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<document id="EE67A2D987AA64968DC367620DB63987" ID-CLB-Dataset="55429" ID-DOI="10.11646/zootaxa.5336.3.8" ID-GBIF-Dataset="c8c3de88-444a-45f7-975a-a0a36f26d9be" ID-ISSN="1175-5326" ID-Zenodo-Dep="8281813" ID-ZooBank="597932E9-187A-453D-9D0D-3FBBA59BEDC2" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="julia" checkinTime="1692951331172" checkinUser="plazi" docAuthor="Zilberman, Bruno &amp; Pires-Silva, Carlos M." docDate="2023" docId="03B087EE3C76FFD4AE9CFC51FDC2FDD4" docLanguage="en" docName="zootaxa.5336.3.8.pdf" docOrigin="Zootaxa 5336 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5336.3.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Austrospirachtha carrijoi Zilberman &amp; Pires-Silva 2023, sp. nov." docType="treatment" docVersion="4" lastPageNumber="432" masterDocId="FF89FF963C77FFDCAE0BFFF8FF8EFFB5" masterDocTitle="A new species and morphological notes on the remarkable termitophilous genus Austrospirachtha Watson from Australia (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="432" masterPageNumber="424" pageNumber="425" updateTime="1692978800748" updateUser="julia" zenodo-license-document="CLOSED">
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<mods:title id="A6A303A6C45283E64A31213E2A392D6B">A new species and morphological notes on the remarkable termitophilous genus Austrospirachtha Watson from Australia (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
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<mods:namePart id="BCEC19793DF893F0FFC7FEB3467BAEA3">Zilberman, Bruno</mods:namePart>
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<mods:affiliation id="69F33C651CED98E7EB23CC237D44E364">Programa de pós graduação em Sistemática, Taxonomia animal e Biodiversidade, Museu de Zoologia da Universidade de São Paulo, Laboratório de Coleoptera, Avenida Nazaré 481, 04263 - 000 São Paulo, SP, Brazil. &amp; https: // orcid. org / 0000 - 0003 - 2613 - 4827</mods:affiliation>
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<mods:namePart id="60CDAA93C93C952A0A1517A5BC341D56">Pires-Silva, Carlos M.</mods:namePart>
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<mods:affiliation id="DEB6E196430DB4D4AD03286A2A330B84">Programa de pós graduação em Sistemática, Taxonomia animal e Biodiversidade, Museu de Zoologia da Universidade de São Paulo, Laboratório de Coleoptera, Avenida Nazaré 481, 04263 - 000 São Paulo, SP, Brazil. &amp; https: // orcid. org / 0000 - 0002 - 6195 - 8648</mods:affiliation>
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<treatment id="03B087EE3C76FFD4AE9CFC51FDC2FDD4" ID-DOI="http://doi.org/10.5281/zenodo.8282444" ID-Zenodo-Dep="8282444" LSID="urn:lsid:plazi:treatment:03B087EE3C76FFD4AE9CFC51FDC2FDD4" httpUri="http://treatment.plazi.org/id/03B087EE3C76FFD4AE9CFC51FDC2FDD4" lastPageId="8" lastPageNumber="432" pageId="1" pageNumber="425">
<subSubSection id="C30365733C76FFDDAE9CFC51FCE4FC76" box="[151,874,937,964]" pageId="1" pageNumber="425" type="nomenclature">
<paragraph id="8BA636F83C76FFDDAE9CFC51FCE4FC76" blockId="1.[151,874,937,964]" box="[151,874,937,964]" pageId="1" pageNumber="425">
<heading id="D0EE81943C76FFDDAE9CFC51FCE4FC76" bold="true" box="[151,874,937,964]" fontSize="11" level="1" pageId="1" pageNumber="425" reason="1">
