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<document id="920C52788EC8175CAF13A4566C979970" ID-DOI="10.5281/zenodo.213946" ID-GBIF-Dataset="2a54e127-4d40-404e-ab7f-5b48f6e37e83" ID-ISSN="1175-5326" ID-Zenodo-Dep="213946" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460564534622" checkinUser="plazi" docAuthor="Grisales, Diana, Wolff, Marta &amp; De, Claudio J. B." docDate="2012" docId="03BE87F85234D42FC09425F6FCAFFD65" docLanguage="en" docName="zt03591p046.pdf" docOrigin="Zootaxa 3591" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Fannia canicularis Linnaeus" docType="treatment" docVersion="8" lastPageNumber="7" masterDocId="FF87FF805231D429C0032033FFBBFFA9" masterDocTitle="Neotropical Fanniidae (Insecta, Diptera): new species of Fannia from Colombia" masterLastPageNumber="46" masterPageNumber="1" pageNumber="6" updateTime="1698320797981" updateUser="plazi">
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<mods:title id="DE8048E6B79E1529547A6D07D89DA03F">Neotropical Fanniidae (Insecta, Diptera): new species of Fannia from Colombia</mods:title>
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<mods:namePart id="CE72F0052FA68A17AA1AB365F82EA4ED">Grisales, Diana</mods:namePart>
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<mods:namePart id="A49E58118FBBA51ED0DC501A80D4BFE8">Wolff, Marta</mods:namePart>
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<mods:namePart id="6A99B4635712E67B0860B8AD0A250046">De, Claudio J. B.</mods:namePart>
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<mods:title id="DD30191D1DA0509FC8C63969A262B642">Zootaxa</mods:title>
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<mods:part id="BD96683CED03A1F7ED8BEE48425B8139">
<mods:date id="A1B7A85673205549784246DB3E63F4CB">2012</mods:date>
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<mods:number id="3ABC337CBB0535D0DC9EA0CE3A003903">3591</mods:number>
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<mods:classification id="468293282A797EC7FFE98C069C2EB55A">journal article</mods:classification>
<mods:identifier id="9F3457DE7B5233215FC76827427CE9AF" type="DOI">10.5281/zenodo.213946</mods:identifier>
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<treatment id="03BE87F85234D42FC09425F6FCAFFD65" ID-DOI="http://doi.org/10.5281/zenodo.6175041" ID-GBIF-Taxon="119557549" ID-Zenodo-Dep="6175041" LSID="urn:lsid:plazi:treatment:03BE87F85234D42FC09425F6FCAFFD65" httpUri="http://treatment.plazi.org/id/03BE87F85234D42FC09425F6FCAFFD65" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<subSubSection id="C30D65655234D42CC09425F6FE44FA56" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="8BA836EE5234D42CC09425F6FDB5FA76" blockId="5.[151,526,1477,1535]" box="[151,526,1477,1503]" pageId="5" pageNumber="6">
<heading id="D0E081825234D42CC09425F6FDB5FA76" bold="true" box="[151,526,1477,1503]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
<taxonomicName id="4C174D6D5234D42CC09425F6FDB5FA76" ID-CoL="8894G" authority="Linnaeus" authorityName="Linnaeus" box="[151,526,1477,1503]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="canicularis">
<emphasis id="B963EAFC5234D42CC09425F6FDB5FA76" bold="true" box="[151,526,1477,1503]" pageId="5" pageNumber="6">
<emphasis id="B963EAFC5234D42CC09425F6FEC4FA76" bold="true" box="[151,383,1477,1503]" italics="true" pageId="5" pageNumber="6">Fannia canicularis</emphasis>
(Linnaeus)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BA836EE5234D42CC09425D4FE44FA56" blockId="5.[151,526,1477,1535]" box="[151,511,1511,1535]" pageId="5" pageNumber="6">
(
<figureCitation id="132C2A6B5234D42CC09C25D4FF57FA56" box="[159,236,1511,1535]" captionStart="FIGURES 1 – 12" captionStartId="8.[151,264,1845,1868]" captionTargetBox="[171,1399,339,1823]" captionTargetId="figure@8.[171,1417,339,1823]" captionTargetPageId="8" captionText="FIGURES 1 – 12. Fannia: (1) F. canicularis, male, head, anterior view; (2) F. chibcha sp. nov., male holotype, hind leg, anterior view; (3) F. dodgei, male, abdomen, posterior view; (4) F. dorsomaculata sp. nov., male holotype, lateral view; (5) F. dorsomaculata sp. nov., male holotype, dorsal view; (6) F. grandis sp. nov., male paratype, lateral view; (7) F. grandis, male, dorsal view; (8) F. iguaque sp. nov., male holotype, head, anterior view; (9) F. katios sp. nov., female paratype, head, anterior view; (10) F. katios sp. nov., male holotype, posterior view; (11) F. katios sp. nov., female paratype, posterior view; (12) F. lamosca sp. nov., male holotype, dorsal view. Scale: 1 mm." httpUri="https://zenodo.org/record/213947/files/figure.png" pageId="5" pageNumber="6">Figs. 1</figureCitation>
