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<document ID-DOI="http://dx.doi.org/10.3897/subtbiol.37.61573" ID-GBIF-Dataset="d4970e14-1aa4-4857-94d2-c8fa31d2b288" ID-GBIF-Taxon="177423716" ID-Pensoft-Pub="1314-2615-37-89" ID-Pensoft-UUID="008ACFFD461C511893E5F656C8C555B9" ID-ZooBank="A6AD0D9D2EAB4AC3ADB8696EBA177B41" LSID="urn:lsid:zoobank.org:act:77EA36ED-B25F-4E95-9DC5-20C8B3513B40" ModsDocID="1314-2615-37-89" checkinTime="1613660849660" checkinUser="pensoft" docAuthor="Falniowski 1, Andrzej, Grego 2, Jozef, Rysiewska 1, Aleksandra, Osikowski 3, Artur &amp; Hofman 4, Sebastian" docDate="2021" docId="171E927DEE35581B84CAACFE5E9E13AC" docLanguage="en" docName="SubterBiol 37: 89-104" docOrigin="Subterranean Biology 37" docPubDate="2021-02-16" docSource="http://dx.doi.org/10.3897/subtbiol.37.61573" docTitle="Horatia ozimeci Grego &amp; Falniowski, sp. nov." docType="treatment" docVersion="3" id="008ACFFD461C511893E5F656C8C555B9" lastPageNumber="89" masterDocId="008ACFFD461C511893E5F656C8C555B9" masterDocTitle="Two new stygobiotic species of Horatia Bourguignat, 1887 (Hydrobiidae) from Croatia" masterLastPageNumber="104" masterPageNumber="89" pageId="0" pageNumber="89" updateTime="1643437583660" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Two new stygobiotic species of Horatia Bourguignat, 1887 (Hydrobiidae) from Croatia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Falniowski 1, Andrzej</mods:namePart>
<mods:affiliation>Department of Malacology, Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Grego 2, Jozef</mods:namePart>
<mods:affiliation>Department of Malacology, Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rysiewska 1, Aleksandra</mods:namePart>
<mods:affiliation>Department of Malacology, Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Osikowski 3, Artur</mods:namePart>
<mods:affiliation>Department of Malacology, Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hofman 4, Sebastian</mods:namePart>
<mods:affiliation>Department of Malacology, Institute of Zoology and Biomedical Research, Jagiellonian University, Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Subterranean Biology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-02-16</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>37</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>89</mods:start>
<mods:end>104</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/subtbiol.37.61573</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/subtbiol.37.61573</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2615-37-89</mods:identifier>
<mods:identifier type="ZooBank">A6AD0D9D2EAB4AC3ADB8696EBA177B41</mods:identifier>
<mods:identifier type="Pensoft-UUID">008ACFFD461C511893E5F656C8C555B9</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="177423716" LSID="urn:lsid:zoobank.org:act:77EA36ED-B25F-4E95-9DC5-20C8B3513B40" httpUri="http://treatment.plazi.org/id/171E927DEE35581B84CAACFE5E9E13AC" lastPageNumber="89" pageId="0" pageNumber="89">
<subSubSection pageId="0" pageNumber="89" type="nomenclature">
<paragraph pageId="0" pageNumber="89">
<taxonomicName LSID="http://zoobank.org/77EA36ED-B25F-4E95-9DC5-20C8B3513B40" authority="Grego &amp; Falniowski, sp. nov." authorityName="Grego &amp; Falniowski, sp. nov." class="Gastropoda" family="Hydrobiidae" genus="Horatia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatia ozimeci" order="Littorinimorpha" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="ozimeci">Horatia ozimeci Grego &amp; Falniowski, sp. nov.</taxonomicName>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Shell variability of two Horatia species A-E H. ozimeci A sequenced specimen (2 H 76) B holotype OZRM 11603 C-E paratypes E NHMW 113607, (Photo NHMW by Ivo Gallmetzer) F-H H. stygorumina F holotype OZRM 11600 G-H paratypes NHMW 113608, (Photo NHMW by Gallmetzer) I Cf. Horatia knorri Schuett, 1961, spring of Ombla in Komolac, holotype SMF 164247 (Photo SMF by Sigrid Hof) J-P sequenced specimens of H. klecakiana J Vrelo Kosinac 2 K H. klecakiana, Gala spring L H. klecakiana, Ruda Begusa M, N, P H. klecakiana, Vrelo Kosinac 1 O H. klecakiana, Grabske Mlinice. The shell measurements are also shown: a shell height, b body whorl breadth, c aperture height, d spire height, e aperture breadth, α apex angle, β angle between body whorl suture and horizontal surface. Scale bars: 1 mm." figureDoi="10.3897/subtbiol.37.61573.figure4" httpUri="https://binary.pensoft.net/fig/509943" pageId="0" pageNumber="89">Figs 4A-E</figureCitation>