<emphasis id="B96DEAEA3C76FFDDAE9CFC51FCE4FC76" bold="true" box="[151,874,937,964]" pageId="1" pageNumber="425">
<taxonomicName id="4C194D7B3C76FFDDAE9CFC51FC88FC76" authority="Zilberman &amp; Pires-Silva" authorityName="Zilberman &amp; Pires-Silva" authorityYear="2023" box="[151,774,937,964]" class="Insecta" family="Staphylinidae" genus="Austrospirachtha" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="425" phylum="Arthropoda" rank="species" species="carrijoi" status="sp. nov.">
<emphasis id="B96DEAEA3C76FFDDAE9CFC51FE45FC76" bold="true" box="[151,459,937,964]" italics="true" pageId="1" pageNumber="425">Austrospirachtha carrijoi</emphasis>
Zilberman &amp; Pires-Silva
</taxonomicName>
<taxonomicNameLabel id="A25E57913C76FFDDAD06FC51FCE4FC76" box="[781,874,937,963]" pageId="1" pageNumber="425" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C30365733C76FFDAAE9CFC09FD2AFDD4" lastPageId="6" lastPageNumber="430" pageId="1" pageNumber="425" type="description">
<paragraph id="8BA636F83C76FFDDAE9CFC09FA1AFBBE" blockId="1.[151,1437,1009,2008]" box="[151,1428,1009,1036]" pageId="1" pageNumber="425">
<emphasis id="B96DEAEA3C76FFDDAE9CFC09FEDEFBBE" bold="true" box="[151,336,1009,1035]" pageId="1" pageNumber="425">Measurements:</emphasis>
Female (
<figureCitation id="13222A7D3C76FFDDAFB3FC09FD90FBBE" box="[440,542,1009,1035]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[152,255,1513,1537]" captionStartId-1="3.[152,255,1943,1967]" captionTargetBox-0="[151,1435,237,1489]" captionTargetBox-1="[156,1431,181,1919]" captionTargetId-0="figure-56@2.[151,1435,237,1489]" captionTargetId-1="figure-37@3.[156,1431,181,1919]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Austrospirachtha carrijoi sp. nov., female, holotype: a, habitus lateral; b, habitus dorsal. About 2.4 mm from the curvature of the abdomen to the apex of mandibula." captionText-1="FIGURE 2. Austrospirachtha carrijoi sp. nov., female, paratype: a, tergite II and sternite II “ring” modification by secondary sclerotization; b, habitus ventral." figureDoi-0="http://doi.org/10.5281/zenodo.8281819" figureDoi-1="http://doi.org/10.5281/zenodo.8281823" httpUri-0="https://zenodo.org/record/8281819/files/figure.png" httpUri-1="https://zenodo.org/record/8281823/files/figure.png" pageId="1" pageNumber="425">Figs. 12</figureCitation>
)—about 2.4 mm from the curvature of the abdomen to the apex of mandibula.
</paragraph>
<paragraph id="8BA636F83C76FFDDAECCFBEDFEF6FB9A" blockId="1.[151,1437,1009,2008]" box="[199,376,1045,1071]" pageId="1" pageNumber="425">Male unknown.</paragraph>
<paragraph id="8BA636F83C76FFDDAECCFBC1FD57FB56" blockId="1.[151,1437,1009,2008]" pageId="1" pageNumber="425">
Body highly physogastric, with constrictions and three pairs of pseudoappeandages; sclerotized parts light to dark brown. Maxilla with first palpomere distinct (