,
<figureCitation id="132C2A6B5234D42CC0F925D4FEA2FA56" box="[250,281,1511,1535]" captionStart="FIGURES 20 – 37" captionStartId="13.[151,264,1716,1739]" captionTargetBox="[151,1435,193,1694]" captionTargetId="figure@13.[151,1436,193,1695]" captionTargetPageId="13" captionText="FIGURES 20 – 37. Fannia: male, hind leg, anterior view: (20) F. canicularis; (21) F. chibcha sp. nov., holotype; (22) F. chingaza sp. nov., holotype; (23) F. dodgei; (24) F. dorsomaculata sp. nov., holotype; (25) F. e m b e r a sp. nov., paratype; (26) F. grandis; (27) F. iguaque sp. nov., holotype; (28) F. katios sp. nov., holotype; (29) F. l a m o s c a sp. nov., holotype; (30) F. magdalena sp. nov., holotype; (31) F. pijao sp. nov., holotype; (32) F. p o rc e sp. nov., holotype; (33) F. pu s i o; (34) F. quimbaya sp. nov., holotype; (35) F. sumapaz sp. nov., holotype; (36) F. sutagao sp. nov., holotype; (37) F. trimaculata. Scale: 1 mm." httpUri="https://zenodo.org/record/213949/files/figure.png" pageId="5" pageNumber="6">20</figureCitation>
,
<figureCitation id="132C2A6B5234D42CC12725D4FEF9FA56" box="[292,322,1511,1535]" captionStart="FIGURES 38 – 55" captionStartId="14.[151,264,1744,1767]" captionTargetBox="[151,1435,193,1723]" captionTargetId="figure@14.[151,1436,193,1723]" captionTargetPageId="14" captionText="FIGURES 38 – 55. Fannia: male, hind leg, posterior view: (38) F. canicularis; (39) F. chibcha sp. nov., holotype; (40) F. chingaza sp. nov., holotype; (41) F. dodgei; (42) F. dorsomaculata sp. nov., holotype; (43) F. e m b e r a sp. nov., holotype; (44) F. grandis; (45) F. iguaque sp. nov., holotype; (46) F. katios sp. nov., holotype; (47) F. l a m o s c a sp. nov., holotype; (48) F. magdalena sp. nov., holotype; (49) F. pijao sp. nov., holotype; (50) F. p o rc e sp. nov., holotype; (51) F. pu s i o; (52) F. quimbaya sp. nov., holotype; (53) F. sumapaz sp. nov., holotype; (54) F. sutagao sp. nov., holotype; (55) F. trimaculata. Scale: 1 mm." httpUri="https://zenodo.org/record/213950/files/figure.png" pageId="5" pageNumber="6">38</figureCitation>
,
<figureCitation id="132C2A6B5234D42CC14D25D4FED7FA56" box="[334,364,1511,1535]" captionStart="FIGURES 56 – 73" captionStartId="15.[151,264,1875,1898]" captionTargetBox="[200,1393,193,1844]" captionTargetId="figure@15.[200,1394,193,1845]" captionTargetPageId="15" captionText="FIGURES 56 – 73. Fannia: male, sternite 5, ventral view: (56) F. c a ni c ul a ri s; (57) F. c h i b c h a sp. nov., paratype; (58) F. chingaza sp. nov., holotype; (59) F. dodgei; (60) F. dorsomaculata sp. nov., paratype; (61) F. e m b e r a sp. nov., paratype; (62) F. grandis; (63) F. iguaque sp. nov., paratype; (64) F. katios sp. nov., paratype; (65) F. l a m o s c a sp. nov., paratype; (66) F. magdalena sp. nov., paratype; (67) F. pijao sp. nov., paratype; (68) F. porce sp. nov., holotype; (69) F. p u s i o; (70) F. quimbaya sp. nov., paratype; (71) F. sumapaz sp. nov., paratype; (72) F. sutagao sp. nov., paratype; (73) F. trimaculata. Scale: 0.1 mm." httpUri="https://zenodo.org/record/213951/files/figure.png" pageId="5" pageNumber="6">56</figureCitation>
,
<figureCitation id="132C2A6B5234D42CC17B25D4FE2DFA56" box="[376,406,1511,1535]" captionStart="FIGURES 74 – 82" captionStartId="16.[151,264,1809,1832]" captionTargetBox="[152,1435,193,1787]" captionTargetId="figure@16.[151,1436,193,1789]" captionTargetPageId="16" captionText="FIGURES 74 – 82. Fannia: male, epandrium, cercal plate and surstyli, dorsal view: (74) F. c an i c ul a r i s; (75) F. c hi b c ha sp. nov., paratype; (76) F. c h i n g a z a sp. nov., holotype; (77) F. dodgei; (78) F. dorsomaculata sp. nov., paratype; (79) F. e m b e r a sp. nov., paratype; (80) F. grandis; (81) F. iguaque sp. nov., paratype; (82) F. katios sp. nov., paratype. Scale: 0.1 mm." httpUri="https://zenodo.org/record/213952/files/figure.png" pageId="5" pageNumber="6">74</figureCitation>