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. SEM images of H. ozimeci (A) juvenile specimen from the type locality (SBMNH 635080) and H. stygorumina (B) juvenile shell from cave Mali Rumin (SBMNH 635079). SEM SBMNH by Vanessa Delnavaz." figureDoi="10.3897/subtbiol.37.61573.figure5" httpUri="https://binary.pensoft.net/fig/509944" pageId="0" pageNumber="89">, 5A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="89" type="type locality">
<paragraph pageId="0" pageNumber="89">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="89">
Spring Izvor Ruda -
<normalizedToken originalValue="Beguša">Begusa</normalizedToken>
, Ruda, sand at the stream bottom below the spring lake, 13 km ESE of Sinj, Split district, Croatia;
<geoCoordinate degrees="43" direction="north" minutes="40" orientation="latitude" precision="1" seconds="06.6" value="43.668503">43°40'06.6&quot;N</geoCoordinate>
,
<geoCoordinate degrees="16" direction="east" minutes="47" orientation="longitude" precision="1" seconds="45.6" value="16.796">16°47'45.6&quot;E</geoCoordinate>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="89" type="holotype">
<paragraph pageId="0" pageNumber="89">Holotype.</paragraph>
<paragraph pageId="0" pageNumber="89">
Dry shell with operculum (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Shell variability of two Horatia species A-E H. ozimeci A sequenced specimen (2 H 76) B holotype OZRM 11603 C-E paratypes E NHMW 113607, (Photo NHMW by Ivo Gallmetzer) F-H H. stygorumina F holotype OZRM 11600 G-H paratypes NHMW 113608, (Photo NHMW by Gallmetzer) I Cf. Horatia knorri Schuett, 1961, spring of Ombla in Komolac, holotype SMF 164247 (Photo SMF by Sigrid Hof) J-P sequenced specimens of H. klecakiana J Vrelo Kosinac 2 K H. klecakiana, Gala spring L H. klecakiana, Ruda Begusa M, N, P H. klecakiana, Vrelo Kosinac 1 O H. klecakiana, Grabske Mlinice. The shell measurements are also shown: a shell height, b body whorl breadth, c aperture height, d spire height, e aperture breadth, α apex angle, β angle between body whorl suture and horizontal surface. Scale bars: 1 mm." figureDoi="10.3897/subtbiol.37.61573.figure4" httpUri="https://binary.pensoft.net/fig/509943" pageId="0" pageNumber="89">4B</figureCitation>
), J. Grego, A. Falniowski, M.
<normalizedToken originalValue="Olšavský">Olsavsky</normalizedToken>
and J.
<normalizedToken originalValue="Olšavská">Olsavska</normalizedToken>
leg., August 5-th 2020, OZRM 11600.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="89" type="paratypes">
<paragraph pageId="0" pageNumber="89">Paratypes.</paragraph>
<paragraph pageId="0" pageNumber="89">
The single live collected paratype (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Shell variability of two Horatia species A-E H. ozimeci A sequenced specimen (2 H 76) B holotype OZRM 11603 C-E paratypes E NHMW 113607, (Photo NHMW by Ivo Gallmetzer) F-H H. stygorumina F holotype OZRM 11600 G-H paratypes NHMW 113608, (Photo NHMW by Gallmetzer) I Cf. Horatia knorri Schuett, 1961, spring of Ombla in Komolac, holotype SMF 164247 (Photo SMF by Sigrid Hof) J-P sequenced specimens of H. klecakiana J Vrelo Kosinac 2 K H. klecakiana, Gala spring L H. klecakiana, Ruda Begusa M, N, P H. klecakiana, Vrelo Kosinac 1 O H. klecakiana, Grabske Mlinice. The shell measurements are also shown: a shell height, b body whorl breadth, c aperture height, d spire height, e aperture breadth, α apex angle, β angle between body whorl suture and horizontal surface. Scale bars: 1 mm." figureDoi="10.3897/subtbiol.37.61573.figure4" httpUri="https://binary.pensoft.net/fig/509943" pageId="0" pageNumber="89">4A</figureCitation>
) has been destroyed for DNA extraction; GenBank number: MW448545.
</paragraph>
<paragraph pageId="0" pageNumber="89">
From type locality: J. Grego, A. Falniowski, M.
<normalizedToken originalValue="Olšavský">Olsavsky</normalizedToken>
and J.
