treatments-xml/data/03/BE/87/03BE87F85210D40AC094241DFBCFFE5D.xml

<document id="5A8C6B3AEF0E2E3BC85970B917FE2814" ID-DOI="10.5281/zenodo.213946" ID-GBIF-Dataset="2a54e127-4d40-404e-ab7f-5b48f6e37e83" ID-ISSN="1175-5326" ID-Zenodo-Dep="213946" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460564534622" checkinUser="plazi" docAuthor="Grisales, Diana, Wolff, Marta &amp; De, Claudio J. B." docDate="2012" docId="03BE87F85210D40AC094241DFBCFFE5D" docLanguage="en" docName="zt03591p046.pdf" docOrigin="Zootaxa 3591" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Fannia pijao Grisales, Wolff &amp; De, 2012, sp. nov." docType="treatment" docVersion="8" lastPageNumber="36" masterDocId="FF87FF805231D429C0032033FFBBFFA9" masterDocTitle="Neotropical Fanniidae (Insecta, Diptera): new species of Fannia from Colombia" masterLastPageNumber="46" masterPageNumber="1" pageNumber="34" updateTime="1698320797981" updateUser="plazi">
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<mods:title id="9436A550556DF9FAF0D0A6205B1F6FC3">Neotropical Fanniidae (Insecta, Diptera): new species of Fannia from Colombia</mods:title>
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<mods:namePart id="61B7512615CB7D9942706F7ABF5130C9">Grisales, Diana</mods:namePart>
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<mods:namePart id="3761F82F69BAB36DC1CB1CB535CE8362">Wolff, Marta</mods:namePart>
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<mods:namePart id="1DAF4F0B823A1EA5F010E621E8647F69">De, Claudio J. B.</mods:namePart>
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<mods:title id="2CF3CE8A05EC0FDF2AB92DABE712DE0D">Zootaxa</mods:title>
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<mods:date id="80C497CF85DBEBAE992C57A1AB963C50">2012</mods:date>
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<treatment id="03BE87F85210D40AC094241DFBCFFE5D" ID-DOI="http://doi.org/10.5281/zenodo.6175071" ID-GBIF-Taxon="119557568" ID-Zenodo-Dep="6175071" LSID="urn:lsid:plazi:treatment:03BE87F85210D40AC094241DFBCFFE5D" httpUri="http://treatment.plazi.org/id/03BE87F85210D40AC094241DFBCFFE5D" lastPageId="35" lastPageNumber="36" pageId="33" pageNumber="34">
<subSubSection id="C30D65655210D408C094241DFDC6FBC0" pageId="33" pageNumber="34" type="nomenclature">
<paragraph id="8BA836EE5210D408C094241DFE22FBE1" blockId="33.[151,637,1070,1129]" box="[151,409,1070,1096]" pageId="33" pageNumber="34">
<heading id="D0E081825210D408C094241DFE22FBE1" bold="true" box="[151,409,1070,1096]" fontSize="11" level="1" pageId="33" pageNumber="34" reason="1">
<emphasis id="B963EAFC5210D408C094241DFE22FBE1" bold="true" box="[151,409,1070,1096]" pageId="33" pageNumber="34">
<taxonomicName id="4C174D6D5210D408C094241DFE8FFBE1" ID-CoL="9J2K9" box="[151,308,1070,1096]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="pijao" status="sp. nov.">
<emphasis id="B963EAFC5210D408C094241DFE8FFBE1" bold="true" box="[151,308,1070,1096]" italics="true" pageId="33" pageNumber="34">Fannia pijao</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25057875210D408C138241DFE22FBE1" box="[315,409,1070,1096]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA836EE5210D408C0942462FDC6FBC0" blockId="33.[151,637,1070,1129]" box="[151,637,1105,1129]" pageId="33" pageNumber="34">
(
<figureCitation id="132C2A6B5210D408C09C2462FE9DFBC0" box="[159,294,1105,1129]" captionStart="FIGURES 13 – 19" captionStartId="9.[151,264,858,881]" captionTargetBox="[153,1434,193,836]" captionTargetId="figure@9.[153,1435,193,837]" captionTargetPageId="9" captionText="FIGURES 13 – 19. Fannia: (13) F. magdalena sp. nov., male holotype, lateral view; (14) F. pijao sp. nov., male holotype, head, anterior view; (15) F. pijao sp. nov., male holotype, thorax, lateral view; (16) F. p o rc e sp. nov., male holotype, head, lateral view; (17) F. q u i m b a y a sp. nov., male holotype, head, lateral view; (18) F. q u i m b a y a sp. nov., male holotype, thorax, lateral view; (19) F. sutagao sp. nov., male holotype, head, anterior view. Scale: 1 mm." httpUri="https://zenodo.org/record/213948/files/figure.png" pageId="33" pageNumber="34">Figs. 14, 15</figureCitation>
, 
<figureCitation id="132C2A6B5210D408C1312462FEEBFBC0" box="[306,336,1105,1129]" captionStart="FIGURES 20 – 37" captionStartId="13.[151,264,1716,1739]" captionTargetBox="[151,1435,193,1694]" captionTargetId="figure@13.[151,1436,193,1695]" captionTargetPageId="13" captionText="FIGURES 20 – 37. Fannia: male, hind leg, anterior view: (20) F. canicularis; (21) F. chibcha sp. nov., holotype; (22) F. chingaza sp. nov., holotype; (23) F. dodgei; (24) F. dorsomaculata sp. nov., holotype; (25) F. e m b e r a sp. nov., paratype; (26) F. grandis; (27) F. iguaque sp. nov., holotype; (28) F. katios sp. nov., holotype; (29) F. l a m o s c a sp. nov., holotype; (30) F. magdalena sp. nov., holotype; (31) F. pijao sp. nov., holotype; (32) F. p o rc e sp. nov., holotype; (33) F. pu s i o; (34) F. quimbaya sp. nov., holotype; (35) F. sumapaz sp. nov., holotype; (36) F. sutagao sp. nov., holotype; (37) F. trimaculata. Scale: 1 mm." httpUri="https://zenodo.org/record/213949/files/figure.png" pageId="33" pageNumber="34">31</figureCitation>
, 
<figureCitation id="132C2A6B5210D408C15E2462FEC0FBC0" box="[349,379,1105,1129]" captionStart="FIGURES 38 – 55" captionStartId="14.[151,264,1744,1767]" captionTargetBox="[151,1435,193,1723]" captionTargetId="figure@14.[151,1436,193,1723]" captionTargetPageId="14" captionText="FIGURES 38 – 55. Fannia: male, hind leg, posterior view: (38) F. canicularis; (39) F. chibcha sp. nov., holotype; (40) F. chingaza sp. nov., holotype; (41) F. dodgei; (42) F. dorsomaculata sp. nov., holotype; (43) F. e m b e r a sp. nov., holotype; (44) F. grandis; (45) F. iguaque sp. nov., holotype; (46) F. katios sp. nov., holotype; (47) F. l a m o s c a sp. nov., holotype; (48) F. magdalena sp. nov., holotype; (49) F. pijao sp. nov., holotype; (50) F. p o rc e sp. nov., holotype; (51) F. pu s i o; (52) F. quimbaya sp. nov., holotype; (53) F. sumapaz sp. nov., holotype; (54) F. sutagao sp. nov., holotype; (55) F. trimaculata. Scale: 1 mm." httpUri="https://zenodo.org/record/213950/files/figure.png" pageId="33" pageNumber="34">49</figureCitation>
, 
<figureCitation id="132C2A6B5210D408C1852462FE1FFBC0" box="[390,420,1105,1129]" captionStart="FIGURES 56 – 73" captionStartId="15.[151,264,1875,1898]" captionTargetBox="[200,1393,193,1844]" captionTargetId="figure@15.[200,1394,193,1845]" captionTargetPageId="15" captionText="FIGURES 56 – 73. Fannia: male, sternite 5, ventral view: (56) F. c a ni c ul a ri s; (57) F. c h i b c h a sp. nov., paratype; (58) F. chingaza sp. nov., holotype; (59) F. dodgei; (60) F. dorsomaculata sp. nov., paratype; (61) F. e m b e r a sp. nov., paratype; (62) F. grandis; (63) F. iguaque sp. nov., paratype; (64) F. katios sp. nov., paratype; (65) F. l a m o s c a sp. nov., paratype; (66) F. magdalena sp. nov., paratype; (67) F. pijao sp. nov., paratype; (68) F. porce sp. nov., holotype; (69) F. p u s i o; (70) F. quimbaya sp. nov., paratype; (71) F. sumapaz sp. nov., paratype; (72) F. sutagao sp. nov., paratype; (73) F. trimaculata. Scale: 0.1 mm." httpUri="https://zenodo.org/record/213951/files/figure.png" pageId="33" pageNumber="34">67</figureCitation>
