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<document id= "351FFC033CF1D9015519280262FDAC51" ID-DOI= "10.5281/zenodo.13133503" ID-ISSN= "0253-116X" ID-Zenodo-Dep= "13133503" IM.bibliography_approvedBy= "felipe" IM.illustrations_approvedBy= "felipe" IM.materialsCitations_approvedBy= "felipe" IM.metadata_approvedBy= "felipe" IM.tables_requiresApprovalFor= "GgImagineBatch" IM.taxonomicNames_approvedBy= "felipe" IM.treatmentCitations_approvedBy= "felipe" IM.treatments_approvedBy= "felipe" checkinTime= "1722338148560" checkinUser= "felipe" docAuthor= "Assing, V. & A, North" docDate= "2008" docId= "03F287B95E69FFB4FF6DDF93CC94FDDA" docLanguage= "en" docName= "LinzerbiolBeitr.2008.40.1.5-135.pdf" docOrigin= "Linzer biologische Beiträge 40 (1)" docStyle= "DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.10:LinzerbiolBeitr.2006-.journal_article" docStyleId= "24E5EB772D355DE2EE23B1E8D07853E0" docStyleName= "LinzerbiolBeitr.2006-.journal_article" docStyleVersion= "10" docTitle= "Sunius klapperichi" docType= "treatment" docVersion= "2" lastPageNumber= "73" masterDocId= "FFCBFFC15E2DFFF0FFDFDF4BCE4BFFEC" masterDocTitle= "A revision of the Sunius species of the Western Palaearctic region and Middle Asia (Coleoptera: Staphylinidae: Paederinae)" masterLastPageNumber= "135" masterPageNumber= "5" pageNumber= "73" updateTime= "1722339276815" updateUser= "ExternalLinkService" zenodo-license-document= "CC-BY-4.0" >
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<mods:title id= "D5DC84BA70C2DED3C6F1E9823716E183" > A revision of the Sunius species of the Western Palaearctic region and Middle Asia (Coleoptera: Staphylinidae: Paederinae)</mods:title>
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<mods:namePart id= "A62BFB82F04A9D1D289544276BA6FBF4" > Assing, V.</mods:namePart>
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<mods:namePart id= "BAA5949D7F90910FF161928A05A7B12D" > A, North</mods:namePart>
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<treatment id= "03F287B95E69FFB4FF6DDF93CC94FDDA" ID-DOI= "http://doi.org/10.5281/zenodo.13133657" ID-Zenodo-Dep= "13133657" LSID= "urn:lsid:plazi:treatment:03F287B95E69FFB4FF6DDF93CC94FDDA" httpUri= "http://treatment.plazi.org/id/03F287B95E69FFB4FF6DDF93CC94FDDA" lastPageNumber= "73" pageId= "68" pageNumber= "73" >
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<subSubSection id= "C34165245E69FFB4FF6DDF93CC81FF02" box= "[178,714,216,238]" pageId= "68" pageNumber= "73" type= "nomenclature" >
<paragraph id= "8BE436AF5E69FFB4FF6DDF93CC81FF02" blockId= "68.[178,714,216,238]" box= "[178,714,216,238]" pageId= "68" pageNumber= "73" >
<heading id= "D0AC81C35E69FFB4FF6DDF93CC81FF02" box= "[178,714,216,238]" fontSize= "8" level= "2" pageId= "68" pageNumber= "73" reason= "2" >
<emphasis id= "B92FEABD5E69FFB4FF6DDF93CC2EFF02" bold= "true" box= "[178,613,216,238]" pageId= "68" pageNumber= "73" >
4.36.