<normalizedToken originalValue="Olšavská">Olsavska</normalizedToken>
leg August 5-th 2020, OZRM 11601/1 specimen; NHMW 113607/1 specimen, JG 1542/28 specimens (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Shell variability of two Horatia species A-E H. ozimeci A sequenced specimen (2 H 76) B holotype OZRM 11603 C-E paratypes E NHMW 113607, (Photo NHMW by Ivo Gallmetzer) F-H H. stygorumina F holotype OZRM 11600 G-H paratypes NHMW 113608, (Photo NHMW by Gallmetzer) I Cf. Horatia knorri Schuett, 1961, spring of Ombla in Komolac, holotype SMF 164247 (Photo SMF by Sigrid Hof) J-P sequenced specimens of H. klecakiana J Vrelo Kosinac 2 K H. klecakiana, Gala spring L H. klecakiana, Ruda Begusa M, N, P H. klecakiana, Vrelo Kosinac 1 O H. klecakiana, Grabske Mlinice. The shell measurements are also shown: a shell height, b body whorl breadth, c aperture height, d spire height, e aperture breadth, α apex angle, β angle between body whorl suture and horizontal surface. Scale bars: 1 mm." figureDoi="10.3897/subtbiol.37.61573.figure4" httpUri="https://binary.pensoft.net/fig/509943" pageId="0" pageNumber="89">4C-E</figureCitation>
); Type locality, J. Grego, G. Jakab and B.
<normalizedToken originalValue="Šmída">Smida</normalizedToken>
leg. March 17-th 2017, JG F0724/24 specimens.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="89" type="diagnosis">
<paragraph pageId="0" pageNumber="89">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="89">
Shell minute, valvatoid, distinguished from the other species of
<taxonomicName authorityName="Bourguignat" authorityYear="1887" class="Gastropoda" family="Hydrobiidae" genus="Horatia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatia" order="Littorinimorpha" pageId="0" pageNumber="89" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="89">Horatia</emphasis>
</taxonomicName>
by its body whorl separated at its terminal sector from the penultimate one (scalarity at this part), the circular and complete peristome and extremely wide umbilicus showing earlier whorls inside.
</paragraph>
<caption doi="10.3897/subtbiol.37.61573.figure4" httpUri="https://binary.pensoft.net/fig/509943" pageId="0" pageNumber="89" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="89">
<emphasis bold="true" pageId="0" pageNumber="89">Figure 4.</emphasis>
Shell variability of two
<taxonomicName authorityName="Bourguignat" authorityYear="1887" class="Gastropoda" family="Hydrobiidae" genus="Horatia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatia" order="Littorinimorpha" pageId="0" pageNumber="89" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="89">Horatia</emphasis>
</taxonomicName>
species
<emphasis bold="true" pageId="0" pageNumber="89">A-E</emphasis>
<taxonomicName genus="H." lsidName="H. ozimeci" pageId="0" pageNumber="89" rank="species" species="ozimeci">
<emphasis italics="true" pageId="0" pageNumber="89">H. ozimeci</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="89">A</emphasis>
sequenced specimen (2H76)
<emphasis bold="true" pageId="0" pageNumber="89">B</emphasis>
holotype OZRM 11603
<emphasis bold="true" pageId="0" pageNumber="89">C-E</emphasis>
paratypes
<emphasis bold="true" pageId="0" pageNumber="89">E</emphasis>
NHMW 113607, (Photo NHMW by Ivo Gallmetzer)
<emphasis bold="true" pageId="0" pageNumber="89">F-H</emphasis>
<taxonomicName genus="H." lsidName="H. stygorumina" pageId="0" pageNumber="89" rank="species" species="stygorumina">
<emphasis italics="true" pageId="0" pageNumber="89">H. stygorumina</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="89">F</emphasis>
holotype OZRM 11600
<emphasis bold="true" pageId="0" pageNumber="89">G-H</emphasis>
paratypes NHMW 113608, (Photo NHMW by Gallmetzer)
<emphasis bold="true" pageId="0" pageNumber="89">I</emphasis>
Cf.