,
<figureCitation id="132C2A6B5234D42CC1A125D4FE7AFA56" box="[418,449,1511,1535]" captionStart="FIGURES 92 – 100" captionStartId="18.[151,264,1950,1973]" captionTargetBox="[246,1354,193,1930]" captionTargetId="figure@18.[237,1355,193,1931]" captionTargetPageId="18" captionText="FIGURES 92 – 100. Fannia: male, epandrium, cercal plate and surstyli, lateral view: (92) F. canicularis; (93) F. c h i b c h a sp. nov., paratype; (94) F. c h i n g a z a sp. nov., holotype; (95) F. dodgei; (96) F. dorsomaculata sp. nov., paratype; (97) F. e m b e r a sp. nov., paratype; (98) F. grandis; (99) F. iguaque sp. nov., paratype; (100) F. katios sp. nov., paratype. Scale: 0.1 mm." httpUri="https://zenodo.org/record/213954/files/figure.png" pageId="5" pageNumber="6">92</figureCitation>
,
<figureCitation id="132C2A6B5234D42CC1CF25D4FE4DFA56" box="[460,502,1511,1535]" captionStart="FIGURES 110 – 118" captionStartId="20.[151,264,1635,1658]" captionTargetBox="[165,1427,193,1608]" captionTargetId="figure@20.[163,1428,193,1609]" captionTargetPageId="20" captionText="FIGURES 110 – 118. Fannia: male, hypandrium and associated structures, dorsal view: (110) F. canicularis; (111) F. chibcha sp. nov., paratype; (112) F. c h i n g a z a sp. nov., holotype; (113) F. dodgei; (114) F. dorsomaculata sp. nov., paratype; (115) F. embera sp. nov., paratype; (116) F. grandis; (117) F. iguaque sp. nov., paratype. Scale: 0.1 mm." httpUri="https://zenodo.org/record/213956/files/figure.png" pageId="5" pageNumber="6">110</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30D65655234D42CC0942602FBF9F9CD" pageId="5" pageNumber="6" type="reference_group">
<paragraph id="8BA836EE5234D42CC0942602FBF9F9CD" blockId="5.[151,1436,1584,1637]" pageId="5" pageNumber="6">
<treatmentCitationGroup id="AB0711C05234D42CC0942602FBF9F9CD" pageId="5" pageNumber="6">
<taxonomicName id="4C174D6D5234D42CC0942602FD34F9EF" ID-CoL="8894G" authority="Linnaeus, 1761: 454" authorityName="Linnaeus" authorityPageNumber="454" authorityYear="1761" box="[151,655,1584,1606]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="canicularis">
<emphasis id="B963EAFC5234D42CC0942602FEE2F9EF" box="[151,345,1584,1606]" italics="true" pageId="5" pageNumber="6">Fannia canicularis</emphasis>
<treatmentCitation id="0AB610FF5234D42CC1632603FD89F9EF" author="Linnaeus" box="[352,562,1584,1606]" page="454" pageId="5" pageNumber="6" year="1761">
<bibRefCitation id="EF864B1F5234D42CC1632603FDBAF9EF" author="Linnaeus" box="[352,513,1584,1606]" pageId="5" pageNumber="6" refString="Linnaeus, C. (1761) Fauna Svecica Sistens Animalia Sveciae Regni: Mammalia, aves, amphibia, pisces, insecta, vermes. Distributa per classes &amp; ordines, genera &amp; species, cum differentiis specierum, synonymis auctorum, nominibus incalorum, locis natalium, descriptionibus insec-torum, Editio altera, auctior. 2 nd Ed. Laurenti Salvii, Stockholmiae, 578 pp." type="book" year="1761">Linnaeus, 1761</bibRefCitation>
: 454
</treatmentCitation>
(
<emphasis id="B963EAFC5234D42CC2422602FD3FF9EF" box="[577,644,1585,1606]" italics="true" pageId="5" pageNumber="6">Musca</emphasis>
)
</taxonomicName>
. New name for
<emphasis id="B963EAFC5234D42CC3362603FC32F9EF" box="[821,905,1584,1606]" italics="true" pageId="5" pageNumber="6">lateralis</emphasis>
<bibRefCitation id="EF864B1F5234D42CC3922603FB8AF9EF" author="Linnaeus" box="[913,1073,1584,1606]" pageId="5" pageNumber="6" refString="Linnaeus, C. (1758) Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum caracteribus, differentiis, synonymis, locis. 10 th Ed. Vol. 1. Salvius, Holmiae [= Stockholm], 824 pp." type="book" year="1758">Linnaeus, 1758</bibRefCitation>
.
<typeStatus id="54AC884C5234D42CC4382602FB1BF9EF" box="[1083,1184,1585,1606]" pageId="5" pageNumber="6" type="lectotype">Lectotype</typeStatus>
male of
<taxonomicName id="4C174D6D5234D42CC4FE2602FA29F9EF" box="[1277,1426,1584,1606]" class="Insecta" family="Anthomyiidae" genus="Musca" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="laterali">
<emphasis id="B963EAFC5234D42CC4FE2602FA29F9EF" box="[1277,1426,1584,1606]" italics="true" pageId="5" pageNumber="6">Musca laterali</emphasis>
</taxonomicName>
s (des.
