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<mods:title id="82BEAE35750DE1D026E7633A235B6072">Preliminary re-examination of genus-level taxonomy of the pollen beetle subfamily Meligethinae (Coleoptera: Nitidulidae)</mods:title>
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<mods:namePart id="1D2CF5D903203E2879CE0766A77C0F09">Audisio, Paolo</mods:namePart>
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<mods:namePart id="187D96A1709D1F539DDC5079AFAC0006">Cline, Andrew Richard</mods:namePart>
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<mods:namePart id="07DA03B2EC7E3957B9F6D6FBE541A2C0">Biase, Alessio De</mods:namePart>
<mods:affiliation id="1F8C02972E8526A7102833F72DA3F027">Dipartimento di Biologia Animale e dell’Uomo, Sapienza Rome University, viale Dell’Università, 32, I- 00185 Rome, Italy; e-mail: alessio. debiase @ uniroma 1. it</mods:affiliation>
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<mods:namePart id="EE1FB5B90CFABF127D399313A2140F54">Antonini, Gloria</mods:namePart>
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<mods:namePart id="2C41ABF1558AF1FC54CF52F4257E7F98">Mancini, Emiliano</mods:namePart>
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<mods:namePart id="1477FB68483FE442A3B4560B185826DB">Trizzino, Marco</mods:namePart>
<mods:affiliation id="1DB733AFDAC104BBBDEBA06742186AF3">Dipartimento di Biologia Animale e dell’Uomo, Sapienza Rome University, via A. Borelli, 50, I- 00161 Rome, Italy; e-mail: paolo. audisio @ uniroma 1. it</mods:affiliation>
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<mods:namePart id="A93CFAFB299DF6A7536CC845666DDEE3">Costantini, Lorenzo</mods:namePart>
<mods:affiliation id="DBC23241288815CC97E84D53E6FB1260">Museo Nazionale d’Arte Orientale, Servizio di Bioarcheologia e Microscopia, via Merulana 248, I- 00185 Rome, Italy; e-mail: l. costantin @ mclink. it</mods:affiliation>
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<mods:namePart id="ABE92AE30EFC97102A901ED0436762B7">Strika, Sirio</mods:namePart>
<mods:affiliation id="DC2FD9DB2C7618994DAD315B68BD562C">Museo Nazionale d’Arte Orientale, Servizio di Bioarcheologia e Microscopia, via Merulana 248, I- 00185 Rome, Italy; e-mail: l. costantin @ mclink. it</mods:affiliation>
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<mods:affiliation id="C82E5ED17635800A17F5FD0B9F2A43AD">Dipartimento di Biologia Animale e dell’Uomo, Sapienza Rome University, via A. Borelli, 50, I- 00161 Rome, Italy; e-mail: paolo. audisio @ uniroma 1. it</mods:affiliation>
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<mods:namePart id="73FDD5D778734DC8DC7DA8AF3D01592B">Cerretti, Pierfilippo</mods:namePart>
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39. 
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<emphasis id="B963EAC8F6E6FF08BAB1FB70FDF7FB27" bold="true" box="[442,578,1156,1180]" italics="true" pageId="140" pageNumber="481">Cornutopria</emphasis>
S. 
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(
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)
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<emphasis id="B963EAC8F6E6FF08BB9CFB32FEB7FB63" box="[151,258,1222,1240]" italics="true" pageId="140" pageNumber="481">Cornutopria</emphasis>
S. 
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, 305.
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species.
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<emphasis id="B963EAC8F6E6FF08BA3EFB06FD88FAB2" box="[309,573,1266,1289]" italics="true" pageId="140" pageNumber="481">Cornutopria basilewskyi</emphasis>
S. 
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(by original designation).
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<emphasis id="B963EAC8F6E6FF08BB9CFACBFED7FAEC" bold="true" box="[151,354,1343,1367]" pageId="140" pageNumber="481">Generic diagnosis.</emphasis>
The single known species (
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length; 1.0– 
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width) exhibits the combination of characters listed by ENDRÖDY- YOUNGA (1978) (
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; all drawings from ENDRÖDY- YOUNGA 1978).