<taxonomicName authorityName="Schutt" authorityYear="1961" class="Gastropoda" family="Hydrobiidae" genus="Horatia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatia knorri" order="Littorinimorpha" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="knorri">
<emphasis italics="true" pageId="0" pageNumber="89">Horatia knorri</emphasis>
</taxonomicName>
<normalizedToken originalValue="Schütt">Schuett</normalizedToken>
, 1961, spring of Ombla in Komolac, holotype SMF 164247 (Photo SMF by Sigrid Hof)
<emphasis bold="true" pageId="0" pageNumber="89">J-P</emphasis>
sequenced specimens of
<taxonomicName genus="H." lsidName="H. klecakiana" pageId="0" pageNumber="89" rank="species" species="klecakiana">
<emphasis italics="true" pageId="0" pageNumber="89">H. klecakiana</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="89">J</emphasis>
Vrelo Kosinac 2
<emphasis bold="true" pageId="0" pageNumber="89">K</emphasis>
<taxonomicName genus="H." lsidName="H. klecakiana" pageId="0" pageNumber="89" rank="species" species="klecakiana">
<emphasis italics="true" pageId="0" pageNumber="89">H. klecakiana</emphasis>
</taxonomicName>
, Gala spring
<emphasis bold="true" pageId="0" pageNumber="89">L</emphasis>
<taxonomicName genus="H." lsidName="H. klecakiana" pageId="0" pageNumber="89" rank="species" species="klecakiana">
<emphasis italics="true" pageId="0" pageNumber="89">H. klecakiana</emphasis>
</taxonomicName>
, Ruda
<normalizedToken originalValue="Beguša">Begusa</normalizedToken>
<emphasis bold="true" pageId="0" pageNumber="89">M, N, P</emphasis>
<taxonomicName genus="H." lsidName="H. klecakiana" pageId="0" pageNumber="89" rank="species" species="klecakiana">
<emphasis italics="true" pageId="0" pageNumber="89">H. klecakiana</emphasis>
</taxonomicName>
, Vrelo Kosinac 1
<emphasis bold="true" pageId="0" pageNumber="89">O</emphasis>
<taxonomicName genus="H." lsidName="H. klecakiana" pageId="0" pageNumber="89" rank="species" species="klecakiana">
<emphasis italics="true" pageId="0" pageNumber="89">H. klecakiana</emphasis>
</taxonomicName>
, Grabske Mlinice. The shell measurements are also shown: a shell height, b body whorl breadth, c aperture height, d spire height, e aperture breadth,
<normalizedToken originalValue="α">α</normalizedToken>
apex angle,
<normalizedToken originalValue="β">β</normalizedToken>
angle between body whorl suture and horizontal surface. Scale bars: 1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="89" type="description">
<paragraph pageId="0" pageNumber="89">Description.</paragraph>
<paragraph pageId="0" pageNumber="89">
<emphasis bold="true" italics="true" pageId="0" pageNumber="89">Shell</emphasis>
(Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Shell variability of two Horatia species A-E H. ozimeci A sequenced specimen (2 H 76) B holotype OZRM 11603 C-E paratypes E NHMW 113607, (Photo NHMW by Ivo Gallmetzer) F-H H. stygorumina F holotype OZRM 11600 G-H paratypes NHMW 113608, (Photo NHMW by Gallmetzer) I Cf. Horatia knorri Schuett, 1961, spring of Ombla in Komolac, holotype SMF 164247 (Photo SMF by Sigrid Hof) J-P sequenced specimens of H. klecakiana J Vrelo Kosinac 2 K H. klecakiana, Gala spring L H. klecakiana, Ruda Begusa M, N, P H. klecakiana, Vrelo Kosinac 1 O H. klecakiana, Grabske Mlinice. The shell measurements are also shown: a shell height, b body whorl breadth, c aperture height, d spire height, e aperture breadth, α apex angle, β angle between body whorl suture and horizontal surface. Scale bars: 1 mm." figureDoi="10.3897/subtbiol.37.61573.figure4" httpUri="https://binary.pensoft.net/fig/509943" pageId="0" pageNumber="89">4B</figureCitation>
) 2.14 mm high and 1.62 mm broad, valvatoid, whitish, translucent, rather thin-walled, consisted of about 3
<normalizedToken originalValue="½">1/2</normalizedToken>
whorls, growing rapidly and separated by moderately deep suture, more prominent closer to the aperture: the terminal part of the body whorl completely separated from the penultimate one. Spire low and narrow, body whorl large. Aperture prosocline, circular in shape, peristome complete and separated from the columella, swollen, umbilicus extremely wide, with the earlier whorls visible inside. Shell surface smooth, with growth lines delicate but visible. Operculum reddish-brown, translucent, paucispiral.