, 
<figureCitation id="132C2A6B5210D408C1B32462FE75FBC0" box="[432,462,1105,1129]" captionStart="FIGURES 83 – 91" captionStartId="17.[151,264,1695,1718]" captionTargetBox="[168,1431,195,1670]" captionTargetId="figure@17.[168,1432,195,1670]" captionTargetPageId="17" captionText="FIGURES 83 – 91. Fannia: male, epandrium, cercal plate and surstyli, dorsal view: (83) F. l a m o s c a sp. nov., paratype; (84) F. magdalena sp. nov., paratype; (85) F. pijao sp. nov., paratype; (86) F. porce sp. nov., holotype; (87) F. p u s i o; (88) F. quimbaya sp. nov., paratype; (89) F. sumapaz sp. nov., paratype; (90) F. sutagao sp. nov., paratype; (91) F. trimaculata. Scale: 0.1 mm." httpUri="https://zenodo.org/record/213953/files/figure.png" pageId="33" pageNumber="34">85</figureCitation>
, 
<figureCitation id="132C2A6B5210D408C1D92462FDBDFBC0" box="[474,518,1105,1129]" captionStart="FIGURES 101 – 109" captionStartId="19.[151,264,1921,1944]" captionTargetBox="[186,1406,193,1902]" captionTargetId="figure@19.[183,1407,193,1903]" captionTargetPageId="19" captionText="FIGURES 101 – 109. Fannia: male, epandrium, cercal plate and surstyli, lateral view: (101) F. l a m os c a sp. nov., paratype; (102) F. magdalena sp. nov., paratype; (103) F. pijao sp. nov., paratype; (104) F. porce sp. nov., holotype; (105) F. p us i o; (106) F. quimbaya sp. nov., partype; (107) F. sumapaz sp. nov., paratype; (108) F. sutagao sp. nov., paratype; (109) F. trimaculata. Scale: 0.1 mm." httpUri="https://zenodo.org/record/213955/files/figure.png" pageId="33" pageNumber="34">103</figureCitation>
, 
<figureCitation id="132C2A6B5210D408C2112462FD85FBC0" box="[530,574,1105,1129]" captionStart="FIGURES 118 – 127" captionStartId="21.[151,264,1599,1622]" captionTargetBox="[168,1429,193,1576]" captionTargetId="figure@21.[165,1429,193,1577]" captionTargetPageId="21" captionText="FIGURES 118 – 127. Fannia: male, hypandrium and associated structures, dorsal view: (118) F. katios sp. nov., paratype; (119) F. l a m o s c a sp. nov., paratype; (120) F. magdalena sp. nov., paratype; (121) F. pijao sp. nov., paratype; (122) F. po rc e sp. nov., holotype; (123) F. p u s i o; (124) F. quimbaya sp. nov., paratype; (125) F. sumapaz sp. nov., paratype; (126) F. sutagao sp. nov., paratype; (127) F. trimaculata. Scale: 0.1 mm." httpUri="https://zenodo.org/record/213957/files/figure.png" pageId="33" pageNumber="34">121</figureCitation>
, 134)
</paragraph>
</subSubSection>
<subSubSection id="C30D65655210D408C09424AAFF54FACC" pageId="33" pageNumber="34" type="diagnosis">
<paragraph id="8BA836EE5210D408C09424AAFF54FACC" blockId="33.[151,1437,1177,2032]" pageId="33" pageNumber="34">
<emphasis id="B963EAFC5210D408C09424AAFEAAFB1B" bold="true" box="[151,273,1177,1202]" pageId="33" pageNumber="34">Diagnosis.</emphasis>
these characters apply only to the male sex. Scutum setulose; 
<emphasis id="B963EAFC5210D408C41724A9FB82FB18" box="[1044,1081,1178,1201]" italics="true" pageId="33" pageNumber="34">acr</emphasis>
0:0; 
<emphasis id="B963EAFC5210D408C47C24AAFB21FB18" box="[1151,1178,1177,1201]" italics="true" pageId="33" pageNumber="34">dc</emphasis>
0:2; fore tarsus with tarsomeres 4 and 5 slightly flattened; hind femur on ventral surface weakly curved, dorsal surface with weak preapical protuberance; anteroventral surface with weak protuberance on apical half and 1 row of setae that increase in length progressively; set of 
<emphasis id="B963EAFC5210D408C1D72535FE5BFAB4" box="[468,480,1286,1309]" italics="true" pageId="33" pageNumber="34">v</emphasis>
setae forming a weak tuft with 
<emphasis id="B963EAFC5210D408C3532535FCD1FAB4" box="[848,874,1286,1309]" italics="true" pageId="33" pageNumber="34">av</emphasis>
; posterior surface on basal third setulose bearing weak setae with hooked apices, apical third with row of more developed setae running towards posteroventral surface.
</paragraph>
</subSubSection>
<subSubSection id="C30D65655210D40BC0C42542FB20F985" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34" type="description">
<paragraph id="8BA836EE5210D408C0C42542FC55FA20" blockId="33.[151,1437,1177,2032]" box="[199,1006,1393,1418]" pageId="33" pageNumber="34">
<emphasis id="B963EAFC5210D408C0C42542FEEAFA23" bold="true" box="[199,337,1393,1418]" pageId="33" pageNumber="34">Description</emphasis>
. 
<emphasis id="B963EAFC5210D408C15C2541FDB7FA23" bold="true" box="[351,524,1394,1418]" pageId="33" pageNumber="34">
<typeStatus id="54AC884C5210D408C15C2541FE77FA23" box="[351,460,1394,1418]" pageId="33" pageNumber="34" type="holotype">Holotype</typeStatus>
male
</emphasis>
: body length 
<quantity id="4CEF9B0B5210D408C2A42541FD50FA20" box="[679,747,1394,1417]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="33" pageNumber="34" unit="mm" value="7.0">7 mm</quantity>
; wing length 5.75 mm.
</paragraph>
<paragraph id="8BA836EE5210D408C0C425A6FD42F9C8" blockId="33.[151,1437,1177,2032]" pageId="33" pageNumber="34">
<emphasis id="B963EAFC5210D408C0C425A6FEBFFA04" box="[199,260,1429,1453]" italics="true" pageId="33" pageNumber="34">Head</emphasis>
(
<figureCitation id="132C2A6B5210D408C11025A6FEDDFA04" box="[275,358,1429,1453]" captionStart="FIGURES 13 – 19" captionStartId="9.[151,264,858,881]" captionTargetBox="[153,1434,193,836]" captionTargetId="figure@9.[153,1435,193,837]" captionTargetPageId="9" captionText="FIGURES 13 – 19. Fannia: (13) F. magdalena sp. nov., male holotype, lateral view; (14) F. pijao sp. nov., male holotype, head, anterior view; (15) F. pijao sp. nov., male holotype, thorax, lateral view; (16) F. p o rc e sp. nov., male holotype, head, lateral view; (17) F. q u i m b a y a sp. nov., male holotype, head, lateral view; (18) F. q u i m b a y a sp. nov., male holotype, thorax, lateral view; (19) F. sutagao sp. nov., male holotype, head, anterior view. Scale: 1 mm." httpUri="https://zenodo.org/record/213948/files/figure.png" pageId="33" pageNumber="34">Fig. 14</figureCitation>
): eye densely setulose. Frontal vitta velvety brown. Interocular space ample, 0.15 mm. 23–24 
<emphasis id="B963EAFC5210D408C58725A6FA2EFA04" box="[1412,1429,1429,1453]" italics="true" pageId="33" pageNumber="34">fr</emphasis>
. Ocellar triangle black; 
<emphasis id="B963EAFC5210D408C1942589FE0AFA78" box="[407,433,1466,1489]" italics="true" pageId="33" pageNumber="34">oc</emphasis>
proclinate with 5 less developed setulae posteriorly. 