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<emphasis id="B963EAC8F6E6FF08BB9CFA54FEA2FA03" bold="true" box="[151,279,1440,1464]" pageId="140" pageNumber="481">Etymology.</emphasis>
The generic name was derived from the combination of the Latin name ‘ 
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<emphasis id="B963EAC8F6E6FF08BF08FA55FBEAFA03" box="[1027,1119,1441,1464]" italics="true" pageId="140" pageNumber="481">cornutus</emphasis>
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’ (= bearing horns), and 
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<emphasis id="B963EAC8F6E6FF08BA8FFA35FE06FA63" box="[388,435,1473,1496]" italics="true" pageId="140" pageNumber="481">Pria</emphasis>
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, a widespread genus of 
<taxonomicName id="4C174D59F6E6FF08B9A0FA4BFC96FA6C" authorityName="Latreille" authorityYear="1802" box="[683,803,1471,1495]" class="Insecta" family="Nitidulidae" kingdom="Animalia" order="Coleoptera" pageId="140" pageNumber="481" phylum="Arthropoda" rank="family">Nitidulidae</taxonomicName>
<taxonomicName id="4C174D59F6E6FF08B823FA4BFC02FA6C" authorityName="C. G. Thomson" authorityYear="1859" box="[808,951,1471,1495]" class="Insecta" family="Nitidulidae" kingdom="Animalia" order="Coleoptera" pageId="140" pageNumber="481" phylum="Arthropoda" rank="subFamily" subFamily="Meligethinae">Meligethinae</taxonomicName>
, which is indicative of the long antennae of the single species attributed to this taxon, and the phylogenetic relationships with 
<taxonomicName id="4C174D59F6E6FF08BA57F9F5FE3EF9A3" authorityName="Stephens" authorityYear="1830" box="[348,395,1537,1560]" class="Insecta" family="Nitidulidae" genus="Pria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="140" pageNumber="481" phylum="Arthropoda" rank="genus">
<emphasis id="B963EAC8F6E6FF08BA57F9F5FE3EF9A3" box="[348,395,1537,1560]" italics="true" pageId="140" pageNumber="481">Pria</emphasis>
</taxonomicName>
. Gender feminine.
</paragraph>
</subSubSection>
<caption id="DF686652F6E7FF09BB7AFBA8FD99FB7B" ID-DOI="http://doi.org/10.5281/zenodo.5363214" ID-Zenodo-Dep="5363214" httpUri="https://zenodo.org/record/5363214/files/figure.png" pageId="141" pageNumber="482" startId="141.[113,144,1116,1136]" targetBox="[264,947,223,1093]" targetPageId="141">
<paragraph id="8BA836DAF6E7FF09BB7AFBA8FD99FB7B" blockId="141.[113,1089,1116,1217]" pageId="141" pageNumber="482">
<emphasis id="B963EAC8F6E7FF09BB7AFBA8FF00FBCB" bold="true" box="[113,181,1116,1136]" pageId="141" pageNumber="482">Fig. 39.</emphasis>
<taxonomicName id="4C174D59F6E7FF09BBB7FBAAFDA5FBCB" authority="S. Endrody-Younga, 1978" authorityName="S. Endrody-Younga" authorityYear="1978" box="[188,528,1117,1136]" class="Insecta" family="Nitidulidae" genus="Cornutopria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="141" pageNumber="482" phylum="Arthropoda" rank="genus">
<emphasis id="B963EAC8F6E7FF09BBB7FBAAFE92FBCB" box="[188,295,1118,1136]" italics="true" pageId="141" pageNumber="482">Cornutopria</emphasis>
S. 
<bibRefCitation id="EF864B2BF6E7FF09BA4FFBA9FDA5FBCB" author="ENDRODY-YOUNGA S." box="[324,528,1117,1136]" pageId="141" pageNumber="482" pagination="295 - 316" refId="ref85938" refString="ENDRODY-YOUNGA S. 1978: Systematic revision and phylogeny of some Meligethinae genera from the Ethiopian Region (Coleoptera: Nitidulidae). Entomologica Germanica 4: 295 - 316." type="journal article" year="1978">Endrödy-Younga, 1978</bibRefCitation>
</taxonomicName>
: 
<emphasis id="B963EAC8F6E7FF09B912FBA8FDF2FBCB" bold="true" box="[537,583,1116,1136]" pageId="141" pageNumber="482">a–f –</emphasis>
<taxonomicName id="4C174D59F6E7FF09B946FBAAFC04FBCB" authority="S. Endrody-Younga, 1978" authorityName="S. Endrody-Younga" authorityYear="1978" box="[589,945,1117,1136]" class="Insecta" family="Nitidulidae" genus="Cornutopria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="141" pageNumber="482" phylum="Arthropoda" rank="species" species="basilewskyi">
<emphasis id="B963EAC8F6E7FF09B946FBAAFD7DFBCB" box="[589,712,1118,1136]" italics="true" pageId="141" pageNumber="482">C. basilewskyi</emphasis>
S. 
<bibRefCitation id="EF864B2BF6E7FF09B9ECFBA9FC04FBCB" author="ENDRODY-YOUNGA S." box="[743,945,1117,1136]" pageId="141" pageNumber="482" pagination="295 - 316" refId="ref85938" refString="ENDRODY-YOUNGA S. 1978: Systematic revision and phylogeny of some Meligethinae genera from the Ethiopian Region (Coleoptera: Nitidulidae). Entomologica Germanica 4: 295 - 316." type="journal article" year="1978">Endrödy-Younga, 1978</bibRefCitation>
</taxonomicName>
. 