<taxonomicName id= "4C5B4D2C5E69FFB4FF34DF92CC2EFF02" ID-CoL= "53GP5" authority= "(COIFFAIT 1981)" baseAuthorityName= "COIFFAIT" baseAuthorityYear= "1981" box= "[235,613,216,238]" class= "Insecta" family= "Staphylinidae" genus= "Sunius" kingdom= "Animalia" order= "Coleoptera" pageId= "68" pageNumber= "73" phylum= "Arthropoda" rank= "species" species= "klapperichi" >
<emphasis id= "B92FEABD5E69FFB4FF34DF92CFFAFF02" bold= "true" box= "[235,433,216,238]" italics= "true" pageId= "68" pageNumber= "73" > Sunius klapperichi</emphasis>
(COIFFAIT 1981)
</taxonomicName>
</emphasis>
(
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<figureCitation id= "13602A2A5E69FFB4FDADDF93CC89FF02" box= "[626,706,216,238]" captionStart= "Map 14" captionStartId= "67.[178,223,1400,1420]" captionTargetBox= "[178,1093,836,1388]" captionTargetId= "figure-289@67.[178,1093,836,1388]" captionTargetPageId= "67" captionText= "Map 14: Distributions of species of the S. seminiger group in the Middle East: Sunius renouardi (COIFFAIT) (filled circles), S. hypogaeus (FAUVEL) (open circles), S. spinosissimus nov.sp. (filled square), and S. klapperichi (COIFFAIT) (open square)." figureDoi= "http://doi.org/10.5281/zenodo.13133563" httpUri= "https://zenodo.org/record/13133563/files/figure.png" pageId= "68" pageNumber= "73" > Map 14</figureCitation>
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)
</heading>
</paragraph>
</subSubSection>
<subSubSection id= "C34165245E69FFB4FF6DDE4BCCCAFE86" pageId= "68" pageNumber= "73" type= "reference_group" >
<paragraph id= "8BE436AF5E69FFB4FF6DDE4BCC2EFEF8" blockId= "68.[178,613,256,276]" box= "[178,613,256,276]" pageId= "68" pageNumber= "73" >
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<treatmentCitation id= "0AFA10BE5E69FFB4FF6DDE4BCC01FEF8" author= "COIFFAIT H" box= "[178,586,256,276]" page= "16" pageId= "68" pageNumber= "73" year= "1981" >
<taxonomicName id= "4C5B4D2C5E69FFB4FF6DDE4BCC01FEF8" ID-CoL= "3NXB6" authority= "COIFFAIT 1981 b: 16" authorityName= "COIFFAIT" authorityPageNumber= "16" authorityYear= "1981" box= "[178,586,256,276]" class= "Insecta" family= "Staphylinidae" genus= "Hypomedon" kingdom= "Animalia" order= "Coleoptera" pageId= "68" pageNumber= "73" phylum= "Arthropoda" rank= "species" species= "klapperichi" >
<emphasis id= "B92FEABD5E69FFB4FF6DDE4BCFC6FEF8" box= "[178,397,256,276]" italics= "true" pageId= "68" pageNumber= "73" > Hypomedon klapperichi</emphasis>
<bibRefCitation id= "EFCA4B5E5E69FFB4FE4CDE4BCC01FEF8" author= "COIFFAIT H" box= "[403,586,256,276]" pageId= "68" pageNumber= "73" pagination= "15 - 23" refId= "ref92463" refString= "COIFFAIT H. (1981 b): Staphylinidae nouveaux du Moyen Orient appartenant au Musee Hongrois d'Histoire Naturelle (Coleoptera). - Folia Entomologica Hungarica 34: 15 - 23." type= "journal article" year= "1981" >
<smallCapsWord id= "8D02A0735E69FFB4FE4CDE4BCFA1FEFE" baselines= "271,270" box= "[403,490,256,276]" lowerCaseFontSize= "7" mainFontSize= "9" normCase= "title" normString= "Coiffait" pageId= "68" pageNumber= "73" > COIFFAIT</smallCapsWord>
1981b: 16
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</taxonomicName>
</treatmentCitation>
ff.
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</paragraph>
<paragraph id= "8BE436AF5E69FFB4FF6DDE62CCCAFE86" blockId= "68.[178,1086,297,362]" pageId= "68" pageNumber= "73" >
<materialsCitation id= "3B333CF25E69FFB4FF6DDE62CCCAFE86" collectingDate= "1958-02-09" collectionCode= "MHNG" collectorName= "J. Klapperich" country= "Jordan" determinerName= "V. Assing" elevation= "500" location= "Wadi Wala" pageId= "68" pageNumber= "73" specimenCount= "1" specimenCount-female= "1" stateProvince= "Madaba" typeStatus= "holotype" >
T y p e m a t e r i a l e x a m i n e d:
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<specimenCount id= "9D5DFD265E69FFB4FD71DE66CC8AFEAE" box= "[686,705,301,322]" count= "1" pageId= "68" pageNumber= "73" type= "female" > ♀</specimenCount>
: " S.
<collectingCountry id= "F34C763F5E69FFB4FD2DDE62CD7AFED1" box= "[754,817,297,317]" name= "Jordan" pageId= "68" pageNumber= "73" > Jordan</collectingCountry>
,
<collectorName id= "26AE53795E69FFB4FCE2DE62CDF3FED1" box= "[829,952,297,317]" pageId= "68" pageNumber= "73" > J. Klapperich</collectorName>
/
<location id= "8E8460745E69FFB4FC11DE62CA71FED1" LSID= "urn:lsid:plazi:treatment:03F287B95E69FFB4FF6DDF93CC94FDDA:8E8460745E69FFB4FC11DE62CA71FED1" box= "[974,1082,297,317]" country= "Jordan" name= "Wadi Wala" pageId= "68" pageNumber= "73" stateProvince= "Madaba" > Wadi Wala</location>
, südl.
