" pageId="22" pageNumber="323" refString="LeConte, J. L. (1876) Family IX, Scolytidae. In: LeConte, J. L. &amp; Horn, G. H. (Eds.), The Rhynchophora of America North of Mexico. Proceedings of the American Philosophical Society, 15 (96), pp. 341 - 391." type="journal article" year="1876">LeConte (1876)</bibRefCitation>
placed 
<taxonomicName id="4C4310A3FFF4332B072DFB7F5BBEFB00" box="[1119,1279,1200,1223]" class="Insecta" family="Curculionidae" genus="Hylurgus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="rugipennis">
<emphasis id="B937B732FFF4332B072DFB7F5BBEFB00" box="[1119,1279,1200,1223]" italics="true" pageId="22" pageNumber="323">H. rugipennis</emphasis>
</taxonomicName>
in his genus 
<taxonomicName id="4C4310A3FFF4332B03E5FB1D5E4DFB2C" box="[151,268,1234,1259]" class="Insecta" family="Curculionidae" genus="Hylurgops" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="genus">
<emphasis id="B937B732FFF4332B03E5FB1D5E4DFB2C" box="[151,268,1234,1259]" italics="true" pageId="22" pageNumber="323">Hylurgops</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BFC6B20FFF4332B03B5FB395DC2F9CC" blockId="22.[151,1436,151,2016]" pageId="22" pageNumber="323">
<bibRefCitation id="EFD216D1FFF4332B03B5FB395E2DFAC8" author="Wood" box="[199,364,1270,1295]" pageId="22" pageNumber="323" refString="Wood, S. L. (1971 a) Family Scolytidae (Ipidae). In: Hatch (Ed.), The Beetles of the Pacific Northwest: Part V: Rhipiceroidea, Sternoxi, Phytophaga, Rhynchophora, and Lamellicornia. University of Washington Press, Seattle, pp. 395 - 428." type="book chapter" year="1971" yearSuffix="a">Wood (1971a)</bibRefCitation>
considered that 
<taxonomicName id="4C4310A3FFF4332B0147FB375D95FAC8" box="[565,724,1272,1295]" class="Insecta" family="Curculionidae" genus="Hylurgus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="rugipennis">
<emphasis id="B937B732FFF4332B0147FB375D95FAC8" box="[565,724,1272,1295]" italics="true" pageId="22" pageNumber="323">H. rugipennis</emphasis>
</taxonomicName>
hybridized with 
<taxonomicName id="4C4310A3FFF4332B00D1FB375B56FAC8" box="[931,1047,1271,1295]" class="Insecta" family="Curculionidae" genus="Hylurgops" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="pinifex">
<emphasis id="B937B732FFF4332B00D1FB375B56FAC8" box="[931,1047,1271,1295]" italics="true" pageId="22" pageNumber="323">H. pinifex</emphasis>
</taxonomicName>
in an area east of the Canadian Cascades to Vernon in British 
<collectingCountry id="F3542BB0FFF4332B0172FAD55D30FAF4" box="[512,625,1306,1331]" name="United States of America" pageId="22" pageNumber="323">Columbia</collectingCountry>
, later 
<bibRefCitation id="EFD216D1FFF4332B01CCFAD55C1AFAF4" author="Bright" box="[702,859,1306,1331]" pageId="22" pageNumber="323" refString="Bright, D. E. (1976) The Insects and Arachnids of Canada, Part 2: The Bark Beetles of Canada and Alaska (Coleoptera: Scolytidae). Canada Department of Agriculture, Ottawa, Ontario, 241 pp." type="book" year="1976">Bright (1976)</bibRefCitation>
considered this hybridization area to extend into Alberta. 