<emphasis id="B963EAFC5210D408C3FC2589FB90FA78" box="[1023,1067,1466,1489]" italics="true" pageId="33" pageNumber="34">Poc</emphasis>
parallel. Fronto-orbital plate and parafacial brown grey pollinose. Parafacial bare, narrowing on ventral third. Fronto-genal suture conspicuously brownish pollinose. Gena grey. Scape dark brown, 3 short setae reaching the pedicel. Pedicel dark brown, with dorsal setae short. Postpedicel golden-brownish pilose, 2.3 times the length of pedicel. Arista brown, pubescent. Palpus brown, claviform and long, with apex concave.
</paragraph>
<paragraph id="8BA836EE5210D408C0C4265EFCE2F958" blockId="33.[151,1437,1177,2032]" pageId="33" pageNumber="34">
<emphasis id="B963EAFC5210D408C0C4265EFEA3F92C" box="[199,280,1645,1669]" italics="true" pageId="33" pageNumber="34">Thorax</emphasis>
: dark brown; scutum without vittae. 
<emphasis id="B963EAFC5210D408C2BB265EFD5BF92C" box="[696,736,1645,1669]" italics="true" pageId="33" pageNumber="34">Acr</emphasis>
0:0, with ground setulae along 
<emphasis id="B963EAFC5210D408C447265DFBD2F92C" box="[1092,1129,1646,1669]" italics="true" pageId="33" pageNumber="34">acr</emphasis>
; pre-scutellar setae weakly developed. 
<emphasis id="B963EAFC5210D408C11926A1FE80F900" box="[282,315,1682,1705]" italics="true" pageId="33" pageNumber="34">Dc</emphasis>
0:2, ground setulae along 
<emphasis id="B963EAFC5210D408C26D26A2FD33F900" box="[622,648,1681,1705]" italics="true" pageId="33" pageNumber="34">dc</emphasis>
row (
<figureCitation id="132C2A6B5210D408C2CC26A2FC9FF900" box="[719,804,1681,1705]" captionStart="FIGURES 13 – 19" captionStartId="9.[151,264,858,881]" captionTargetBox="[153,1434,193,836]" captionTargetId="figure@9.[153,1435,193,837]" captionTargetPageId="9" captionText="FIGURES 13 – 19. Fannia: (13) F. magdalena sp. nov., male holotype, lateral view; (14) F. pijao sp. nov., male holotype, head, anterior view; (15) F. pijao sp. nov., male holotype, thorax, lateral view; (16) F. p o rc e sp. nov., male holotype, head, lateral view; (17) F. q u i m b a y a sp. nov., male holotype, head, lateral view; (18) F. q u i m b a y a sp. nov., male holotype, thorax, lateral view; (19) F. sutagao sp. nov., male holotype, head, anterior view. Scale: 1 mm." httpUri="https://zenodo.org/record/213948/files/figure.png" pageId="33" pageNumber="34">Fig. 15</figureCitation>
). Two differentiated 
<emphasis id="B963EAFC5210D408C41626A1FBF2F900" box="[1045,1097,1682,1705]" italics="true" pageId="33" pageNumber="34">pprn</emphasis>
. One developed 
<emphasis id="B963EAFC5210D408C50E26A1FA98F900" box="[1293,1315,1682,1705]" italics="true" pageId="33" pageNumber="34">ia</emphasis>
. Two 
<emphasis id="B963EAFC5210D408C56D26A1FA2FF900" box="[1390,1428,1682,1705]" italics="true" pageId="33" pageNumber="34">pra</emphasis>
, longer seta near suture. One differentiated 
<emphasis id="B963EAFC5210D408C2722685FD0CF964" box="[625,695,1718,1741]" italics="true" pageId="33" pageNumber="34">prepm</emphasis>
. Pre-basal scutellar area setulose. Subapical scutellar setae absent. Discal scutellar setae absent. One row of lateral scutellar setae.
</paragraph>
<paragraph id="8BA836EE5210D408C0C426CDFCE8F895" blockId="33.[151,1437,1177,2032]" pageId="33" pageNumber="34">
<emphasis id="B963EAFC5210D408C0C426CDFF44F8BC" box="[199,255,1790,1813]" italics="true" pageId="33" pageNumber="34">Wing</emphasis>
: brown, with the upper third, between C and R2+3 veins, darker. Calypters brownish with margins slightly darker. Haltere brown with base yellowish and stem brownish.
</paragraph>
<paragraph id="8BA836EE5210D40BC0C4277AFC84FD0E" blockId="33.[151,1437,1177,2032]" lastBlockId="34.[151,1436,151,2012]" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
<emphasis id="B963EAFC5210D408C0C4277AFF47F8C9" box="[199,252,1865,1888]" italics="true" pageId="33" pageNumber="34">Legs</emphasis>
: dark brown, with apex of femur and tibiae slightly yellowish, pulvilli yellowish. Fore femur with 1 row of long 
<emphasis id="B963EAFC5210D408C0EC275FFF45F82D" box="[239,254,1900,1924]" italics="true" pageId="33" pageNumber="34">d</emphasis>
with straight apices; ventral surface bare; 1 row of long 
<emphasis id="B963EAFC5210D408C383275EFC22F82D" box="[896,921,1901,1924]" italics="true" pageId="33" pageNumber="34">pv</emphasis>
, setae on apical half longer; posterior surface densely setulose, bearing long and weak setae with curved apices. Fore tibia with 1 pre-apical 
<emphasis id="B963EAFC5210D408C4C827A3FB62F801" box="[1227,1241,1936,1960]" italics="true" pageId="33" pageNumber="34">d</emphasis>
; 1 
<emphasis id="B963EAFC5210D408C4FC27A3FAA0F801" box="[1279,1307,1936,1960]" italics="true" pageId="33" pageNumber="34">ad</emphasis>
on edge of apical third; 1 apical 
<emphasis id="B963EAFC5210D408C18B2786FE2EF865" box="[392,405,1973,1996]" italics="true" pageId="33" pageNumber="34">v</emphasis>
; 1 pre-apical 
<emphasis id="B963EAFC5210D408C2302786FDECF865" box="[563,599,1973,1996]" italics="true" pageId="33" pageNumber="34">pv;</emphasis>
1 
<emphasis id="B963EAFC5210D408C2762786FD38F865" box="[629,643,1973,1996]" italics="true" pageId="33" pageNumber="34">p</emphasis>
on edge of apical third. Fore tarsus with strong 
<emphasis id="B963EAFC5210D408C4AA2786FB0EF865" box="[1193,1205,1973,1996]" italics="true" pageId="33" pageNumber="34">v</emphasis>
setae followed by 1 thin seta on first tarsomere; tarsomeres 4 and 5 slightly flattened. Mid femur with a weak constriction on pre-apical ventral surface; 1 row of 
<emphasis id="B963EAFC5213D40BC1B320ABFE77FF19" box="[432,460,152,176]" italics="true" pageId="34" pageNumber="35">ad</emphasis>
, long and developed on pre-apex;1 row of 
<emphasis id="B963EAFC5213D40BC3AF20AAFC7CFF19" box="[940,967,153,176]" italics="true" pageId="34" pageNumber="35">av</emphasis>