<treatmentCitation id="0AB610FF5234D42CC0FD267CFE1BF9CC" author="Pont" box="[254,416,1615,1637]" page="170" pageId="5" pageNumber="6" year="1981">
<bibRefCitation id="EF864B1F5234D42CC0FD267CFED4F9CD" author="Pont" box="[254,367,1615,1636]" pageId="5" pageNumber="6" refString="Pont, A. C. (1981) The Linnaean species of the families Fanniidae, Anthomyiidae and Muscidae (Insecta: Diptera). Biological Journal of the Linnean Society, 15, 165 - 175." type="journal article" year="1981">Pont, 1981</bibRefCitation>
: 170
</treatmentCitation>
) in the Linnean Society, London (LSUK). Type-locality: “Europe.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30D65655234D42FC09426A2FDE8FF5E" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="8BA836EE5234D42FC09426A2FDE8FF5E" blockId="5.[151,1436,1681,2029]" lastBlockId="6.[151,1437,151,716]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<emphasis id="B963EAFC5234D42CC09426A2FEAAF903" bold="true" box="[151,273,1681,1706]" pageId="5" pageNumber="6">Diagnosis.</emphasis>
These characters apply only to the male sex. General coloration yellowish; eye bare; frontal vitta greyish and narrow; 8
<emphasis id="B963EAFC5234D42CC1A32686FE09F964" box="[416,434,1717,1741]" italics="true" pageId="5" pageNumber="6">fr</emphasis>
; upper
<emphasis id="B963EAFC5234D42CC20D2686FD8EF964" box="[526,565,1717,1741]" italics="true" pageId="5" pageNumber="6">orb</emphasis>
present (
<figureCitation id="132C2A6B5234D42CC2A02686FD53F964" box="[675,744,1717,1741]" captionStart="FIGURES 1 – 12" captionStartId="8.[151,264,1845,1868]" captionTargetBox="[171,1399,339,1823]" captionTargetId="figure@8.[171,1417,339,1823]" captionTargetPageId="8" captionText="FIGURES 1 – 12. Fannia: (1) F. canicularis, male, head, anterior view; (2) F. chibcha sp. nov., male holotype, hind leg, anterior view; (3) F. dodgei, male, abdomen, posterior view; (4) F. dorsomaculata sp. nov., male holotype, lateral view; (5) F. dorsomaculata sp. nov., male holotype, dorsal view; (6) F. grandis sp. nov., male paratype, lateral view; (7) F. grandis, male, dorsal view; (8) F. iguaque sp. nov., male holotype, head, anterior view; (9) F. katios sp. nov., female paratype, head, anterior view; (10) F. katios sp. nov., male holotype, posterior view; (11) F. katios sp. nov., female paratype, posterior view; (12) F. lamosca sp. nov., male holotype, dorsal view. Scale: 1 mm." httpUri="https://zenodo.org/record/213947/files/figure.png" pageId="5" pageNumber="6">Fig. 1</figureCitation>
); fronto-orbital plate and parafacialia greyish pollinose and without short setae; scape and pedicel black; scape with 1 seta reaching the pedicel; arista dark brown with base black; palpus black and filiform; scutum brown with 3 brown vittae along
<emphasis id="B963EAFC5234D42CC41526CDFB80F8BC" box="[1046,1083,1790,1813]" italics="true" pageId="5" pageNumber="6">acr</emphasis>
and
<emphasis id="B963EAFC5234D42CC47F26CEFB2CF8BC" box="[1148,1175,1789,1813]" italics="true" pageId="5" pageNumber="6">dc</emphasis>
, not reaching base of scutellum; 2
<emphasis id="B963EAFC5234D42CC1282711FEE9F890" box="[299,338,1826,1849]" italics="true" pageId="5" pageNumber="6">pra</emphasis>
, short and near the
<emphasis id="B963EAFC5234D42CC23A2712FDD3F890" box="[569,616,1825,1849]" italics="true" pageId="5" pageNumber="6">spal</emphasis>
; wing and calypters yellowish; haltere yellowish with base brown; legs dark brown with yellow joints; mid tibia constricted on ventral surface of basal half; hind coxa setulose on posterior margin; hind femur without pre-apical protuberance on ventral surface, 1 developed
<emphasis id="B963EAFC5234D42CC4C12759FB67F828" box="[1218,1244,1898,1921]" italics="true" pageId="5" pageNumber="6">av</emphasis>
on apical half, 1 row of
<emphasis id="B963EAFC5234D42CC0EE27BEFEB2F80C" box="[237,265,1933,1957]" italics="true" pageId="5" pageNumber="6">ad</emphasis>
ending in 5 developed
<emphasis id="B963EAFC5234D42CC21F27BEFD91F80C" box="[540,554,1933,1957]" italics="true" pageId="5" pageNumber="6">d</emphasis>
(
<figureCitation id="132C2A6B5234D42CC23F27BEFD28F80C" box="[572,659,1933,1957]" captionStart="FIGURES 20 – 37" captionStartId="13.[151,264,1716,1739]" captionTargetBox="[151,1435,193,1694]" captionTargetId="figure@13.[151,1436,193,1695]" captionTargetPageId="13" captionText="FIGURES 20 – 37. Fannia: male, hind leg, anterior view: (20) F. canicularis; (21) F. chibcha sp. nov., holotype; (22) F. chingaza sp. nov., holotype; (23) F. dodgei; (24) F. dorsomaculata sp. nov., holotype; (25) F. e m b e r a sp. nov., paratype; (26) F. grandis; (27) F. iguaque sp. nov., holotype; (28) F. katios sp. nov., holotype; (29) F. l a m o s c a sp. nov., holotype; (30) F. magdalena sp. nov., holotype; (31) F. pijao sp. nov., holotype; (32) F. p o rc e sp. nov., holotype; (33) F. pu s i o; (34) F. quimbaya sp. nov., holotype; (35) F. sumapaz sp. nov., holotype; (36) F. sutagao sp. nov., holotype; (37) F. trimaculata. Scale: 1 mm." httpUri="https://zenodo.org/record/213949/files/figure.png" pageId="5" pageNumber="6">Fig. 20</figureCitation>