</paragraph>
<caption doi="10.3897/subtbiol.37.61573.figure5" httpUri="https://binary.pensoft.net/fig/509944" pageId="0" pageNumber="89" start="Figure 5" startId="F5">
<paragraph pageId="0" pageNumber="89">
<emphasis bold="true" pageId="0" pageNumber="89">Figure 5.</emphasis>
SEM images of
<taxonomicName genus="H." lsidName="H. ozimeci" pageId="0" pageNumber="89" rank="species" species="ozimeci">
<emphasis italics="true" pageId="0" pageNumber="89">H. ozimeci</emphasis>
</taxonomicName>
(
<emphasis bold="true" pageId="0" pageNumber="89">A</emphasis>
) juvenile specimen from the type locality (SBMNH 635080) and
<taxonomicName genus="H." lsidName="H. stygorumina" pageId="0" pageNumber="89" rank="species" species="stygorumina">
<emphasis italics="true" pageId="0" pageNumber="89">H. stygorumina</emphasis>
</taxonomicName>
(
<emphasis bold="true" pageId="0" pageNumber="89">B</emphasis>
) juvenile shell from cave Mali Rumin (SBMNH 635079). SEM SBMNH by Vanessa Delnavaz.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="89">
<emphasis bold="true" italics="true" pageId="0" pageNumber="89">Shell variability</emphasis>
restricted (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Shell variability of two Horatia species A-E H. ozimeci A sequenced specimen (2 H 76) B holotype OZRM 11603 C-E paratypes E NHMW 113607, (Photo NHMW by Ivo Gallmetzer) F-H H. stygorumina F holotype OZRM 11600 G-H paratypes NHMW 113608, (Photo NHMW by Gallmetzer) I Cf. Horatia knorri Schuett, 1961, spring of Ombla in Komolac, holotype SMF 164247 (Photo SMF by Sigrid Hof) J-P sequenced specimens of H. klecakiana J Vrelo Kosinac 2 K H. klecakiana, Gala spring L H. klecakiana, Ruda Begusa M, N, P H. klecakiana, Vrelo Kosinac 1 O H. klecakiana, Grabske Mlinice. The shell measurements are also shown: a shell height, b body whorl breadth, c aperture height, d spire height, e aperture breadth, α apex angle, β angle between body whorl suture and horizontal surface. Scale bars: 1 mm." figureDoi="10.3897/subtbiol.37.61573.figure4" httpUri="https://binary.pensoft.net/fig/509943" pageId="0" pageNumber="89">4A, C-E</figureCitation>
), and shell less variable than in
<taxonomicName genus="H." lsidName="H. klecakiana" pageId="0" pageNumber="89" rank="species" species="klecakiana">
<emphasis italics="true" pageId="0" pageNumber="89">H. klecakiana</emphasis>
</taxonomicName>
(Figs
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Type material of taxa described by Bourguignat, 1887 A syntype of H. klecakiana MHNG - 110592. Croatie, ex Yougoslavie, Ribaric, Vallee de La Cettina (= Cetina) (= Vrijovac spring in the source area of the Cetina River), Vriovac spring in Pasko polje, near to village Cetina. The following species are junior synonyms of H. klecakiana: B syntype of H. verlikana MHNG 110600 Marais entre Verlika (= Vrlika = Verlicca) et Ribaric C syntype of H. palustris MHNG 110597 Fontaine a Ervac (= Hrvace) D syntype of H. obtusa MHNG 110596 Sorgente de La Cetina E syntype of H. letourneuxi MHNG 110593 Fontaine du moulin a Ervac (= Hrvace) F syntype of H. fontinalis MHNG 110589, Sorgente de La Cettina G syntype of H. albanica MHNG 110587, Source du moulin Cetina, a Durazzo (= Durres), not Durres in Albania, likely the misspelling of the watermill name in village Cetina H syntype of H. obliquq MHNG 110594 Fontaine du moulin a Ervac (= Hrvace). (Photo MHNG by Estee Bochud and Eike Neubert, kindly provided by Emmanuel Tardy)." figureDoi="10.3897/subtbiol.37.61573.figure1" httpUri="https://binary.pensoft.net/fig/509940" pageId="0" pageNumber="89">1</figureCitation>
,
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Shell variability of two Horatia species A-E H. ozimeci A sequenced specimen (2 H 76) B holotype OZRM 11603 C-E paratypes E NHMW 113607, (Photo NHMW by Ivo Gallmetzer) F-H H. stygorumina F holotype OZRM 11600 G-H paratypes NHMW 113608, (Photo NHMW by Gallmetzer) I Cf. Horatia knorri Schuett, 1961, spring of Ombla in Komolac, holotype SMF 164247 (Photo SMF by Sigrid Hof) J-P sequenced specimens of H. klecakiana J Vrelo Kosinac 2 K H. klecakiana, Gala spring L H. klecakiana, Ruda Begusa M, N, P H. klecakiana, Vrelo Kosinac 1 O H. klecakiana, Grabske Mlinice. The shell measurements are also shown: a shell height, b body whorl breadth, c aperture height, d spire height, e aperture breadth, α apex angle, β angle between body whorl suture and horizontal surface. Scale bars: 1 mm." figureDoi="10.3897/subtbiol.37.61573.figure4" httpUri="https://binary.pensoft.net/fig/509943" pageId="0" pageNumber="89">4J-P</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="89">
<emphasis bold="true" italics="true" pageId="0" pageNumber="89">Protoconch</emphasis>
(Fig.