<bibRefCitation id="EFD216D1FFF4332B038BFAF15ECCFA90" author="Wood" box="[249,397,1342,1367]" pageId="22" pageNumber="323" refString="Wood, S. L. (1982) The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Naturalist Memoirs, 6, 1 - 1359." type="journal article" year="1982">Wood (1982)</bibRefCitation>
reduced 
<taxonomicName id="4C4310A3FFF4332B0286FA8F5DCFFA90" box="[500,654,1344,1367]" class="Insecta" family="Curculionidae" genus="Hylurgus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="rugipennis">
<emphasis id="B937B732FFF4332B0286FA8F5DCFFA90" box="[500,654,1344,1367]" italics="true" pageId="22" pageNumber="323">H. rugipennis</emphasis>
</taxonomicName>
to a subspecies because of the apparent intergrades in the above area, but the characters used to define the intergrades were not detailed by 
<bibRefCitation id="EFD216D1FFF4332B00C6FAAD5B0EFABC" author="Wood" box="[948,1103,1378,1403]" pageId="22" pageNumber="323" refString="Wood, S. L. (1971 a) Family Scolytidae (Ipidae). In: Hatch (Ed.), The Beetles of the Pacific Northwest: Part V: Rhipiceroidea, Sternoxi, Phytophaga, Rhynchophora, and Lamellicornia. University of Washington Press, Seattle, pp. 395 - 428." type="book chapter" year="1971" yearSuffix="a">Wood (1971a</bibRefCitation>
, 
<bibRefCitation id="EFD216D1FFF4332B072EFAAC5BD9FABC" author="Wood" box="[1116,1176,1379,1403]" pageId="22" pageNumber="323" refString="Wood, S. L. (1982) The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Naturalist Memoirs, 6, 1 - 1359." type="journal article" year="1982">1982</bibRefCitation>
). Bright (pers. comm.) mentioned that the character used by Wood to determine the hybrids of the two species was the distribution of the scale-like setae cover in the elytra. This suggests that in the “intergradation zone,” 
<taxonomicName id="4C4310A3FFF4332B0729FA635BB9FA04" box="[1115,1272,1452,1475]" class="Insecta" family="Curculionidae" genus="Hylurgus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="rugipennis">
<emphasis id="B937B732FFF4332B0729FA635BB9FA04" box="[1115,1272,1452,1475]" italics="true" pageId="22" pageNumber="323">H. rugipennis</emphasis>
</taxonomicName>
has scale-like setae extending anteriorly into the elytral disc as in 
<taxonomicName id="4C4310A3FFF4332B019DFA1F5C21FA20" box="[751,864,1487,1511]" class="Insecta" family="Curculionidae" genus="Hylurgops" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="pinifex">
<emphasis id="B937B732FFF4332B019DFA1F5C21FA20" box="[751,864,1487,1511]" italics="true" pageId="22" pageNumber="323">H. pinifex</emphasis>
</taxonomicName>
, in contrast to the coastal forms of the species in which the scales are limited to the declivity.
</paragraph>
<paragraph id="8BFC6B20FFF4332B03B5F9D95B9AF8B4" blockId="22.[151,1436,151,2016]" pageId="22" pageNumber="323">
While considering it a subspecies, 
<bibRefCitation id="EFD216D1FFF4332B011BF9D95C42F9E8" author="Wood" box="[617,771,1558,1583]" pageId="22" pageNumber="323" refString="Wood, S. L. (1982) The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Naturalist Memoirs, 6, 1 - 1359." type="journal article" year="1982">Wood (1982)</bibRefCitation>
described the phenotypes of 
<taxonomicName id="4C4310A3FFF4332B071AF9D75A65F9E8" box="[1128,1316,1560,1583]" class="Insecta" family="Curculionidae" genus="Hylurgus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="subSpecies" species="rugipennis" subSpecies="rugipennis">
<emphasis id="B937B732FFF4332B071AF9D75A65F9E8" box="[1128,1316,1560,1583]" italics="true" pageId="22" pageNumber="323">H. r. rugipennis</emphasis>
</taxonomicName>
occurring throughout the Pacific Northwest as morphologically and biologically distinct from 
<taxonomicName id="4C4310A3FFF4332B07FBF9F35A5AF994" box="[1161,1307,1595,1619]" class="Insecta" family="Curculionidae" genus="Hylurgus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="subSpecies" species="rugipennis" subSpecies="pinifex">
<emphasis id="B937B732FFF4332B07FBF9F35A5AF994" box="[1161,1307,1595,1619]" italics="true" pageId="22" pageNumber="323">H. r. pinifex</emphasis>
</taxonomicName>