); hind tibia with 1 row of short
<emphasis id="B963EAFC5234D42CC40F27BEFB93F80C" box="[1036,1064,1933,1957]" italics="true" pageId="5" pageNumber="6">ad</emphasis>
on basal half, ending in 1 short median seta, 2 median
<emphasis id="B963EAFC5234D42CC1952781FE0AF860" box="[406,433,1970,1993]" italics="true" pageId="5" pageNumber="6">av</emphasis>
(
<figureCitation id="132C2A6B5234D42CC1BC2782FDA8F860" box="[447,531,1969,1993]" captionStart="FIGURES 20 – 37" captionStartId="13.[151,264,1716,1739]" captionTargetBox="[151,1435,193,1694]" captionTargetId="figure@13.[151,1436,193,1695]" captionTargetPageId="13" captionText="FIGURES 20 – 37. Fannia: male, hind leg, anterior view: (20) F. canicularis; (21) F. chibcha sp. nov., holotype; (22) F. chingaza sp. nov., holotype; (23) F. dodgei; (24) F. dorsomaculata sp. nov., holotype; (25) F. e m b e r a sp. nov., paratype; (26) F. grandis; (27) F. iguaque sp. nov., holotype; (28) F. katios sp. nov., holotype; (29) F. l a m o s c a sp. nov., holotype; (30) F. magdalena sp. nov., holotype; (31) F. pijao sp. nov., holotype; (32) F. p o rc e sp. nov., holotype; (33) F. pu s i o; (34) F. quimbaya sp. nov., holotype; (35) F. sumapaz sp. nov., holotype; (36) F. sutagao sp. nov., holotype; (37) F. trimaculata. Scale: 1 mm." httpUri="https://zenodo.org/record/213949/files/figure.png" pageId="5" pageNumber="6">Fig. 20</figureCitation>
); abdomen elongate, brown-greyish with syntergite 1+2, tergites 2–3 translucentyellow with median vitta brown; sternite 1 setulose; sternite 5 as in
<figureCitation id="132C2A6B5234D42CC39C27E6FC48F844" box="[927,1011,2005,2029]" captionStart="FIGURES 56 – 73" captionStartId="15.[151,264,1875,1898]" captionTargetBox="[200,1393,193,1844]" captionTargetId="figure@15.[200,1394,193,1845]" captionTargetPageId="15" captionText="FIGURES 56 – 73. Fannia: male, sternite 5, ventral view: (56) F. c a ni c ul a ri s; (57) F. c h i b c h a sp. nov., paratype; (58) F. chingaza sp. nov., holotype; (59) F. dodgei; (60) F. dorsomaculata sp. nov., paratype; (61) F. e m b e r a sp. nov., paratype; (62) F. grandis; (63) F. iguaque sp. nov., paratype; (64) F. katios sp. nov., paratype; (65) F. l a m o s c a sp. nov., paratype; (66) F. magdalena sp. nov., paratype; (67) F. pijao sp. nov., paratype; (68) F. porce sp. nov., holotype; (69) F. p u s i o; (70) F. quimbaya sp. nov., paratype; (71) F. sumapaz sp. nov., paratype; (72) F. sutagao sp. nov., paratype; (73) F. trimaculata. Scale: 0.1 mm." httpUri="https://zenodo.org/record/213951/files/figure.png" pageId="5" pageNumber="6">Fig. 56</figureCitation>
; terminalia (
<figureCitation id="132C2A6B5234D42CC48127E6FB59F844" box="[1154,1250,2005,2029]" captionStart="FIGURES 74 – 82" captionStartId="16.[151,264,1809,1832]" captionTargetBox="[152,1435,193,1787]" captionTargetId="figure@16.[151,1436,193,1789]" captionTargetPageId="16" captionText="FIGURES 74 – 82. Fannia: male, epandrium, cercal plate and surstyli, dorsal view: (74) F. c an i c ul a r i s; (75) F. c hi b c ha sp. nov., paratype; (76) F. c h i n g a z a sp. nov., holotype; (77) F. dodgei; (78) F. dorsomaculata sp. nov., paratype; (79) F. e m b e r a sp. nov., paratype; (80) F. grandis; (81) F. iguaque sp. nov., paratype; (82) F. katios sp. nov., paratype. Scale: 0.1 mm." httpUri="https://zenodo.org/record/213952/files/figure.png" pageId="5" pageNumber="6">Figs. 74</figureCitation>
,
<figureCitation id="132C2A6B5234D42CC4ED27E6FAB4F844" box="[1262,1295,2005,2029]" captionStart="FIGURES 92 – 100" captionStartId="18.[151,264,1950,1973]" captionTargetBox="[246,1354,193,1930]" captionTargetId="figure@18.[237,1355,193,1931]" captionTargetPageId="18" captionText="FIGURES 92 – 100. Fannia: male, epandrium, cercal plate and surstyli, lateral view: (92) F. canicularis; (93) F. c h i b c h a sp. nov., paratype; (94) F. c h i n g a z a sp. nov., holotype; (95) F. dodgei; (96) F. dorsomaculata sp. nov., paratype; (97) F. e m b e r a sp. nov., paratype; (98) F. grandis; (99) F. iguaque sp. nov., paratype; (100) F. katios sp. nov., paratype. Scale: 0.1 mm." httpUri="https://zenodo.org/record/213954/files/figure.png" pageId="5" pageNumber="6">92</figureCitation>