that increase in length towards apex, then decrease in length on apical half where they have hooked apices; 1 row of 
<emphasis id="B963EAFC5213D40BC420208EFB94FF7D" box="[1059,1071,189,212]" italics="true" pageId="34" pageNumber="35">v</emphasis>
with hooked apices; 1 row of developed 
<emphasis id="B963EAFC5213D40BC11120D2FE96FF51" box="[274,301,225,248]" italics="true" pageId="34" pageNumber="35">pv</emphasis>
with hooked apices, 
<emphasis id="B963EAFC5213D40BC22320D2FD81FF51" box="[544,570,225,248]" italics="true" pageId="34" pageNumber="35">pv</emphasis>
on apical third short and forming a small ctenidium; 1 row of weak, long 
<emphasis id="B963EAFC5213D40BC58D20D2FA27FF51" box="[1422,1436,225,248]" italics="true" pageId="34" pageNumber="35">p</emphasis>
with curved apices, on apical third with longer setae with straight apices running towards posteriodorsal surface. Mid tibia on ventral surface on basal third with a weak constriction, dorsal surface densely setulose on apical half, 1 long apical 
<emphasis id="B963EAFC5213D40BC12D217EFE81FECD" box="[302,314,333,356]" italics="true" pageId="34" pageNumber="35">v</emphasis>
; 1 sub-median 
<emphasis id="B963EAFC5213D40BC1EB217FFDBFFECD" box="[488,516,332,356]" italics="true" pageId="34" pageNumber="35">ad</emphasis>
and 1 developed pre-apical 
<emphasis id="B963EAFC5213D40BC343217FFCF5FECD" box="[832,846,332,356]" italics="true" pageId="34" pageNumber="35">d</emphasis>
; 1 short pre-apical and 1 long apical 
<emphasis id="B963EAFC5213D40BC4EC217EFB46FECD" box="[1263,1277,333,356]" italics="true" pageId="34" pageNumber="35">a</emphasis>
; 1 long apical 
<emphasis id="B963EAFC5213D40BC0942142FF09FE21" box="[151,178,369,392]" italics="true" pageId="34" pageNumber="35">av</emphasis>
; 1 median 
<emphasis id="B963EAFC5213D40BC1322142FE84FE21" box="[305,319,369,392]" italics="true" pageId="34" pageNumber="35">p</emphasis>
and 1 on edge of apical third; 1 short pre-apical 
<emphasis id="B963EAFC5213D40BC3642143FC38FE21" box="[871,899,368,392]" italics="true" pageId="34" pageNumber="35">pd</emphasis>
; 1 apical 
<emphasis id="B963EAFC5213D40BC3F02142FBB7FE21" box="[1011,1036,369,392]" italics="true" pageId="34" pageNumber="35">pv</emphasis>
. Mid tarsus with 2 strong 
<emphasis id="B963EAFC5213D40BC5342142FAF8FE21" box="[1335,1347,369,392]" italics="true" pageId="34" pageNumber="35">v</emphasis>
on fore tarsomere. 
<collectingCountry id="F300767E5213D40BC11721A7FEF7FE05" box="[276,332,404,428]" name="India" pageId="34" pageNumber="35">Hind</collectingCountry>
coxa on posterior margin bare. 
<collectingCountry id="F300767E5213D40BC2B621A7FD56FE05" box="[693,749,404,428]" name="India" pageId="34" pageNumber="35">Hind</collectingCountry>
femur on ventral surface weakly curved; dorsal surface with weak pre-apical protuberance; anteroventral surface with weak protuberance on apical half, 1 row of 
<emphasis id="B963EAFC5213D40BC548218AFADDFE79" box="[1355,1382,441,464]" italics="true" pageId="34" pageNumber="35">av</emphasis>
that increase in length progressively (
<figureCitation id="132C2A6B5213D40BC21121EFFDDDFE5D" box="[530,614,476,500]" captionStart="FIGURES 20 – 37" captionStartId="13.[151,264,1716,1739]" captionTargetBox="[151,1435,193,1694]" captionTargetId="figure@13.[151,1436,193,1695]" captionTargetPageId="13" captionText="FIGURES 20 – 37. Fannia: male, hind leg, anterior view: (20) F. canicularis; (21) F. chibcha sp. nov., holotype; (22) F. chingaza sp. nov., holotype; (23) F. dodgei; (24) F. dorsomaculata sp. nov., holotype; (25) F. e m b e r a sp. nov., paratype; (26) F. grandis; (27) F. iguaque sp. nov., holotype; (28) F. katios sp. nov., holotype; (29) F. l a m o s c a sp. nov., holotype; (30) F. magdalena sp. nov., holotype; (31) F. pijao sp. nov., holotype; (32) F. p o rc e sp. nov., holotype; (33) F. pu s i o; (34) F. quimbaya sp. nov., holotype; (35) F. sumapaz sp. nov., holotype; (36) F. sutagao sp. nov., holotype; (37) F. trimaculata. Scale: 1 mm." httpUri="https://zenodo.org/record/213949/files/figure.png" pageId="34" pageNumber="35">Fig. 31</figureCitation>
), on apical third with 1 row of weak setae which end with a developed seta; ventral surface with a set of setae forming a small tuft with the 
<emphasis id="B963EAFC5213D40BC3942232FC0AFDB1" box="[919,945,513,536]" italics="true" pageId="34" pageNumber="35">av</emphasis>
; 1 row of 
<emphasis id="B963EAFC5213D40BC4252233FBF9FDB1" box="[1062,1090,512,536]" italics="true" pageId="34" pageNumber="35">ad</emphasis>
that increase in length and run towards dorsal surface; posterior surface on basal third setulose, bearing weak setae with hooked apices, apical third with 1 row of more developed setae going towards posteroventral surface (
<figureCitation id="132C2A6B5213D40BC4402274FB20FDF6" box="[1091,1179,583,607]" captionStart="FIGURES 38 – 55" captionStartId="14.[151,264,1744,1767]" captionTargetBox="[151,1435,193,1723]" captionTargetId="figure@14.[151,1436,193,1723]" captionTargetPageId="14" captionText="FIGURES 38 – 55. Fannia: male, hind leg, posterior view: (38) F. canicularis; (39) F. chibcha sp. nov., holotype; (40) F. chingaza sp. nov., holotype; (41) F. dodgei; (42) F. dorsomaculata sp. nov., holotype; (43) F. e m b e r a sp. nov., holotype; (44) F. grandis; (45) F. iguaque sp. nov., holotype; (46) F. katios sp. nov., holotype; (47) F. l a m o s c a sp. nov., holotype; (48) F. magdalena sp. nov., holotype; (49) F. pijao sp. nov., holotype; (50) F. p o rc e sp. nov., holotype; (51) F. pu s i o; (52) F. quimbaya sp. nov., holotype; (53) F. sumapaz sp. nov., holotype; (54) F. sutagao sp. nov., holotype; (55) F. trimaculata. Scale: 1 mm." httpUri="https://zenodo.org/record/213950/files/figure.png" pageId="34" pageNumber="35">Fig. 49</figureCitation>
). 