<collectingRegion id= "499FF84D5E69FFB4FF39DE0BCF79FEB8" box= "[230,306,320,340]" country= "Jordan" name= "Madaba" pageId= "68" pageNumber= "73" > Madaba</collectingRegion>
,
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<elevation id= "0076D19C5E69FFB4FEE1DE0BCF37FEB8" box= "[318,380,320,340]" metricMagnitude= "2" metricUnit= "m" metricValue= "5.0" pageId= "68" pageNumber= "73" unit= "m" value= "500.0" > 500 m</elevation>
</quantity>
,
<date id= "FFE5106F5E69FFB4FE57DE0BCF93FEB8" box= "[392,472,320,340]" pageId= "68" pageNumber= "73" value= "1958-02-09" >
<collectingDate id= "EFA1E9875E69FFB4FE57DE0BCF93FEB8" box= "[392,472,320,340]" pageId= "68" pageNumber= "73" value= "1958-02-09" > 9.2.1958</collectingDate>
</date>
/ u. Steinen / Type /
<taxonomicName id= "4C5B4D2C5E69FFB4FD74DE0BCA65FEB8" ID-CoL= "3NXB6" authority= "H. Coiffait 1980" authorityName= "H. Coiffait" authorityYear= "1980" box= "[683,1070,320,340]" class= "Insecta" family= "Staphylinidae" genus= "Hypomedon" kingdom= "Animalia" order= "Coleoptera" pageId= "68" pageNumber= "73" phylum= "Arthropoda" rank= "species" species= "klapperichi" >
Hypomedon klapperichi H.
<bibRefCitation id= "EFCA4B5E5E69FFB4FC6EDE0BCA65FEB8" author= "COIFFAIT H" box= "[945,1070,320,340]" pageId= "68" pageNumber= "73" pagination= "37 - 49" refId= "ref92417" refString= "COIFFAIT H. (1980): Staphylinides nouveaux. - Nouvelle Revue d'Entomologie 10: 37 - 49." type= "journal article" year= "1980" > Coiffait 1980</bibRefCitation>
</taxonomicName>
/
<taxonomicName id= "4C5B4D2C5E69FFB4FF6DDE1DCF17FE86" ID-CoL= "53GP5" baseAuthorityName= "COIFFAIT" baseAuthorityYear= "1981" box= "[178,348,342,362]" class= "Insecta" family= "Staphylinidae" genus= "Sunius" kingdom= "Animalia" order= "Coleoptera" pageId= "68" pageNumber= "73" phylum= "Arthropoda" rank= "species" species= "klapperichi" > Sunius klapperichi</taxonomicName>
det.
<determinerName id= "7E38DA6B5E69FFB4FE56DE1DCFAFFE86" box= "[393,484,342,362]" pageId= "68" pageNumber= "73" > V. Assing</determinerName>
2007" (
<collectionCode id= "ED4AAE6A5E69FFB4FDF3DE1DCC3CFE86" box= "[556,631,342,362]" country= "Switzerland" httpUri= "http://grbio.org/cool/2jyg-xmd2" name= "Museum d'Histoire Naturelle" pageId= "68" pageNumber= "73" type= "Museum" > MHNG</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id= "C34165245E69FFB4FF6DDE33CC94FDDA" pageId= "68" pageNumber= "73" type= "description" >
<paragraph id= "8BE436AF5E69FFB4FF6DDE33CC94FDDA" blockId= "68.[178,1086,376,566]" pageId= "68" pageNumber= "73" >
C o m m e n t s: The original description is based on a
<typeStatus id= "54E0880D5E69FFB4FCDADE33CD15FE62" box= "[773,862,376,398]" pageId= "68" pageNumber= "73" type= "holotype" > holotype</typeStatus>
and a
<typeStatus id= "54E0880D5E69FFB4FC76DE33CA48FE61" box= "[937,1027,376,397]" pageId= "68" pageNumber= "73" type= "paratype" > paratype</typeStatus>
, both females, from "
<collectingCountry id= "F34C763F5E69FFB4FE91DEDFCFECFE46" box= "[334,423,404,426]" name= "Jordan" pageId= "68" pageNumber= "73" > Jordanie</collectingCountry>
méridionale, Wadi Wala au sud de
<collectingRegion id= "499FF84D5E69FFB4FCC1DEDFCD38FE46" box= "[798,883,404,426]" country= "Jordan" name= "Madaba" pageId= "68" pageNumber= "73" > Madaba</collectingRegion>
" (