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. SEM images of H. ozimeci (A) juvenile specimen from the type locality (SBMNH 635080) and H. stygorumina (B) juvenile shell from cave Mali Rumin (SBMNH 635079). SEM SBMNH by Vanessa Delnavaz." figureDoi="10.3897/subtbiol.37.61573.figure5" httpUri="https://binary.pensoft.net/fig/509944" pageId="0" pageNumber="89">5A</figureCitation>
) smooth, similar as in
<taxonomicName genus="H." lsidName="H. klecakiana" pageId="0" pageNumber="89" rank="species" species="klecakiana">
<emphasis italics="true" pageId="0" pageNumber="89">H. klecakiana</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="89">
<emphasis bold="true" italics="true" pageId="0" pageNumber="89">Measurements</emphasis>
of holotype, illustrated paratypes, and shells of
<taxonomicName genus="H." lsidName="H. klecakiana" pageId="0" pageNumber="89" rank="species" species="klecakiana">
<emphasis italics="true" pageId="0" pageNumber="89">H. klecakiana</emphasis>
</taxonomicName>
: Table
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Shell measurements of the Horatia: I-IV H. ozimeci (I sequenced specimen, II holotype, in bold; III-IV paratypes); V-VII sequenced specimens of H. klecakiana; VIII H. stygorumina, holotype, in bold. Measurements are shown in Fig. 4 J." httpUri="http://table.plazi.org/id/B70765E2784600EB05E676C7B53632D9" pageId="0" pageNumber="89" tableUuid="B70765E2784600EB05E676C7B53632D9">2</tableCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="89">
<emphasis bold="true" pageId="0" pageNumber="89">
<emphasis italics="true" pageId="0" pageNumber="89">Soft parts morphology and anatomy</emphasis>
.
</emphasis>
The body white, pigmentless, with no eyes. The anatomy unknown.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="89" type="etymology">
<paragraph pageId="0" pageNumber="89">Derivatio nominis</paragraph>
<paragraph pageId="0" pageNumber="89">
. The specific epithet
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Horatia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatia ozimeci" order="Littorinimorpha" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="ozimeci">
<emphasis italics="true" pageId="0" pageNumber="89">Horatia ozimeci</emphasis>
</taxonomicName>
refers to our friend Mr.sc. Roman Ozimec, a biospeleologist from Bast, Croatia, deeply devoted to the study and protection of the subterranean habitats in the Dinarides and Balkans.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="89" type="distribution">
<paragraph pageId="0" pageNumber="89">Distribution and habitat.</paragraph>
<paragraph pageId="0" pageNumber="89">
Known from the type locality. The type locality is a karst spring lake surrounded by three outflow caves intermittently draining the karst conduit at high water saturation. The spring draining water from sinkholes at
<normalizedToken originalValue="Buško">Busko</normalizedToken>
Jezero (Bosna and Hercegovina) and supports the river Ruda, a left tributary of Cetina River. The following
<taxonomicName class="Gastropoda" family="Hydrobiidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="0" pageNumber="89" phylum="Mollusca" rank="family">Hydrobiidae</taxonomicName>
species were detected in the habitat:
<taxonomicName authorityName="Bourguignat" authorityYear="1887" class="Gastropoda" family="Hydrobiidae" genus="Horatia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatia klecakiana" order="Littorinimorpha" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="klecakiana">
<emphasis italics="true" pageId="0" pageNumber="89">Horatia klecakiana</emphasis>
</taxonomicName>
,
<taxonomicName baseAuthorityName="Frauenfeld" baseAuthorityYear="1863" genus="Orientalina" lsidName="Orientalina curta subsp. germari" pageId="0" pageNumber="89" rank="subSpecies" species="curta" subSpecies="germari">
<emphasis italics="true" pageId="0" pageNumber="89">Orientalina curta germari</emphasis>
</taxonomicName>
(Frauenfeld, 1863),
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Montenegrospeum" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Montenegrospeum sketi" order="Littorinimorpha" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="sketi">
<emphasis italics="true" pageId="0" pageNumber="89">Montenegrospeum sketi</emphasis>
</taxonomicName>
Grego &amp;
<normalizedToken originalValue="Glöer">Gloeer</normalizedToken>
, 2018,
<taxonomicName baseAuthorityName="Kuščer" baseAuthorityYear="1933" class="Gastropoda" family="Hydrobiidae" genus="Kerkia" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Kerkia jadertina subsp. sinjana" order="Littorinimorpha" pageId="0" pageNumber="89" phylum="Mollusca" rank="subSpecies" species="jadertina" subSpecies="sinjana">
<emphasis italics="true" pageId="0" pageNumber="89">Kerkia jadertina sinjana</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="Kuščer">Kuscer</normalizedToken>
, 1933).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="89" type="molecular distinctness and relationships of horatia ozimeci">
<paragraph pageId="0" pageNumber="89">
Molecular distinctness and relationships of
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Horatia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatia ozimeci" order="Littorinimorpha" pageId="0" pageNumber="89" phylum="Mollusca" rank="species" species="ozimeci">
<emphasis italics="true" pageId="0" pageNumber="89">Horatia ozimeci</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph pageId="0" pageNumber="89">
We obtained eight new sequences of COI (409 bp, GenBank Accession Numbers MW448545-MW448552). The tests by
<bibRefCitation DOI="https://doi.org/10.1016/S1055-7903(02)00326-3" author="Xia, X" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="89" pagination="1 - 7" refId="B53" refString="Xia, X, Xie, Z, Salemi, M, Chen, L, Wang, Y, 2003. An index of substitution saturation and its application. Molecular Phylogenetics and Evolution 26: 1 - 7, DOI: https://doi.org/10.1016/S1055-7903(02)00326-3" title="An index of substitution saturation and its application." url="https://doi.org/10.1016/S1055-7903(02)00326-3" volume="26" year="2003">Xia et al. (2003)</bibRefCitation>
revealed no saturation. The topologies of the resulting phylograms were identical in both the ML and BI. All the seven sequences of
<taxonomicName genus="H." lsidName="H. klecakiana" pageId="0" pageNumber="89" rank="species" species="klecakiana">
<emphasis italics="true" pageId="0" pageNumber="89">H. klecakiana</emphasis>
</taxonomicName>
, collected at five localities, were identical.