<collectingCountry id="F300767E5213D40BC4B12274FB51FDF6" box="[1202,1258,583,607]" name="India" pageId="34" pageNumber="35">Hind</collectingCountry>
tibia on dorsal surface curved, 1 median and 1 pre-apical 
<emphasis id="B963EAFC5213D40BC277225FFD39FD2D" box="[628,642,620,644]" italics="true" pageId="34" pageNumber="35">d</emphasis>
developed; 1 sub-median 
<emphasis id="B963EAFC5213D40BC3A9225FFC7DFD2D" box="[938,966,620,644]" italics="true" pageId="34" pageNumber="35">ad</emphasis>
and 1 row of long 
<emphasis id="B963EAFC5213D40BC49F225FFB11FD2D" box="[1180,1194,620,644]" italics="true" pageId="34" pageNumber="35">d</emphasis>
on basal half; 1 short pre-apical 
<emphasis id="B963EAFC5213D40BC10D22A2FEA7FD01" box="[270,284,657,680]" italics="true" pageId="34" pageNumber="35">a</emphasis>
; 2 median 
<emphasis id="B963EAFC5213D40BC19A22A2FE1CFD01" box="[409,423,657,680]" italics="true" pageId="34" pageNumber="35">a</emphasis>
and 1 apical 
<emphasis id="B963EAFC5213D40BC23F22A2FDF3FD01" box="[572,584,657,680]" italics="true" pageId="34" pageNumber="35">v</emphasis>
(
<figureCitation id="132C2A6B5213D40BC25B22BCFD10FD0E" box="[600,683,655,679]" captionStart="FIGURES 20 – 37" captionStartId="13.[151,264,1716,1739]" captionTargetBox="[151,1435,193,1694]" captionTargetId="figure@13.[151,1436,193,1695]" captionTargetPageId="13" captionText="FIGURES 20 – 37. Fannia: male, hind leg, anterior view: (20) F. canicularis; (21) F. chibcha sp. nov., holotype; (22) F. chingaza sp. nov., holotype; (23) F. dodgei; (24) F. dorsomaculata sp. nov., holotype; (25) F. e m b e r a sp. nov., paratype; (26) F. grandis; (27) F. iguaque sp. nov., holotype; (28) F. katios sp. nov., holotype; (29) F. l a m o s c a sp. nov., holotype; (30) F. magdalena sp. nov., holotype; (31) F. pijao sp. nov., holotype; (32) F. p o rc e sp. nov., holotype; (33) F. pu s i o; (34) F. quimbaya sp. nov., holotype; (35) F. sumapaz sp. nov., holotype; (36) F. sutagao sp. nov., holotype; (37) F. trimaculata. Scale: 1 mm." httpUri="https://zenodo.org/record/213949/files/figure.png" pageId="34" pageNumber="35">Fig. 31</figureCitation>
); 1 apical 
<emphasis id="B963EAFC5213D40BC31C22A2FC8CFD01" box="[799,823,657,680]" italics="true" pageId="34" pageNumber="35">pv</emphasis>
.
</paragraph>
<paragraph id="8BA836EE5213D40BC0C42287FAE3FC0D" blockId="34.[151,1436,151,2012]" pageId="34" pageNumber="35">
<emphasis id="B963EAFC5213D40BC0C42287FE8AFD65" box="[199,305,692,716]" italics="true" pageId="34" pageNumber="35">Abdomen</emphasis>
: elongate, in general with developed setae which are not differentiated from the ground setulae, brown and pollinose. Syntergite 1+2, tergites 3 and 4 translucent-brown, with central vitta dark brown. Syntergite 1+2 with a set of lateral basal long setae. Sternite 1 densely setulose. Sternite 5 as in 
<figureCitation id="132C2A6B5213D40BC45022CFFB1EFCBD" box="[1107,1189,764,788]" captionStart="FIGURES 56 – 73" captionStartId="15.[151,264,1875,1898]" captionTargetBox="[200,1393,193,1844]" captionTargetId="figure@15.[200,1394,193,1845]" captionTargetPageId="15" captionText="FIGURES 56 – 73. Fannia: male, sternite 5, ventral view: (56) F. c a ni c ul a ri s; (57) F. c h i b c h a sp. nov., paratype; (58) F. chingaza sp. nov., holotype; (59) F. dodgei; (60) F. dorsomaculata sp. nov., paratype; (61) F. e m b e r a sp. nov., paratype; (62) F. grandis; (63) F. iguaque sp. nov., paratype; (64) F. katios sp. nov., paratype; (65) F. l a m o s c a sp. nov., paratype; (66) F. magdalena sp. nov., paratype; (67) F. pijao sp. nov., paratype; (68) F. porce sp. nov., holotype; (69) F. p u s i o; (70) F. quimbaya sp. nov., paratype; (71) F. sumapaz sp. nov., paratype; (72) F. sutagao sp. nov., paratype; (73) F. trimaculata. Scale: 0.1 mm." httpUri="https://zenodo.org/record/213951/files/figure.png" pageId="34" pageNumber="35">Fig. 67</figureCitation>
. Terminalia (
<figureCitation id="132C2A6B5213D40BC53A22CFFA2CFCBD" box="[1337,1431,764,788]" captionStart="FIGURES 83 – 91" captionStartId="17.[151,264,1695,1718]" captionTargetBox="[168,1431,195,1670]" captionTargetId="figure@17.[168,1432,195,1670]" captionTargetPageId="17" captionText="FIGURES 83 – 91. Fannia: male, epandrium, cercal plate and surstyli, dorsal view: (83) F. l a m o s c a sp. nov., paratype; (84) F. magdalena sp. nov., paratype; (85) F. pijao sp. nov., paratype; (86) F. porce sp. nov., holotype; (87) F. p u s i o; (88) F. quimbaya sp. nov., paratype; (89) F. sumapaz sp. nov., paratype; (90) F. sutagao sp. nov., paratype; (91) F. trimaculata. Scale: 0.1 mm." httpUri="https://zenodo.org/record/213953/files/figure.png" pageId="34" pageNumber="35">Figs. 85</figureCitation>
, 
<figureCitation id="132C2A6B5213D40BC094232CFF7EFC9E" box="[151,197,799,823]" captionStart="FIGURES 101 – 109" captionStartId="19.[151,264,1921,1944]" captionTargetBox="[186,1406,193,1902]" captionTargetId="figure@19.[183,1407,193,1903]" captionTargetPageId="19" captionText="FIGURES 101 – 109. Fannia: male, epandrium, cercal plate and surstyli, lateral view: (101) F. l a m os c a sp. nov., paratype; (102) F. magdalena sp. nov., paratype; (103) F. pijao sp. nov., paratype; (104) F. porce sp. nov., holotype; (105) F. p us i o; (106) F. quimbaya sp. nov., partype; (107) F. sumapaz sp. nov., paratype; (108) F. sutagao sp. nov., paratype; (109) F. trimaculata. Scale: 0.1 mm." httpUri="https://zenodo.org/record/213955/files/figure.png" pageId="34" pageNumber="35">103</figureCitation>
): epandrium wider than long with two median sclerotized and pointed projections, basal half with large amounts of thin setae, not very long; cercal plate sclerotized and weakly divided basally, elongate apically and with short setae; surstylus laterally articulated with epandrium, without basal sclerotized projections, apical half slender, curved and with short setae; bacilliform process absent; hypandrium and associated structures as in 
<figureCitation id="132C2A6B5213D40BC4F023BFFAEFFC0D" box="[1267,1364,908,932]" captionStart="FIGURES 118 – 127" captionStartId="21.[151,264,1599,1622]" captionTargetBox="[168,1429,193,1576]" captionTargetId="figure@21.[165,1429,193,1577]" captionTargetPageId="21" captionText="FIGURES 118 – 127. Fannia: male, hypandrium and associated structures, dorsal view: (118) F. katios sp. nov., paratype; (119) F. l a m o s c a sp. nov., paratype; (120) F. magdalena sp. nov., paratype; (121) F. pijao sp. nov., paratype; (122) F. po rc e sp. nov., holotype; (123) F. p u s i o; (124) F. quimbaya sp. nov., paratype; (125) F. sumapaz sp. nov., paratype; (126) F. sutagao sp. nov., paratype; (127) F. trimaculata. Scale: 0.1 mm." httpUri="https://zenodo.org/record/213957/files/figure.png" pageId="34" pageNumber="35">Fig. 121</figureCitation>
.