. Also, the distribution of the scale-like setae used in his subspecies determination represents a variation occurring across the two species’ distributions that was not observed by 
<bibRefCitation id="EFD216D1FFF4332B01A8F94D5C30F95C" author="Wood" box="[730,881,1666,1691]" pageId="22" pageNumber="323" refString="Wood, S. L. (1982) The bark and ambrosia beetles of North and Central America (Coleoptera: Scolytidae), a taxonomic monograph. Great Basin Naturalist Memoirs, 6, 1 - 1359." type="journal article" year="1982">Wood (1982)</bibRefCitation>
. Other characters such as the reticulate surface of the pronotal interspaces and the more regularly sized pronotal punctures of 
<taxonomicName id="4C4310A3FFF4332B009DF9675BCBF978" box="[1007,1162,1704,1727]" class="Insecta" family="Curculionidae" genus="Hylurgus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="rugipennis">
<emphasis id="B937B732FFF4332B009DF9675BCBF978" box="[1007,1162,1704,1727]" italics="true" pageId="22" pageNumber="323">H. rugipennis</emphasis>
</taxonomicName>
are constant throughout the two species’ distributions. The fact that 
<taxonomicName id="4C4310A3FFF4332B01F9F9035C67F924" box="[651,806,1740,1763]" class="Insecta" family="Curculionidae" genus="Hylurgus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="rugipennis">
<emphasis id="B937B732FFF4332B01F9F9035C67F924" box="[651,806,1740,1763]" italics="true" pageId="22" pageNumber="323">H. rugipennis</emphasis>
</taxonomicName>
was found in areas where its host 
<taxonomicName id="4C4310A3FFF4332B07C1F9035A2AF924" box="[1203,1387,1738,1763]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="22" pageNumber="323" phylum="Tracheophyta" rank="species" species="monticola">
<emphasis id="B937B732FFF4332B07C1F9035A2AF924" box="[1203,1387,1738,1763]" italics="true" pageId="22" pageNumber="323">Pinus monticola</emphasis>
</taxonomicName>
and some of the known hosts of 
<taxonomicName id="4C4310A3FFF4332B0290F93F5D13F8C0" box="[482,594,1775,1799]" class="Insecta" family="Curculionidae" genus="Hylurgops" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="pinifex">
<emphasis id="B937B732FFF4332B0290F93F5D13F8C0" box="[482,594,1775,1799]" italics="true" pageId="22" pageNumber="323">H. pinifex</emphasis>
</taxonomicName>
co-occur, but apparently not in areas where these conditions are not met explains the species’ sympatry in areas in Alberta, British 
<collectingCountry id="F3542BB0FFF4332B0050F8DD5CD1F8EC" box="[802,912,1810,1835]" name="United States of America" pageId="22" pageNumber="323">Columbia</collectingCountry>
and the Sierra Nevada of California (see black arrows, 
<figureCitation id="137877A5FFF4332B0384F8F95E0CF888" box="[246,333,1846,1871]" captionStart="FIGURE 19" captionStartId="21.[151,250,999,1021]" captionTargetBox="[153,1436,194,977]" captionTargetId="figure@21.[151,1436,193,978]" captionTargetPageId="21" captionText="FIGURE 19. Species distribution with main hosts: Hylurgops rugipennis (orange) and H. pinifex (black-gray). Arrows indicate sympatric areas where the two species and their respective main hosts co-occur." httpUri="https://zenodo.org/record/252633/files/figure.png" pageId="22" pageNumber="323">Fig. 19</figureCitation>
). In this revision, 
<taxonomicName id="4C4310A3FFF4332B015EF8F75D8AF888" box="[556,715,1848,1871]" class="Insecta" family="Curculionidae" genus="Hylurgus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="rugipennis">
<emphasis id="B937B732FFF4332B015EF8F75D8AF888" box="[556,715,1848,1871]" italics="true" pageId="22" pageNumber="323">H. rugipennis</emphasis>
</taxonomicName>
is treated as a distinct species from 
<taxonomicName id="4C4310A3FFF4332B07F4F8F75BB8F888" box="[1158,1273,1847,1871]" class="Insecta" family="Curculionidae" genus="Hylurgops" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="pinifex">
<emphasis id="B937B732FFF4332B07F4F8F75BB8F888" box="[1158,1273,1847,1871]" italics="true" pageId="22" pageNumber="323">H. pinifex</emphasis>
</taxonomicName>
based on the characters specified in the key and other characters included in the description of the two species.