<figureCitation id="13222A7D3C76FFDDACD9FBA5FCA5FBC2" box="[722,811,1117,1143]" captionStart="FIGURE 3" captionStartId="4.[152,255,1650,1674]" captionTargetBox="[160,1426,181,1625]" captionTargetId="figure-29@4.[160,1426,181,1625]" captionTargetPageId="4" captionText="FIGURE 3. Austrospirachtha carrijoi sp. nov., female, paratype: a, head ventral; b, mandibles (dorsal view); c, labrum (dorsal view); d, maxilla (dorsal view); e, meso-metathorax and first tergite attached to metanotum (seen through transparence as the meso-metanotum); f, elytron (dorsal view); g, elytron, close-up (dorsal view), details with arrows pointing to the three main bristles; h, tergite VIII (dorsal view); i, tergite VII (dorsal view), distal region, detail of chaetotaxy; j, sternite VIII (dorsal view); k, tergites IX and X (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.8281827" httpUri="https://zenodo.org/record/8281827/files/figure.png" pageId="1" pageNumber="425">Fig. 3D</figureCitation>
), and post-mentum with two distinguishable main pairs of bristles (
<figureCitation id="13222A7D3C76FFDDAF10FB79FEF9FB29" box="[283,375,1153,1180]" captionStart="FIGURE 5" captionStartId="6.[152,255,2008,2032]" captionTargetBox="[221,1366,1287,1980]" captionTargetId="figure-560@6.[216,1371,1281,1985]" captionTargetPageId="6" captionText="FIGURE 5. Chaetotaxy of mentum-submentum. a, Austrospirachta carrijoi sp. nov.; b, A. mimetes." figureDoi="http://doi.org/10.5281/zenodo.8281833" httpUri="https://zenodo.org/record/8281833/files/figure.png" pageId="1" pageNumber="425">Fig. 5A</figureCitation>
). Abdominal segment III inflated into an ovoid region, longer than the subsequent segments combined; sternite V with secondary sclerotization on distal and lateral margins (
<figureCitation id="13222A7D3C76FFDDAA28FB5DFBF2FB0A" box="[1059,1148,1189,1215]" captionStart="FIGURE 4" captionStartId="5.[152,255,1904,1928]" captionTargetBox="[226,1361,191,1873]" captionTargetId="figure-37@5.[216,1372,181,1880]" captionTargetPageId="5" captionText="FIGURE 4. Austrospirachtha carrijoi sp. nov., and A. mimetes illustrations. ae, Austrospirachtha carrijoi sp. nov.: a, dorsal view of abdomen; b, sternite IV (dorsal view); c, sternite V (dorsal view); d, sterniteVI (dorsal view); e, sterniteVII (dorsal view); fj, A. mimetes: f, dorsal view of abdomen; g, sternite IV (dorsal view); h, sternite V (dorsal view); i, sternite VI (dorsal view); j, sternite VII (dorsal view). ps: primary sclerotization, se: secondary sclerotization. Blue arrows indicate pseudoappendages." figureDoi="http://doi.org/10.5281/zenodo.8281829" httpUri="https://zenodo.org/record/8281829/files/figure.png" pageId="1" pageNumber="425">Fig. 4C</figureCitation>
); sternite VI with sinuous secondary sclerotization on distal margin (
<figureCitation id="13222A7D3C76FFDDAC79FB31FD43FB56" box="[626,717,1225,1251]" captionStart="FIGURE 4" captionStartId="5.[152,255,1904,1928]" captionTargetBox="[226,1361,191,1873]" captionTargetId="figure-37@5.[216,1372,181,1880]" captionTargetPageId="5" captionText="FIGURE 4. Austrospirachtha carrijoi sp. nov., and A. mimetes illustrations. ae, Austrospirachtha carrijoi sp. nov.: a, dorsal view of abdomen; b, sternite IV (dorsal view); c, sternite V (dorsal view); d, sterniteVI (dorsal view); e, sterniteVII (dorsal view); fj, A. mimetes: f, dorsal view of abdomen; g, sternite IV (dorsal view); h, sternite V (dorsal view); i, sternite VI (dorsal view); j, sternite VII (dorsal view). ps: primary sclerotization, se: secondary sclerotization. Blue arrows indicate pseudoappendages." figureDoi="http://doi.org/10.5281/zenodo.8281829" httpUri="https://zenodo.org/record/8281829/files/figure.png" pageId="1" pageNumber="425">Fig. 4D</figureCitation>
).