</paragraph>
<paragraph id="8BA836EE5213D40BC0C4239CFDDFFC45" blockId="34.[151,1436,151,2012]" pageId="34" pageNumber="35">
<emphasis id="B963EAFC5213D40BC0C4239CFE85FC61" bold="true" box="[199,318,943,968]" pageId="34" pageNumber="35">Variation:</emphasis>
fronto-orbital plate greyish pollinose; parafacial brownish pollinose; legs completely dark brown; fore tarsus with tarsomeres not modified.
</paragraph>
<paragraph id="8BA836EE5213D40BC0C423CBFB20F985" blockId="34.[151,1436,151,2012]" pageId="34" pageNumber="35">
<emphasis id="B963EAFC5213D40BC0C423CBFEA5FBB9" bold="true" box="[199,286,1016,1040]" pageId="34" pageNumber="35">Female</emphasis>
: similar to male except: body length 5.5– 
<quantity id="4CEF9B0B5213D40BC31B23C4FCDBFBB9" box="[792,864,1015,1040]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="34" pageNumber="35" unit="mm" value="6.0">6 mm</quantity>
; wing length 5.5–5.7 mm. Setae in general less developed. Eye with smaller and sparse setulae. Interocular space ample, 0.6 mm. 11 
<emphasis id="B963EAFC5213D40BC475242FFB33FB9D" box="[1142,1160,1052,1076]" italics="true" pageId="34" pageNumber="35">fr</emphasis>
, with 1–2 pairs slightly more developed. Interfrontal setae 2. Ocellar triangle grey. Parafacial greyish and brownish pollinose. Lunule brown. Scape with two setae. Thorax with 2–3 
<emphasis id="B963EAFC5213D40BC2A92456FD6AFBD5" box="[682,721,1125,1148]" italics="true" pageId="34" pageNumber="35">pra</emphasis>
, longer seta near the suture. Differentiated proepimeral setae 2. Wing paler, but with upper third as in male. Calypters yellowish with margin darker. Haltere with base brown and stem yellowish. Fore leg with 1 row of long 
<emphasis id="B963EAFC5213D40BC288249EFD1DFB6D" box="[651,678,1197,1220]" italics="true" pageId="34" pageNumber="35">pv</emphasis>
; posterior surface with setae, not densely setulose. Fore tarsus with tarsomere 5 slightly flattened. Mid femur with weak constriction on pre-apical ventral surface; anteroventral surface without differentiated setae; ventral surface without row of setae; 1 row of weak, long and uniform 
<emphasis id="B963EAFC5213D40BC54424C6FAD9FAA5" box="[1351,1378,1269,1292]" italics="true" pageId="34" pageNumber="35">pv</emphasis>
; mid tibia on ventral surface without constriction and weakly setulose; 1 median and 1 pre-apical developed 
<emphasis id="B963EAFC5213D40BC55D252BFAC1FA99" box="[1374,1402,1304,1328]" italics="true" pageId="34" pageNumber="35">ad</emphasis>
; 2 median 
<emphasis id="B963EAFC5213D40BC0F1250EFEBBFAFD" box="[242,256,1341,1364]" italics="true" pageId="34" pageNumber="35">p</emphasis>
or 1 median 
<emphasis id="B963EAFC5213D40BC19B250EFE1DFAFD" box="[408,422,1341,1364]" italics="true" pageId="34" pageNumber="35">p</emphasis>
and 2 extra on apical third. 
<collectingCountry id="F300767E5213D40BC2EA250FFC9AFAFD" box="[745,801,1340,1364]" name="India" pageId="34" pageNumber="35">Hind</collectingCountry>
femur more widened than in male; ventral surface with ground setulae up to basal half, not forming a tuft; posterior surface without differentiated setae. 
<collectingCountry id="F300767E5213D40BC4DF256CFAAFFADE" box="[1244,1300,1375,1399]" name="India" pageId="34" pageNumber="35">Hind</collectingCountry>
tibia with 1 row of short 
<emphasis id="B963EAFC5213D40BC12A25B7FEFEFA35" box="[297,325,1412,1436]" italics="true" pageId="34" pageNumber="35">ad</emphasis>
on basal half. Abdomen completely dark brown. Sternites weakly setulose. Terminalia (Fig. 134): cercus narrower and long than hypoproct, with long setae on apex; hypoproct more or less triangular and with short and sinuous setae; sternite 8 reduced to two membranous plates with short and curved setae and a long seta, two large more sclerotized and bare basal plates, near sternite 7; sternites 7 and 6 sub equal in size, wider than long and with long apical setae. Spermathecae 3, semicircular, one smaller than the others (Fig. 134).
</paragraph>
</subSubSection>
<subSubSection id="C30D65655213D40BC0C42604FE5FF8AD" pageId="34" pageNumber="35" type="biology_ecology">
<paragraph id="8BA836EE5213D40BC0C42604FE5FF8AD" blockId="34.[151,1436,151,2012]" pageId="34" pageNumber="35">
<emphasis id="B963EAFC5213D40BC0C42604FE9CF9F9" bold="true" box="[199,295,1591,1616]" pageId="34" pageNumber="35">Biology.</emphasis>
according to label data, 
<taxonomicName id="4C174D6D5213D40BC23F260AFD75F9F9" box="[572,718,1593,1616]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="pijao" status="sp. nov.">
<emphasis id="B963EAFC5213D40BC23F260AFD75F9F9" box="[572,718,1593,1616]" italics="true" pageId="34" pageNumber="35">Fannia pijao</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25057875213D40BC2D6260BFC95F9F9" box="[725,814,1592,1616]" pageId="34" pageNumber="35" rank="species">
<emphasis id="B963EAFC5213D40BC2D6260BFD54F9F9" bold="true" box="[725,751,1592,1616]" pageId="34" pageNumber="35">sp</emphasis>
. 
<emphasis id="B963EAFC5213D40BC2FE260BFC9CF9F9" bold="true" box="[765,807,1592,1616]" pageId="34" pageNumber="35">nov</emphasis>
.