</paragraph>
<paragraph id="8BFC6B20FFF4332A03B5F8B15E8AFEA2" blockId="22.[151,1436,151,2016]" lastBlockId="23.[151,1436,151,357]" lastPageId="23" lastPageNumber="324" pageId="22" pageNumber="323">
<taxonomicName id="4C4310A3FFF4332B03B5F8B15EFEF850" box="[199,447,1918,1943]" class="Insecta" family="Curculionidae" genus="Hylurgops" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="323" phylum="Arthropoda" rank="species" species="rugipennis">
<emphasis id="B937B732FFF4332B03B5F8B15EFEF850" box="[199,447,1918,1943]" italics="true" pageId="22" pageNumber="323">Hylurgops rugipennis</emphasis>
</taxonomicName>
, as defined herein, is restricted to the coastal forests of the Pacific Northwest from Kodiak Island in Alaska to Monterey in California and east to the Northern Rockies in Alberta, British 
<collectingCountry id="F3542BB0FFF4332B0654F86D5AD9F87C" box="[1318,1432,1954,1979]" name="United States of America" pageId="22" pageNumber="323">Columbia</collectingCountry>
; Idaho; Washington; and the Sierra Nevada of California (
<figureCitation id="137877A5FFF4332B0041F8095CCBF818" box="[819,906,1990,2015]" captionStart="FIGURE 19" captionStartId="21.[151,250,999,1021]" captionTargetBox="[153,1436,194,977]" captionTargetId="figure@21.[151,1436,193,978]" captionTargetPageId="21" captionText="FIGURE 19. Species distribution with main hosts: Hylurgops rugipennis (orange) and H. pinifex (black-gray). Arrows indicate sympatric areas where the two species and their respective main hosts co-occur." httpUri="https://zenodo.org/record/252633/files/figure.png" pageId="22" pageNumber="323">Fig. 19</figureCitation>
). In its coastal range from Alaska to extreme northern California, the species occurs primarily in 
<taxonomicName id="4C4310A3FFF5332A019BFF565CDEFF77" box="[745,927,151,176]" class="Pinopsida" family="Pinaceae" genus="Picea" kingdom="Plantae" order="Pinales" pageId="23" pageNumber="324" phylum="Tracheophyta" rank="species" species="sitchensis">
<emphasis id="B937B732FFF5332A019BFF565CDEFF77" box="[745,927,151,176]" italics="true" pageId="23" pageNumber="324">Picea sitchensis</emphasis>
</taxonomicName>
. South from there, the species’ range seems determined by the occurrence of the coastal “closed cone pine” species, 
<taxonomicName id="4C4310A3FFF5332A0081FF725BE8FF13" box="[1011,1193,189,212]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="23" pageNumber="324" phylum="Tracheophyta" rank="species" species="attenuata">
<emphasis id="B937B732FFF5332A0081FF725BE8FF13" box="[1011,1193,189,212]" italics="true" pageId="23" pageNumber="324">Pinus attenuata</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C4310A3FFF5332A07C9FF725A00FF13" box="[1211,1345,189,212]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="23" pageNumber="324" phylum="Tracheophyta" rank="species" species="muricata">
<emphasis id="B937B732FFF5332A07C9FF725A00FF13" box="[1211,1345,189,212]" italics="true" pageId="23" pageNumber="324">P. muricata</emphasis>
</taxonomicName>
, and 
<taxonomicName id="4C4310A3FFF5332A06F5FF725FABFF3F" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="23" pageNumber="324" phylum="Tracheophyta" rank="species" species="radiata">
<emphasis id="B937B732FFF5332A06F5FF725FABFF3F" italics="true" pageId="23" pageNumber="324">P. radiata</emphasis>
</taxonomicName>
. The distribution of 
<taxonomicName id="4C4310A3FFF5332A0292FF2E5D3CFF3F" box="[480,637,225,248]" class="Insecta" family="Curculionidae" genus="Hylurgops" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="324" phylum="Arthropoda" rank="species" species="rugipennis">
<emphasis id="B937B732FFF5332A0292FF2E5D3CFF3F" box="[480,637,225,248]" italics="true" pageId="23" pageNumber="324">H. rugipennis</emphasis>
</taxonomicName>
east of the Canadian Cascade Mountains into the Northern Rockies primarily follows the distribution of 
<taxonomicName id="4C4310A3FFF5332A0138FECA5D9AFEDA" box="[586,731,260,285]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="23" pageNumber="324" phylum="Tracheophyta" rank="species" species="monticola">
<emphasis id="B937B732FFF5332A0138FECA5D9AFEDA" box="[586,731,260,285]" italics="true" pageId="23" pageNumber="324">P. monticola</emphasis>
</taxonomicName>
, which extends up to the Waterton Lakes National Park in Alberta. In the Sierra Nevada, the southernmost distribution of the species into central California appears delimited by this conifer host as well.
</paragraph>
</subSubSection>
</treatment>
</document>