</paragraph>
<paragraph id="8BA636F83C76FFDDAECCFB15FE6AF902" blockId="1.[151,1437,1009,2008]" pageId="1" pageNumber="425">
<emphasis id="B96DEAEA3C76FFDDAECCFB15FE89FAB2" bold="true" box="[199,263,1261,1287]" pageId="1" pageNumber="425">Head</emphasis>
slightly wider than long, overall, seemingly subquadrate (
<figureCitation id="13222A7D3C76FFDDADBEFB15FB9DFAB2" box="[949,1043,1261,1287]" captionStart="FIGURE 3" captionStartId="4.[152,255,1650,1674]" captionTargetBox="[160,1426,181,1625]" captionTargetId="figure-29@4.[160,1426,181,1625]" captionTargetPageId="4" captionText="FIGURE 3. Austrospirachtha carrijoi sp. nov., female, paratype: a, head ventral; b, mandibles (dorsal view); c, labrum (dorsal view); d, maxilla (dorsal view); e, meso-metathorax and first tergite attached to metanotum (seen through transparence as the meso-metanotum); f, elytron (dorsal view); g, elytron, close-up (dorsal view), details with arrows pointing to the three main bristles; h, tergite VIII (dorsal view); i, tergite VII (dorsal view), distal region, detail of chaetotaxy; j, sternite VIII (dorsal view); k, tergites IX and X (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.8281827" httpUri="https://zenodo.org/record/8281827/files/figure.png" pageId="1" pageNumber="425">Fig. 3A</figureCitation>
); anterior region of vertex highly impressed, leaving areas above eyes seemingly carinated; posterior margin on lateral view with one notch (
<figureCitation id="13222A7D3C76FFDDAB64FAE9FF32FAFA" captionStart="FIGURE 1" captionStartId="2.[152,255,1513,1537]" captionTargetBox="[151,1435,237,1489]" captionTargetId="figure-56@2.[151,1435,237,1489]" captionTargetPageId="2" captionText="FIGURE 1. Austrospirachtha carrijoi sp. nov., female, holotype: a, habitus lateral; b, habitus dorsal. About 2.4 mm from the curvature of the abdomen to the apex of mandibula." figureDoi="http://doi.org/10.5281/zenodo.8281819" httpUri="https://zenodo.org/record/8281819/files/figure.png" pageId="1" pageNumber="425">Fig. 1A</figureCitation>
).
<emphasis id="B96DEAEA3C76FFDDAED9FACDFE83FAFA" bold="true" box="[210,269,1333,1359]" pageId="1" pageNumber="425">Gula</emphasis>
broad, expanding posteriorly with occipital region occupying greater portion of total head width (
<figureCitation id="13222A7D3C76FFDDAB64FACDFF29FAC6" captionStart="FIGURE 3" captionStartId="4.[152,255,1650,1674]" captionTargetBox="[160,1426,181,1625]" captionTargetId="figure-29@4.[160,1426,181,1625]" captionTargetPageId="4" captionText="FIGURE 3. Austrospirachtha carrijoi sp. nov., female, paratype: a, head ventral; b, mandibles (dorsal view); c, labrum (dorsal view); d, maxilla (dorsal view); e, meso-metathorax and first tergite attached to metanotum (seen through transparence as the meso-metanotum); f, elytron (dorsal view); g, elytron, close-up (dorsal view), details with arrows pointing to the three main bristles; h, tergite VIII (dorsal view); i, tergite VII (dorsal view), distal region, detail of chaetotaxy; j, sternite VIII (dorsal view); k, tergites IX and X (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.8281827" httpUri="https://zenodo.org/record/8281827/files/figure.png" pageId="1" pageNumber="425">Fig. 3</figureCitation>
).
<emphasis id="B96DEAEA3C76FFDDAECBFAA1FEA9FAC6" bold="true" box="[192,295,1369,1395]" pageId="1" pageNumber="425">Antenna</emphasis>
with 11 antennomeres, scape almost as long as antennomeres 24 combined; antennomeres 510 subquadrate; antennomere 11 elongate, about as long as antennomeres 910 combined.