): epandrium slightly wider than long, with setae predominantly on basal half, 2 setae developed basally; surstylus widened basally with a set of setae on the outer edges, apex slender with short setae; bacilliform process absent; hypandrium and associated structures as in
<figureCitation id="132C2A6B5237D42FC1EC20ECFDF4FF5E" box="[495,591,223,247]" captionStart="FIGURES 110 – 118" captionStartId="20.[151,264,1635,1658]" captionTargetBox="[165,1427,193,1608]" captionTargetId="figure@20.[163,1428,193,1609]" captionTargetPageId="20" captionText="FIGURES 110 – 118. Fannia: male, hypandrium and associated structures, dorsal view: (110) F. canicularis; (111) F. chibcha sp. nov., paratype; (112) F. c h i n g a z a sp. nov., holotype; (113) F. dodgei; (114) F. dorsomaculata sp. nov., paratype; (115) F. embera sp. nov., paratype; (116) F. grandis; (117) F. iguaque sp. nov., paratype. Scale: 0.1 mm." httpUri="https://zenodo.org/record/213956/files/figure.png" pageId="6" pageNumber="7">Fig. 110</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30D65655237D42FC0C42130FB54FE66" pageId="6" pageNumber="7" type="biology_ecology">
<paragraph id="8BA836EE5237D42FC0C42130FB54FE66" blockId="6.[151,1437,151,716]" pageId="6" pageNumber="7">
<emphasis id="B963EAFC5237D42FC0C42130FE9CFEB5" bold="true" box="[199,295,259,284]" pageId="6" pageNumber="7">Biology.</emphasis>
<taxonomicName id="4C174D6D5237D42FC12E2136FDB9FEB5" box="[301,514,260,284]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="canicularis">
<emphasis id="B963EAFC5237D42FC12E2136FDB9FEB5" box="[301,514,260,284]" italics="true" pageId="6" pageNumber="7">Fannia canicularis</emphasis>
</taxonomicName>
is a cosmopolitan and synanthropic species that has been widely studied (
<bibRefCitation id="EF864B1F5237D42FC53B2137FF69FE96" author="Chillcott" pageId="6" pageNumber="7" refString="Chillcott, J. (1961) A Revision of the Neartic Species of Fanniinae (Diptera: Muscidae). Canadian Entomologist, 92, 1 - 295." type="journal article" year="1961">Chillcott 1961</bibRefCitation>
;
<bibRefCitation id="EF864B1F5237D42FC0DC2114FE7EFE96" author="Almeida" box="[223,453,295,320]" pageId="6" pageNumber="7" refString="Almeida, J. R., de Carvalho, C. J. B. &amp; Malkowski, S. R. (1985) Dipteros sinantropicos de Curitiba e arredores (Paran, Brazil). II. Fanniidae e Anthomyiidae. Anais da Sociedade Entomologica Brasileira, 14, 277 - 288." type="journal article" year="1985">
Almeida
<emphasis id="B963EAFC5237D42FC14B211AFE39FEE9" box="[328,386,296,320]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
1985
</bibRefCitation>
; Queiroz &amp; de
<bibRefCitation id="EF864B1F5237D42FC2752114FC9BFE96" author="Carvalho" box="[630,800,295,319]" pageId="6" pageNumber="7" refString="Queiroz, S. M. &amp; Carvalho, C. C. (1987) Chave pictorica e descricoes de larvas de 3 instar de Diptera (Calliphoridae, Muscidae and Fanniidae) em vazadouros de residuos solidos domesticos em Curitiba, Paran. Anais da Sociedade Entomologica Brasileira, 16, 265 - 288." type="journal article" year="1987">Carvalho 1987</bibRefCitation>
;
<bibRefCitation id="EF864B1F5237D42FC32D211BFB86FE96" author="Meyer" box="[814,1085,295,320]" pageId="6" pageNumber="7" refString="Meyer, J. A. &amp; Mullens, B. A. (1988) Development of inmature Fannia spp. (Diptera: Muscidae) at constant laboratory temperatures. Journal Medical of Entomology, 25, 165 - 171." type="journal article" year="1988">Meyer &amp; Mullens 1988</bibRefCitation>
; de
<bibRefCitation id="EF864B1F5237D42FC46E2114FAE7FE96" author="Carvalho" box="[1133,1372,295,320]" pageId="6" pageNumber="7" refString="de Carvalho, C. J. B., Pont A. C., Couri, M. S. &amp; Pamplona, D. (2003) A catalogue of the Fanniidae (Diptera) of the Neotropical Region. Zootaxa, 219, 1 - 32." type="journal article" year="2003">
Carvalho
<emphasis id="B963EAFC5237D42FC4DF211AFAADFEE9" box="[1244,1302,296,320]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
2003
</bibRefCitation>
). It is capable of transporting bacteria such as
<taxonomicName id="4C174D6D5237D42FC264217FFC9BFECD" box="[615,800,332,356]" class="Gammaproteobacteria" family="Enterobacteriaceae" genus="Escherichia" kingdom="Bacteria" order="Enterobacteriales" pageId="6" pageNumber="7" phylum="Proteobacteria" rank="species" species="coli">
<emphasis id="B963EAFC5237D42FC264217FFC9BFECD" box="[615,800,332,356]" italics="true" pageId="6" pageNumber="7">Escherichia coli</emphasis>
</taxonomicName>
and
<taxonomicName id="4C174D6D5237D42FC35F217FFBE6FECD" box="[860,1117,332,356]" class="Bacilli" family="Staphylococcaceae" genus="Staphylococcus" kingdom="Bacteria" order="Bacillales" pageId="6" pageNumber="7" phylum="Firmicutes" rank="species" species="aureus">