<taxonomicName genus="H." lsidName="H. ozimeci" pageId="0" pageNumber="89" rank="species" species="ozimeci">
<emphasis italics="true" pageId="0" pageNumber="89">H. ozimeci</emphasis>
</taxonomicName>
formed a sister clade with
<taxonomicName genus="H." lsidName="H. klecakiana" pageId="0" pageNumber="89" rank="species" species="klecakiana">
<emphasis italics="true" pageId="0" pageNumber="89">H. klecakiana</emphasis>
</taxonomicName>
(bootstrap support 100%, Bayesian probability 1.0), confirming the congenerity of the two taxa (Fig.
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Maximum Likelihood tree inferred from mitochondrial COI. Bootstrap support above 60 % and corresponding Bayesian probabilities are given. Bold indicates newly obtained sequences." figureDoi="10.3897/subtbiol.37.61573.figure6" httpUri="https://binary.pensoft.net/fig/509945" pageId="0" pageNumber="89">6</figureCitation>
). Within genus
<taxonomicName authorityName="Bourguignat" authorityYear="1887" class="Gastropoda" family="Hydrobiidae" genus="Horatia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatia" order="Littorinimorpha" pageId="0" pageNumber="89" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="89">Horatia</emphasis>
</taxonomicName>
the p-distance between the taxa was 0.074. This well supported clade belongs to the
<taxonomicName authorityName="D.W.Taylor" authorityYear="1966" lsidName="" pageId="0" pageNumber="89" rank="subFamily" subFamily="Horatiinae">Horatiinae</taxonomicName>
, subfamily of
<taxonomicName class="Gastropoda" family="Hydrobiidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Littorinimorpha" pageId="0" pageNumber="89" phylum="Mollusca" rank="family">Hydrobiidae</taxonomicName>
. Deeper relationships within the
<taxonomicName authorityName="D.W.Taylor" authorityYear="1966" lsidName="" pageId="0" pageNumber="89" rank="subFamily" subFamily="Horatiinae">Horatiinae</taxonomicName>
remain unresolved, because of the lack of acceptable support for deeper nodes, which is typical of the phylograms based on COI.