<emphasis id="B963EAC8F6E7FF09B8B0FBA9FC73FBCB" bold="true" box="[955,966,1117,1136]" pageId="141" pageNumber="482">a</emphasis>
– dorsal view of male head and first three antennomeres; 
<emphasis id="B963EAC8F6E7FF09BAEBFB8CFE59FB30" bold="true" box="[480,492,1144,1163]" pageId="141" pageNumber="482">b</emphasis>
– male antennal club and antennomeres 4–11; 
<emphasis id="B963EAC8F6E7FF09B877FB8CFC30FB30" bold="true" box="[892,901,1144,1163]" pageId="141" pageNumber="482">c</emphasis>
– mentum; 
<emphasis id="B963EAC8F6E7FF09B8E0FB8CFC42FB30" bold="true" box="[1003,1015,1144,1163]" pageId="141" pageNumber="482">d</emphasis>
– dorsal view of aedeagus; 
<emphasis id="B963EAC8F6E7FF09BA18FB66FE9BFB1D" bold="true" box="[275,302,1170,1190]" pageId="141" pageNumber="482">e–f</emphasis>
– dorsal and lateral view of male pygidium. All drawings modified from ENDRÖDY- YOUNGA (1978). Refer to ENDRÖDY- YOUNGA (1978) for scale.
</paragraph>
</caption>
<subSubSection id="C30D6551F6E7FF09BB7AFB1AFD05F9F5" pageId="141" pageNumber="482" type="biology_ecology">
<paragraph id="8BA836DAF6E7FF09BB7AFB1AFECAFA3D" blockId="141.[113,1090,1262,1574]" pageId="141" pageNumber="482">
<emphasis id="B963EAC8F6E7FF09BB7AFB1AFF78FABD" bold="true" box="[113,205,1262,1286]" pageId="141" pageNumber="482">Biology.</emphasis>
The biology of the single known representative of 
<taxonomicName id="4C174D59F6E7FF09B9D4FB1BFCD1FABD" authorityName="S. Endrody-Younga" authorityYear="1978" box="[735,868,1263,1286]" class="Insecta" family="Nitidulidae" genus="Cornutopria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="141" pageNumber="482" phylum="Arthropoda" rank="genus">
<emphasis id="B963EAC8F6E7FF09B9D4FB1BFCD1FABD" box="[735,868,1263,1286]" italics="true" pageId="141" pageNumber="482">Cornutopria</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C174D59F6E7FF09B87BFB1BFBBDFABD" authorityName="S. Endrody-Younga" authorityYear="1978" box="[880,1032,1263,1286]" class="Insecta" family="Nitidulidae" genus="Cornutopria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="141" pageNumber="482" phylum="Arthropoda" rank="species" species="basilewskyi">
<emphasis id="B963EAC8F6E7FF09B87BFB1BFBBDFABD" box="[880,1032,1263,1286]" italics="true" pageId="141" pageNumber="482">C. basilewskyi</emphasis>
</taxonomicName>
from 
<collectingCountry id="F300764AF6E7FF09BB7AFAFAFF19FA9D" box="[113,172,1294,1318]" name="Democratic Republic of the Congo" pageId="141" pageNumber="482">Zaire</collectingCountry>
, is unknown (ENDRÖDY- YOUNGA 1978). A probable association with male inflorescences of palms (
<taxonomicName id="4C174D59F6E7FF09BBD4FADAFEE7FAFD" box="[223,338,1326,1350]" class="Liliopsida" family="Arecaceae" kingdom="Plantae" order="Arecales" pageId="141" pageNumber="482" phylum="Tracheophyta" rank="family">Arecaceae</taxonomicName>
) may be inferred, as it is analogously exhibited in the related genera 
<taxonomicName id="4C174D59F6E7FF09BB7AFABBFEB3FADD" authorityName="S. Endrody-Younga" authorityYear="1978" box="[113,262,1359,1382]" class="Insecta" family="Nitidulidae" genus="Microporodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="141" pageNumber="482" phylum="Arthropoda" rank="genus">
<emphasis id="B963EAC8F6E7FF09BB7AFABBFEB3FADD" box="[113,262,1359,1382]" italics="true" pageId="141" pageNumber="482">Microporodes</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C174D59F6E7FF09BA36FABBFE1AFADD" authorityName="S. Endrody-Younga" authorityYear="1978" box="[317,431,1359,1382]" class="Insecta" family="Nitidulidae" genus="Palmopria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="141" pageNumber="482" phylum="Arthropoda" rank="genus">
<emphasis id="B963EAC8F6E7FF09BA36FABBFE1AFADD" box="[317,431,1359,1382]" italics="true" pageId="141" pageNumber="482">Palmopria</emphasis>
</taxonomicName>
, however this assumption is speculative and requires further fieldwork to substantiate.