<emphasis id="B963EAFC5237D42FC35F217FFBE6FECD" box="[860,1117,332,356]" italics="true" pageId="6" pageNumber="7">Staphylococcus aureus</emphasis>
</taxonomicName>
, protozoans and nematodes (
<bibRefCitation id="EF864B1F5237D42FC09D215CFEC7FE2E" author="Rozkosny" box="[158,380,367,392]" pageId="6" pageNumber="7" refString="Rozkosny, R., Gregor, F. &amp; Pont, A. C. (1997) The European Fanniidae (Diptera). Acta Scientiarum Bohemicae Brno, 31, 1 - 80." type="journal article" year="1997">
Rozkošný
<emphasis id="B963EAFC5237D42FC1192143FE8CFE21" box="[282,311,368,392]" italics="true" pageId="6" pageNumber="7">al.</emphasis>
1997
</bibRefCitation>
). Recently the role of
<taxonomicName id="4C174D6D5237D42FC2832142FCA7FE21" box="[640,796,368,392]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="canicularis">
<emphasis id="B963EAFC5237D42FC2832142FCA7FE21" box="[640,796,368,392]" italics="true" pageId="6" pageNumber="7">F. canicularis</emphasis>
</taxonomicName>
in vectoring Newcastle disease has been demonstrated. The virus, which lives in the median and distal portions of the intestine of
<taxonomicName id="4C174D6D5237D42FC3E721A6FBC5FE05" box="[996,1150,404,428]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="canicularis">
<emphasis id="B963EAFC5237D42FC3E721A6FBC5FE05" box="[996,1150,404,428]" italics="true" pageId="6" pageNumber="7">F. canicularis</emphasis>
</taxonomicName>
, is transmitted to poultry and other birds, affecting their nervous, respiratory and digestive systems (
<bibRefCitation id="EF864B1F5237D42FC3D82184FB5FFE66" author="Chakrabarti" box="[987,1252,439,464]" pageId="6" pageNumber="7" refString="Chakrabarti, S., King, D. J., Cardona, C. J. &amp; Gerry, A. C. (2008) Persistence of Exotic Newcastle DiseaseVirus (ENDV) in laboratory infected Musca domestica and Fannia canicularis. Avian Diseases, 52, 375 - 379." type="journal article" year="2008">
Chakrabarti
<emphasis id="B963EAFC5237D42FC46B218AFB22FE79" box="[1128,1177,440,464]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
. 2008
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30D65655237D42FC0C421E8FB68FDBE" pageId="6" pageNumber="7" type="discussion">
<paragraph id="8BA836EE5237D42FC0C421E8FB68FDBE" blockId="6.[151,1437,151,716]" pageId="6" pageNumber="7">
<emphasis id="B963EAFC5237D42FC0C421E8FEF4FE5D" bold="true" box="[199,335,475,500]" pageId="6" pageNumber="7">Comments.</emphasis>
<taxonomicName id="4C174D6D5237D42FC15521EEFE4BFE5D" box="[342,496,476,500]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="canicularis">
<emphasis id="B963EAFC5237D42FC15521EEFE4BFE5D" box="[342,496,476,500]" italics="true" pageId="6" pageNumber="7">F. canicularis</emphasis>
</taxonomicName>
belongs to the sub-group
<taxonomicName id="4C174D6D5237D42FC31721EFFC29FE5D" box="[788,914,476,500]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="canicularis">
<emphasis id="B963EAFC5237D42FC31721EFFC29FE5D" box="[788,914,476,500]" italics="true" pageId="6" pageNumber="7">canicularis</emphasis>
</taxonomicName>
of the
<taxonomicName id="4C174D6D5237D42FC3E221EFFBE4FE5D" box="[993,1119,476,500]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="canicularis">
<emphasis id="B963EAFC5237D42FC3E221EFFBE4FE5D" box="[993,1119,476,500]" italics="true" pageId="6" pageNumber="7">canicularis</emphasis>
</taxonomicName>
group (
<bibRefCitation id="EF864B1F5237D42FC4B521EFFAE6FE5D" author="Chillcott" box="[1206,1373,476,500]" pageId="6" pageNumber="7" refString="Chillcott, J. (1961) A Revision of the Neartic Species of Fanniinae (Diptera: Muscidae). Canadian Entomologist, 92, 1 - 295." type="journal article" year="1961">Chillcott 1961</bibRefCitation>
). The monophyly of the group, however, has not been corroborated (
<bibRefCitation id="EF864B1F5237D42FC35121CCFB7CFDBE" author="Dominguez" box="[850,1223,511,535]" pageId="6" pageNumber="7" refString="Dominguez, M. C. &amp; Roig-Junent, S. (2008) A phylogeny of the family Fanniidae Schnabl (Insecta: Diptera: Calyptratae) based on adult morphological characters, with special reference to the Austral species of the genus Fannia. Invertebrate Systematics, 22, 563 - 587." type="journal article" year="2008">Domínguez &amp; Roig-Juñent 2008</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30D65655237D42FC0C42217FAE7FD2D" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="8BA836EE5237D42FC0C42217FD9AFD95" blockId="6.[151,1437,151,716]" box="[199,545,547,572]" pageId="6" pageNumber="7">
<emphasis id="B963EAFC5237D42FC0C42217FECEFD95" bold="true" box="[199,373,547,572]" pageId="6" pageNumber="7">
<typeStatus id="54AC884C5237D42FC0C42217FEB9FD95" box="[199,258,548,572]" pageId="6" pageNumber="7">Type</typeStatus>
material.