</paragraph>
<caption ID-Table-UUID="B70765E2784600EB05E676C7B53632D9" httpUri="http://table.plazi.org/id/B70765E2784600EB05E676C7B53632D9" pageId="0" pageNumber="89" start="Table 2" startId="T2">
<paragraph pageId="0" pageNumber="89">
<emphasis bold="true" pageId="0" pageNumber="89">Table 2.</emphasis>
Shell measurements of the
<taxonomicName authorityName="Bourguignat" authorityYear="1887" class="Gastropoda" family="Hydrobiidae" genus="Horatia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Horatia" order="Littorinimorpha" pageId="0" pageNumber="89" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="89">Horatia</emphasis>
</taxonomicName>
: I-IV
<taxonomicName genus="H." lsidName="H. ozimeci" pageId="0" pageNumber="89" rank="species" species="ozimeci">
<emphasis italics="true" pageId="0" pageNumber="89">H. ozimeci</emphasis>
</taxonomicName>
(I sequenced specimen, II holotype, in bold; III-IV paratypes); V-VII sequenced specimens of
<taxonomicName genus="H." lsidName="H. klecakiana" pageId="0" pageNumber="89" rank="species" species="klecakiana">
<emphasis italics="true" pageId="0" pageNumber="89">H. klecakiana</emphasis>
</taxonomicName>
; VIII
<taxonomicName genus="H." lsidName="H. stygorumina" pageId="0" pageNumber="89" rank="species" species="stygorumina">
<emphasis italics="true" pageId="0" pageNumber="89">H. stygorumina</emphasis>
</taxonomicName>
, holotype, in bold. Measurements are shown in Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Shell variability of two Horatia species A-E H. ozimeci A sequenced specimen (2 H 76) B holotype OZRM 11603 C-E paratypes E NHMW 113607, (Photo NHMW by Ivo Gallmetzer) F-H H. stygorumina F holotype OZRM 11600 G-H paratypes NHMW 113608, (Photo NHMW by Gallmetzer) I Cf. Horatia knorri Schuett, 1961, spring of Ombla in Komolac, holotype SMF 164247 (Photo SMF by Sigrid Hof) J-P sequenced specimens of H. klecakiana J Vrelo Kosinac 2 K H. klecakiana, Gala spring L H. klecakiana, Ruda Begusa M, N, P H. klecakiana, Vrelo Kosinac 1 O H. klecakiana, Grabske Mlinice. The shell measurements are also shown: a shell height, b body whorl breadth, c aperture height, d spire height, e aperture breadth, α apex angle, β angle between body whorl suture and horizontal surface. Scale bars: 1 mm." figureDoi="10.3897/subtbiol.37.61573.figure4" httpUri="https://binary.pensoft.net/fig/509943" pageId="0" pageNumber="89">4J</figureCitation>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="89">
<table pageId="0" pageNumber="89">
<tr pageId="0" pageNumber="89">
<th colspan="1" isEmpty="true" pageId="0" pageNumber="89" rowspan="1">-</th>
<th colspan="1" pageId="0" pageNumber="89" rowspan="1">I</th>
<th colspan="1" pageId="0" pageNumber="89" rowspan="1">II</th>
<th colspan="1" pageId="0" pageNumber="89" rowspan="1">III</th>
<th colspan="1" pageId="0" pageNumber="89" rowspan="1">IV</th>
<th colspan="1" pageId="0" pageNumber="89" rowspan="1">
<taxonomicName genus="H." lsidName="H. ozimeci" pageId="0" pageNumber="89" rank="species" species="ozimeci">
<emphasis italics="true" pageId="0" pageNumber="89">H. ozimeci</emphasis>
</taxonomicName>
</th>
<th colspan="1" pageId="0" pageNumber="89" rowspan="1">V</th>
<th colspan="1" pageId="0" pageNumber="89" rowspan="1">VI</th>
<th colspan="1" pageId="0" pageNumber="89" rowspan="1">VII</th>
<th colspan="1" pageId="0" pageNumber="89" rowspan="1">VIII</th>
</tr>
<tr pageId="0" pageNumber="89">
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="89">a</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">2.14</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="89">1.70</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">2.02</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">1.71</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">
1.89
<normalizedToken originalValue="±">+/-</normalizedToken>
0.22
</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">1.61</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">1.51</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">1.61</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="89">2.08</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="89">
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="89">b</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">1.62</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="89">1.58</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">1.56</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">1.38</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">
1.54
<normalizedToken originalValue="±">+/-</normalizedToken>
0.11
</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">1.33</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">1.19</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">1.26</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="89">1.57</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="89">
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="89">c</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">1.22</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="89">1.12</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">1.29</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">1.05</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">
1.17
<normalizedToken originalValue="±">+/-</normalizedToken>
0.11
</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">1.14</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">0.88</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">1.08</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="89">1.24</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="89">
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="89">d</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">0.23</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="89">0.32</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">0.18</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">0.26</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">
0.25
<normalizedToken originalValue="±">+/-</normalizedToken>
0.06
</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">0.17</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">0.26</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">0.25</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="89">0.49</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="89">
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="89">e</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">1.05</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="89">1.05</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">1.18</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">0.95</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">
1.06
<normalizedToken originalValue="±">+/-</normalizedToken>
0.09
</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">0.97</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">0.91</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">0.92</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="89">1.08</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="89">
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="89">
<normalizedToken originalValue="α">α</normalizedToken>
</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">131</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="89">131</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">123</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">124</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">
127.25
<normalizedToken originalValue="±">+/-</normalizedToken>
4.35
</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">112</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">114</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">113</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="89">104</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="89">
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">
<emphasis italics="true" pageId="0" pageNumber="89">
<normalizedToken originalValue="β">β</normalizedToken>
</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">4</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="89">9</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">8</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">9</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">
7.50
<normalizedToken originalValue="±">+/-</normalizedToken>
2.38
</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">6</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">11</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">10</td>
<td colspan="1" pageId="0" pageNumber="89" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="89">8</emphasis>
</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>
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