<bibRefCitation id= "EFCA4B5E5E69FFB4FC56DEDFCA79FE46" author= "COIFFAIT H" box= "[905,1074,404,426]" pageId= "68" pageNumber= "73" pagination= "15 - 23" refId= "ref92463" refString= "COIFFAIT H. (1981 b): Staphylinidae nouveaux du Moyen Orient appartenant au Musee Hongrois d'Histoire Naturelle (Coleoptera). - Folia Entomologica Hungarica 34: 15 - 23." type= "journal article" year= "1981" > COIFFAIT 1981b</bibRefCitation>
). The
<typeStatus id= "54E0880D5E69FFB4FF3EDEFBCF71FE2A" box= "[225,314,432,454]" pageId= "68" pageNumber= "73" type= "holotype" > holotype</typeStatus>
is externally indistinguishable from
<taxonomicName id= "4C5B4D2C5E69FFB4FD6ADEFBCD77FE2A" baseAuthorityName= "Fauvel" baseAuthorityYear= "1900" box= "[693,828,432,454]" class= "Insecta" family= "Staphylinidae" genus= "Sunius" kingdom= "Animalia" order= "Coleoptera" pageId= "68" pageNumber= "73" phylum= "Arthropoda" rank= "species" species= "hypogaeus" >
<emphasis id= "B92FEABD5E69FFB4FD6ADEFBCD77FE2A" box= "[693,828,432,454]" italics= "true" pageId= "68" pageNumber= "73" > S. hypogaeus</emphasis>
</taxonomicName>
and allied species. Since the type locality is some
<quantity id= "4CA39B4A5E69FFB4FE1BDE87CC5EFE0E" box= "[452,533,460,482]" metricMagnitude= "5" metricUnit= "m" metricValue= "2.0" pageId= "68" pageNumber= "73" unit= "km" value= "200.0" > 200 km</quantity>
away from the known distribution of
<taxonomicName id= "4C5B4D2C5E69FFB4FC6CDE87CA76FE0E" baseAuthorityName= "Fauvel" baseAuthorityYear= "1900" box= "[947,1085,460,482]" class= "Insecta" family= "Staphylinidae" genus= "Sunius" kingdom= "Animalia" order= "Coleoptera" pageId= "68" pageNumber= "73" phylum= "Arthropoda" rank= "species" species= "hypogaeus" >
<emphasis id= "B92FEABD5E69FFB4FC6CDE87CA76FE0E" box= "[947,1085,460,482]" italics= "true" pageId= "68" pageNumber= "73" > S. hypogaeus</emphasis>
</taxonomicName>
(
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<figureCitation id= "13602A2A5E69FFB4FF66DEA3CF41FE12" box= "[185,266,488,510]" captionStart= "Map 14" captionStartId= "67.[178,223,1400,1420]" captionTargetBox= "[178,1093,836,1388]" captionTargetId= "figure-289@67.[178,1093,836,1388]" captionTargetPageId= "67" captionText= "Map 14: Distributions of species of the S. seminiger group in the Middle East: Sunius renouardi (COIFFAIT) (filled circles), S. hypogaeus (FAUVEL) (open circles), S. spinosissimus nov.sp. (filled square), and S. klapperichi (COIFFAIT) (open square)." figureDoi= "http://doi.org/10.5281/zenodo.13133563" httpUri= "https://zenodo.org/record/13133563/files/figure.png" pageId= "68" pageNumber= "73" > Map 14</figureCitation>
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), it seems likely that
<taxonomicName id= "4C5B4D2C5E69FFB4FE3ADEA3CC39FE12" baseAuthorityName= "COIFFAIT" baseAuthorityYear= "1981" box= "[485,626,488,510]" class= "Insecta" family= "Staphylinidae" genus= "Sunius" kingdom= "Animalia" order= "Coleoptera" pageId= "68" pageNumber= "73" phylum= "Arthropoda" rank= "species" species= "klapperichi" >
<emphasis id= "B92FEABD5E69FFB4FE3ADEA3CC39FE12" box= "[485,626,488,510]" italics= "true" pageId= "68" pageNumber= "73" > S. klapperichi</emphasis>
</taxonomicName>
represents a distinct species. However, since the male sexual characters are unknown, its identity must remain doubtful until males from the vicinity of the type locality become available.
</paragraph>
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