</taxonomicNameLabel>
is associated with decomposing organic matter and the well-preserved high Andean forest at Reserva Forestal Nacional Protectora Rio Blanco. This ecosystem houses several species of mammals, birds, reptiles and river sources, as well as altitudes between 
<quantity id="4CEF9B0B5213D40BC4B5264CFAEBF931" box="[1206,1360,1663,1688]" metricMagnitude="3" metricUnit="m" metricValue="2.9699999999999998" metricValueMax="3.7" metricValueMin="2.2399999999999998" pageId="34" pageNumber="35" unit="m" value="2970.0" valueMax="3700.0" valueMin="2240.0">2240-3700 m</quantity>
in the Andine system, such as black cedar (
<taxonomicName id="4C174D6D5213D40BC2962697FCB6F915" box="[661,781,1700,1724]" class="Magnoliopsida" family="Meliaceae" kingdom="Plantae" order="Sapindales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="family">Meliaceae</taxonomicName>
, 
<taxonomicName id="4C174D6D5213D40BC32B2697FBDDF915" authority="Dode" authorityName="Dode" box="[808,1126,1700,1724]" class="Magnoliopsida" family="Juglandaceae" genus="Juglans" kingdom="Plantae" order="Fagales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="species" species="neotropica">
<emphasis id="B963EAFC5213D40BC32B2697FBB5F915" box="[808,1038,1700,1724]" italics="true" pageId="34" pageNumber="35">Juglans neotropica</emphasis>
Dode
</taxonomicName>
), Alstonville 
<taxonomicName id="4C174D6D5213D40BC51E2697FA20F915" box="[1309,1435,1700,1724]" class="Magnoliopsida" family="Melastomataceae" genus="Tibouchina" kingdom="Plantae" order="Myrtales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="genus">Tibouchina</taxonomicName>
(
<taxonomicName id="4C174D6D5213D40BC09D26F4FEDEF976" box="[158,357,1735,1759]" class="Magnoliopsida" family="Melastomataceae" kingdom="Plantae" order="Myrtales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="family">Melastomataceae</taxonomicName>
, 
<taxonomicName id="4C174D6D5213D40BC17326FBFD52F976" authority="Bondpland" authorityName="Bondpland" box="[368,745,1735,1760]" class="Magnoliopsida" family="Melastomataceae" genus="Tibouchina" kingdom="Plantae" order="Myrtales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="species" species="lepidota">
<emphasis id="B963EAFC5213D40BC17326FBFDEAF949" box="[368,593,1736,1760]" italics="true" pageId="34" pageNumber="35">Tibouchina lepidota</emphasis>
(Bondpland)
</taxonomicName>
, Rushfoil (
<taxonomicName id="4C174D6D5213D40BC36026F4FBB5F976" box="[867,1038,1735,1759]" class="Magnoliopsida" family="Euphorbiaceae" kingdom="Plantae" order="Malpighiales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="family">Euphorbiaceae</taxonomicName>
, 
<taxonomicName id="4C174D6D5213D40BC41A26FBFA34F976" authority="Mull. Arg" authorityName="Mull. Arg" box="[1049,1423,1735,1760]" class="Magnoliopsida" family="Euphorbiaceae" genus="Croton" kingdom="Plantae" order="Malpighiales" pageId="34" pageNumber="35" phylum="Tracheophyta" rank="species" species="magdalenensis">
<emphasis id="B963EAFC5213D40BC41A26FBFAAEF949" box="[1049,1301,1736,1760]" italics="true" pageId="34" pageNumber="35">Croton magdalenensis</emphasis>
Müll. Arg
</taxonomicName>
), among others (
<bibRefCitation id="EF864B1F5213D40BC13E26DFFE63F8AD" author="Giraldo" box="[317,472,1772,1796]" pageId="34" pageNumber="35" refString="Giraldo, S. A. (2009) Parque y Reservas Naturales de Manizales. Available from http: // parquesyreservasnaturalesdemanizales. blogspot. com / 2009 / 03 / parques-y-reservas-naturales-de. html (Accessed on 4 th April 2010)." type="journal article" year="2009">Giraldo 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30D65655213D40BC0C4273CFE4EF8D9" pageId="34" pageNumber="35" type="discussion">
<paragraph id="8BA836EE5213D40BC0C4273CFE4EF8D9" blockId="34.[151,1436,151,2012]" pageId="34" pageNumber="35">
<emphasis id="B963EAFC5213D40BC0C4273CFEF4F881" bold="true" box="[199,335,1807,1832]" pageId="34" pageNumber="35">Comments.</emphasis>
<taxonomicName id="4C174D6D5213D40BC15B2722FE56F881" box="[344,493,1809,1832]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="pijao" status="sp. nov.">
<emphasis id="B963EAFC5213D40BC15B2722FE56F881" box="[344,493,1809,1832]" italics="true" pageId="34" pageNumber="35">Fannia pijao</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25057875213D40BC1F52723FDE9F881" box="[502,594,1808,1832]" pageId="34" pageNumber="35" rank="species">
<emphasis id="B963EAFC5213D40BC1F52723FDABF881" bold="true" box="[502,528,1808,1832]" pageId="34" pageNumber="35">sp</emphasis>
. 
<emphasis id="B963EAFC5213D40BC2222723FDF0F881" bold="true" box="[545,587,1808,1832]" pageId="34" pageNumber="35">nov</emphasis>
.
</taxonomicNameLabel>
is morphologically close to 
<taxonomicName id="4C174D6D5213D40BC39F2722FBDEF881" box="[924,1125,1808,1832]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="quimbaya" status="sp. nov.">
<emphasis id="B963EAFC5213D40BC39F2722FBDEF881" box="[924,1125,1808,1832]" italics="true" pageId="34" pageNumber="35">Fannia quimbaya</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25057875213D40BC46C2723FB71F881" box="[1135,1226,1808,1832]" pageId="34" pageNumber="35" rank="species">
<emphasis id="B963EAFC5213D40BC46C2723FB32F881" bold="true" box="[1135,1161,1808,1832]" pageId="34" pageNumber="35">sp</emphasis>
. 
<emphasis id="B963EAFC5213D40BC4992723FB7FF881" bold="true" box="[1178,1220,1808,1832]" pageId="34" pageNumber="35">nov</emphasis>
.
</taxonomicNameLabel>
, but differs in the arrangement of the 
<emphasis id="B963EAFC5213D40BC1712707FE36F8E5" box="[370,397,1844,1868]" italics="true" pageId="34" pageNumber="35">dc</emphasis>
as 0:2. The combination of characters in these two species makes it difficult to include them in any known group of 
<taxonomicName id="4C174D6D5213D40BC19E276AFE55F8D9" box="[413,494,1881,1904]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B963EAFC5213D40BC19E276AFE55F8D9" box="[413,494,1881,1904]" italics="true" pageId="34" pageNumber="35">Fannia</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C30D65655213D40BC0C4274FFC13F875" pageId="34" pageNumber="35" type="etymology">
<paragraph id="8BA836EE5213D40BC0C4274FFC13F875" blockId="34.[151,1436,151,2012]" pageId="34" pageNumber="35">
<emphasis id="B963EAFC5213D40BC0C4274FFEF5F83D" bold="true" box="[199,334,1916,1940]" pageId="34" pageNumber="35">Etymology.</emphasis>
name in apposition. The species epithet refers to the indigenous people 
<taxonomicName id="4C174D6D5213D40BC48B274FFB7FF83D" box="[1160,1220,1916,1940]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="39" phylum="Arthropoda" rank="species" species="pijao" status="sp. nov.">Pijao</taxonomicName>
, who inhabited the Central Cordillera of the Colombian Andes in pre-Colombian times, between the snow peaks of Huila, Quindío and Tolima, the high 
<taxonomicName id="4C174D6D5213D40BC15527F7FE69F875" box="[342,466,1988,2012]" class="Insecta" family="Fanniidae" genus="Fannia" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="33" phylum="Arthropoda" rank="species" species="magdalena" status="sp. nov.">Magdalena</taxonomicName>
Valley and Valle del Cauca, in 
<collectingCountry id="F300767E5213D40BC33127F7FC1FF875" box="[818,932,1988,2012]" name="Colombia" pageId="34" pageNumber="35">Colombia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C30D65655212D40AC0C420ABFC90FE66" pageId="35" pageNumber="36" type="materials_examined">
<paragraph id="8BA836EE5212D40AC0C420ABFC90FE66" blockId="35.[151,1437,151,500]" pageId="35" pageNumber="36">
<emphasis id="B963EAFC5212D40AC0C420ABFEC2FF19" bold="true" box="[199,377,151,176]" pageId="35" pageNumber="36">
<typeStatus id="54AC884C5212D40AC0C420ABFEB9FF19" box="[199,258,152,176]" pageId="35" pageNumber="36">Type</typeStatus>
material
</emphasis>
. 
<emphasis id="B963EAFC5212D40AC19120ABFE44FF19" bold="true" box="[402,511,152,176]" pageId="35" pageNumber="36">
<typeStatus id="54AC884C5212D40AC19120ABFE44FF19" box="[402,511,152,176]" pageId="35" pageNumber="36" type="holotype">Holotype</typeStatus>
</emphasis>
male (
<collectionCode id="ED06AE2B5212D40AC26220A4FD0FFF06" box="[609,692,151,175]" pageId="35" pageNumber="36">CEUA</collectionCode>
). CO [
<collectingCountry id="F300767E5212D40AC31120A4FC31FF06" box="[786,906,151,175]" name="Colombia" pageId="35" pageNumber="36">Colombia</collectingCountry>
]. Caldas. Manizales. Reserva Rio\ Blanco. 5˚ 
<date id="FFA9102E5212D40AC0AA208FFEB8FF7D" box="[169,259,188,212]" pageId="35" pageNumber="36" value="58-07-26">7.26.58</date>
”N 75˚43.7’9.58”W\ 2592 msnm. T VSR - pez [Van Someren-Rydon trap with fish] Bosque\ 
<date id="FFA9102E5212D40AC542208FFF74FF5E" pageId="35" pageNumber="36" value="2005-04-03">Abril 3/ 2005</date>
G. Entomologia\ 
<collectionCode id="ED06AE2B5212D40AC1AF20ECFE42FF5E" box="[428,505,223,247]" pageId="35" pageNumber="36">CEUA</collectionCode>
7498. 