</emphasis>
Not examined.
</paragraph>
<paragraph id="8BA836EE5237D42FC0C42274FAE7FD2D" blockId="6.[151,1437,151,716]" pageId="6" pageNumber="7">
<emphasis id="B963EAFC5237D42FC0C42274FE02FDC9" bold="true" box="[199,441,583,608]" pageId="6" pageNumber="7">Material examined.</emphasis>
<collectingCountry id="F300767E5237D42FC1CA2274FDD9FDF6" box="[457,610,583,607]" name="Colombia" pageId="6" pageNumber="7">COLOMBIA</collectingCountry>
. Boyacá:
<specimenCount id="9D11FD675237D42FC2D62274FC9FFDF6" box="[725,804,583,607]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
Pauna, Manote bajo, La Rochela, Alt. 1.280 mts.,
<date id="FFA9102E5237D42FC094225FFEB0FD2D" box="[151,267,620,644]" pageId="6" pageNumber="7" value="1979-04-28">28–IV–79</date>
, Col. I. de Arévalo (
<collectionCode id="ED06AE2B5237D42FC1EC225FFDEEFD2D" box="[495,597,620,644]" httpUri="http://grbio.org/cool/ct7t-ipa9" name="Museo Entomologico Francisco Luis Gallego" pageId="6" pageNumber="7">MEFLG</collectionCode>
);
<taxonomicName id="4C174D6D5237D42FC264225FFD53FD2D" box="[615,744,620,644]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="33" phylum="Arthropoda" rank="species" species="magdalena" status="sp. nov.">Magdalena</taxonomicName>
:
<specimenCount id="9D11FD675237D42FC2F1225FFC86FD2D" box="[754,829,620,644]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
Santa
<collectingCountry id="F300767E5237D42FC389225EFC76FD2D" box="[906,973,621,644]" name="Malta" pageId="6" pageNumber="7">Marta</collectingCountry>
<date id="FFA9102E5237D42FC3D7225FFB91FD2D" box="[980,1066,620,644]" pageId="6" pageNumber="7" value="1973-03-01">1–3–73</date>
, Silvio Zuluaga (
<collectionCode id="ED06AE2B5237D42FC4E9225FFAEBFD2D" box="[1258,1360,620,644]" httpUri="http://grbio.org/cool/ct7t-ipa9" name="Museo Entomologico Francisco Luis Gallego" pageId="6" pageNumber="7">MEFLG</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C30D65655237D42FC0C422BCFCAFFD65" pageId="6" pageNumber="7" type="distribution">
<paragraph id="8BA836EE5237D42FC0C422BCFCAFFD65" blockId="6.[151,1437,151,716]" pageId="6" pageNumber="7">
<emphasis id="B963EAFC5237D42FC0C422BCFEDBFD01" bold="true" box="[199,352,655,680]" pageId="6" pageNumber="7">Distribution.</emphasis>
Cosmopolitan (de
<bibRefCitation id="EF864B1F5237D42FC24422BCFCF9FD0E" author="Carvalho" box="[583,834,655,680]" pageId="6" pageNumber="7" refString="de Carvalho, C. J. B., Pont A. C., Couri, M. S. &amp; Pamplona, D. (2003) A catalogue of the Fanniidae (Diptera) of the Neotropical Region. Zootaxa, 219, 1 - 32." type="journal article" year="2003">
Carvalho
<emphasis id="B963EAFC5237D42FC2B922A2FD4AFD01" box="[698,753,656,680]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
. 2003
</bibRefCitation>
).
<collectingCountry id="F300767E5237D42FC35922BCFC76FD0E" box="[858,973,655,679]" name="Colombia" pageId="6" pageNumber="7">Colombia</collectingCountry>
: departments of
<taxonomicName id="4C174D6D5237D42FC49522BCFAA9FD0E" box="[1174,1298,655,679]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="33" phylum="Arthropoda" rank="species" species="magdalena" status="sp. nov.">Magdalena</taxonomicName>
(Caribbean region) and Boyacá (Andean region, Eastern Cordillera).
</paragraph>
</subSubSection>
</treatment>
</document>
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