</paragraph>
<paragraph id="8BA836DAF6E7FF09BB7AFA7AFD86FA5D" blockId="141.[113,1090,1262,1574]" pageId="141" pageNumber="482">
<emphasis id="B963EAC8F6E7FF09BB7AFA7AFED7FA1D" bold="true" box="[113,354,1422,1446]" pageId="141" pageNumber="482">Phylogenetic position.</emphasis>
Available morphological data provides possible evidence of a relatively close relationship with members of the [(
<taxonomicName id="4C174D59F6E7FF09B92AFA5BFD26FA7D" authorityName="S. Endrody-Younga" authorityYear="1978" box="[545,659,1455,1478]" class="Insecta" family="Nitidulidae" genus="Palmopria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="141" pageNumber="482" phylum="Arthropoda" rank="genus">
<emphasis id="B963EAC8F6E7FF09B92AFA5BFD26FA7D" box="[545,659,1455,1478]" italics="true" pageId="141" pageNumber="482">Palmopria</emphasis>
</taxonomicName>
+ 
<taxonomicName id="4C174D59F6E7FF09B9A6FA5BFCF7FA7D" authorityName="S. Endrody-Younga" authorityYear="1978" box="[685,834,1455,1478]" class="Insecta" family="Nitidulidae" genus="Microporodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="141" pageNumber="482" phylum="Arthropoda" rank="genus">
<emphasis id="B963EAC8F6E7FF09B9A6FA5BFCF7FA7D" box="[685,834,1455,1478]" italics="true" pageId="141" pageNumber="482">Microporodes</emphasis>
</taxonomicName>
) + 
<taxonomicName id="4C174D59F6E7FF09B86EFA5BFC46FA7D" authorityName="S. Endrody-Younga" authorityYear="1978" box="[869,1011,1455,1478]" class="Insecta" family="Nitidulidae" genus="Lechanteuria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="141" pageNumber="482" phylum="Arthropoda" rank="genus">
<emphasis id="B963EAC8F6E7FF09B86EFA5BFC46FA7D" box="[869,1011,1455,1478]" italics="true" pageId="141" pageNumber="482">Lechanteuria</emphasis>
</taxonomicName>
] clade. No molecular data are currently available.
</paragraph>
<paragraph id="8BA836DAF6E7FF09BB7AFA1AFF08F99D" blockId="141.[113,1090,1262,1574]" pageId="141" pageNumber="482">
<emphasis id="B963EAC8F6E7FF09BB7AFA1AFD86F9BD" bold="true" box="[113,563,1518,1542]" pageId="141" pageNumber="482">Taxonomy and geographic distribution.</emphasis>
This taxon includes a single species from central Africa.
</paragraph>
<paragraph id="8BA836DAF6E7FF09BB7AF9C8FD05F9F5" blockId="141.[113,688,1595,1615]" box="[113,688,1595,1615]" pageId="141" pageNumber="482">
<taxonomicName id="4C174D59F6E7FF09BB7AF9C8FD92F9F4" authority="S. Endrody-Younga, 1978" authorityName="S. Endrody-Younga" authorityYear="1978" box="[113,551,1595,1615]" class="Insecta" family="Nitidulidae" genus="Cornutopria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="141" pageNumber="482" phylum="Arthropoda" rank="species" species="basilewskyi">
<emphasis id="B963EAC8F6E7FF09BB7AF9C8FEF6F9F5" box="[113,323,1596,1614]" italics="true" pageId="141" pageNumber="482">Cornutopria basilewskyi</emphasis>
S. 
<bibRefCitation id="EF864B2BF6E7FF09BA6BF9CFFD92F9F4" author="ENDRODY-YOUNGA S." box="[352,551,1595,1615]" pageId="141" pageNumber="482" pagination="295 - 316" refId="ref85938" refString="ENDRODY-YOUNGA S. 1978: Systematic revision and phylogeny of some Meligethinae genera from the Ethiopian Region (Coleoptera: Nitidulidae). Entomologica Germanica 4: 295 - 316." type="journal article" year="1978">Endrödy-Younga, 1978</bibRefCitation>
</taxonomicName>
<collectingCountry id="F300764AF6E7FF09B988F9CFFD05F9F5" box="[643,688,1595,1614]" name="Democratic Republic of the Congo" pageId="141" pageNumber="482">Zaire</collectingCountry>
</paragraph>
</subSubSection>
</treatment>
</document>