<emphasis id="B963EAFC5212D40AC25620D3FD6DFF51" bold="true" box="[597,726,224,248]" pageId="35" pageNumber="36">
<typeStatus id="54AC884C5212D40AC25620D3FD6AFF51" box="[597,721,224,248]" pageId="35" pageNumber="36" type="paratype">Paratypes</typeStatus>
:
</emphasis>
CO [
<collectingCountry id="F300767E5212D40AC32120ECFC21FF5E" box="[802,922,223,247]" name="Colombia" pageId="35" pageNumber="36">Colombia</collectingCountry>
]. Caldas. Manizales. Reserva Rio\ Blanco. 5˚ 
<date id="FFA9102E5212D40AC0AA2137FEB8FEB5" box="[169,259,260,284]" pageId="35" pageNumber="36" value="58-07-26">7.26.58</date>
”N 75˚43.7’9.58”W\ 2592 msnm.T VSR - pez [Van Someren-Rydon trap with fish] Bosque\ 
<date id="FFA9102E5212D40AC5422137FF74FE96" pageId="35" pageNumber="36" value="2005-04-03">Abril 3/ 2005</date>
G. Entomologia\ 
<collectionCode id="ED06AE2B5212D40AC1992114FE51FE96" box="[410,490,295,319]" pageId="35" pageNumber="36">CEUA</collectionCode>
. (
<specimenCount id="9D11FD675212D40AC1FC2114FDD3FE96" box="[511,616,295,319]" pageId="35" pageNumber="36" type="male">11 males</specimenCount>
, 
<collectionCode id="ED06AE2B5212D40AC2762114FD7EFE96" box="[629,709,295,319]" pageId="35" pageNumber="36">CEUA</collectionCode>
; 
<specimenCount id="9D11FD675212D40AC2D12114FCA4FE96" box="[722,799,295,319]" pageId="35" pageNumber="36" type="male">1 male</specimenCount>
<collectionCode id="ED06AE2B5212D40AC32A2114FCCCFE96" box="[809,887,295,319]" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="35" pageNumber="36">DZUP</collectionCode>
); “idem \ 
<collectionCode id="ED06AE2B5212D40AC3E82114FB8CFE96" box="[1003,1079,295,319]" pageId="35" pageNumber="36">CEUA</collectionCode>
12407 (
<specimenCount id="9D11FD675212D40AC4992114FB53FE96" box="[1178,1256,295,319]" pageId="35" pageNumber="36" type="male">1 male</specimenCount>
, 
<collectionCode id="ED06AE2B5212D40AC4F52114FAFCFE96" box="[1270,1351,295,319]" pageId="35" pageNumber="36">CEUA</collectionCode>
); idem\ Bosque noche Manual\ 
<collectionCode id="ED06AE2B5212D40AC1B7217FFDBBFECD" box="[436,512,332,356]" pageId="35" pageNumber="36">CEUA</collectionCode>
(
<specimenCount id="9D11FD675212D40AC219217FFDD6FECD" box="[538,621,332,356]" pageId="35" pageNumber="36" type="male">1 male</specimenCount>
, 
<collectionCode id="ED06AE2B5212D40AC27C217FFD6BFECD" box="[639,720,332,356]" pageId="35" pageNumber="36">CEUA</collectionCode>
); CO [
<collectingCountry id="F300767E5212D40AC325217FFC25FECD" box="[806,926,332,356]" name="Colombia" pageId="35" pageNumber="36">Colombia</collectingCountry>
]. Caldas. Manizales. Reserva Rio\ Blanco. 5˚ 
<date id="FFA9102E5212D40AC0AA215CFEB8FE2E" box="[169,259,367,391]" pageId="35" pageNumber="36" value="58-07-26">7.26.58</date>
”N 75˚43.7’9.58”W\ 2592 msnm.T VSR - pez [Van Someren-Rydon trap with fish] Bosque\ 
<date id="FFA9102E5212D40AC542215CFF74FE05" pageId="35" pageNumber="36" value="2005-04-03">Abril 3/ 2005</date>
G. Entomologia\ 
<collectionCode id="ED06AE2B5212D40AC19421A7FE58FE05" box="[407,483,404,428]" pageId="35" pageNumber="36">CEUA</collectionCode>
12392 (
<specimenCount id="9D11FD675212D40AC24721A7FD1CFE05" box="[580,679,404,428]" pageId="35" pageNumber="36" type="female">1 female</specimenCount>
, 
<collectionCode id="ED06AE2B5212D40AC2B021A7FCBFFE05" box="[691,772,404,428]" pageId="35" pageNumber="36">CEUA</collectionCode>
); “idem \ 
<collectionCode id="ED06AE2B5212D40AC37721A7FC7BFE05" box="[884,960,404,428]" pageId="35" pageNumber="36">CEUA</collectionCode>
(
<specimenCount id="9D11FD675212D40AC3D221A7FBF9FE05" box="[977,1090,404,428]" pageId="35" pageNumber="36" type="female">2 females</specimenCount>
, 
<collectionCode id="ED06AE2B5212D40AC44D21A7FB24FE05" box="[1102,1183,404,428]" pageId="35" pageNumber="36">CEUA</collectionCode>
); “idem \ 
<collectionCode id="ED06AE2B5212D40AC51321A7FAE7FE05" box="[1296,1372,404,428]" pageId="35" pageNumber="36">CEUA</collectionCode>
7497 (
<specimenCount id="9D11FD675212D40AC0A12184FEBFFE66" box="[162,260,439,463]" pageId="35" pageNumber="36" type="female">1 female</specimenCount>
, 
<collectionCode id="ED06AE2B5212D40AC10C2184FEDBFE66" box="[271,352,439,463]" pageId="35" pageNumber="36">CEUA</collectionCode>
); “idem \ 
<collectionCode id="ED06AE2B5212D40AC1CD2184FDA1FE66" box="[462,538,439,463]" pageId="35" pageNumber="36">CEUA</collectionCode>
7495(
<specimenCount id="9D11FD675212D40AC2642184FD7DFE66" box="[615,710,439,463]" pageId="35" pageNumber="36" type="female">1 female</specimenCount>
, 
<collectionCode id="ED06AE2B5212D40AC2D22184FCA4FE66" box="[721,799,439,463]" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="35" pageNumber="36">DZUP</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C30D65655212D40AC0C421E8FBCFFE5D" box="[199,1140,475,500]" pageId="35" pageNumber="36" type="distribution">
<paragraph id="8BA836EE5212D40AC0C421E8FBCFFE5D" blockId="35.[151,1437,151,500]" box="[199,1140,475,500]" pageId="35" pageNumber="36">
<emphasis id="B963EAFC5212D40AC0C421E8FEDBFE5D" bold="true" box="[199,352,475,500]" pageId="35" pageNumber="36">Distribution.</emphasis>
<collectingCountry id="F300767E5212D40AC16421EFFE61FE5D" box="[359,474,476,500]" name="Colombia" pageId="35" pageNumber="36">Colombia</collectingCountry>
: department of Caldas (Andean region, Central Cordillera).
</paragraph>
</subSubSection>
</treatment>
</document>