<emphasis id="B96DEAEA3C76FFDDAA73FA85FB52FA22" bold="true" box="[1144,1244,1405,1431]" pageId="1" pageNumber="425">Labrum</emphasis>
wider than long, with anterior margin sinuous; two short and closer to each other bristles on extreme anterior region at each side (
<emphasis id="B96DEAEA3C76FFDDAB7CFA59FF3AFA6A" italics="true" pageId="1" pageNumber="425">d2- d1</emphasis>
); right above, one very long bristle on each side (
<emphasis id="B96DEAEA3C76FFDDACEEFA3DFC86FA6A" box="[741,776,1477,1503]" italics="true" pageId="1" pageNumber="425">m1</emphasis>
), next to lateral margin; one to two mid to long bristles on posterior region (
<emphasis id="B96DEAEA3C76FFDDAF51FA11FE15F9B6" box="[346,411,1513,1540]" italics="true" pageId="1" pageNumber="425">p2-p1</emphasis>
) (
<figureCitation id="13222A7D3C76FFDDAFBFFA11FE77F9B6" box="[436,505,1513,1539]" captionStart="FIGURE 3" captionStartId="4.[152,255,1650,1674]" captionTargetBox="[160,1426,181,1625]" captionTargetId="figure-29@4.[160,1426,181,1625]" captionTargetPageId="4" captionText="FIGURE 3. Austrospirachtha carrijoi sp. nov., female, paratype: a, head ventral; b, mandibles (dorsal view); c, labrum (dorsal view); d, maxilla (dorsal view); e, meso-metathorax and first tergite attached to metanotum (seen through transparence as the meso-metanotum); f, elytron (dorsal view); g, elytron, close-up (dorsal view), details with arrows pointing to the three main bristles; h, tergite VIII (dorsal view); i, tergite VII (dorsal view), distal region, detail of chaetotaxy; j, sternite VIII (dorsal view); k, tergites IX and X (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.8281827" httpUri="https://zenodo.org/record/8281827/files/figure.png" pageId="1" pageNumber="425">Fig. 3</figureCitation>
).
<emphasis id="B96DEAEA3C76FFDDAC05FA11FD02F9B6" bold="true" box="[526,652,1513,1539]" pageId="1" pageNumber="425">Mandibles</emphasis>
symmetrical, slim, toothless (
<figureCitation id="13222A7D3C76FFDDADD5FA11FBB8F9B6" box="[990,1078,1513,1539]" captionStart="FIGURE 3" captionStartId="4.[152,255,1650,1674]" captionTargetBox="[160,1426,181,1625]" captionTargetId="figure-29@4.[160,1426,181,1625]" captionTargetPageId="4" captionText="FIGURE 3. Austrospirachtha carrijoi sp. nov., female, paratype: a, head ventral; b, mandibles (dorsal view); c, labrum (dorsal view); d, maxilla (dorsal view); e, meso-metathorax and first tergite attached to metanotum (seen through transparence as the meso-metanotum); f, elytron (dorsal view); g, elytron, close-up (dorsal view), details with arrows pointing to the three main bristles; h, tergite VIII (dorsal view); i, tergite VII (dorsal view), distal region, detail of chaetotaxy; j, sternite VIII (dorsal view); k, tergites IX and X (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.8281827" httpUri="https://zenodo.org/record/8281827/files/figure.png" pageId="1" pageNumber="425">Fig. 3B</figureCitation>
).
<emphasis id="B96DEAEA3C76FFDDAA40FA11FB3CF9B6" bold="true" box="[1099,1202,1513,1539]" pageId="1" pageNumber="425">Mentum</emphasis>
fused to submentum (postmentum), subtrapezoidal, one long main bristle on each side of anterior region, each right in front of a medium sized one (
<figureCitation id="13222A7D3C76FFDDAF19F9C9FED6F9F9" box="[274,344,1585,1612]" captionStart="FIGURE 5" captionStartId="6.[152,255,2008,2032]" captionTargetBox="[221,1366,1287,1980]" captionTargetId="figure-560@6.[216,1371,1281,1985]" captionTargetPageId="6" captionText="FIGURE 5. Chaetotaxy of mentum-submentum. a, Austrospirachta carrijoi sp. nov.; b, A. mimetes." figureDoi="http://doi.org/10.5281/zenodo.8281833" httpUri="https://zenodo.org/record/8281833/files/figure.png" pageId="1" pageNumber="425">Fig. 5</figureCitation>
).
<emphasis id="B96DEAEA3C76FFDDAF65F9C9FE75F9FE" bold="true" box="[366,507,1585,1611]" pageId="1" pageNumber="425">Prementum</emphasis>
extremely reduced, labial palpi indistinct.
<emphasis id="B96DEAEA3C76FFDDADE9F9C9FBB0F9FE" bold="true" box="[994,1086,1585,1611]" pageId="1" pageNumber="425">Maxilla</emphasis>
with galea subquadrate, lacina narrower, and palpi 4-jointed: first palpomere transverse, about four times shorter than second, which is elongate; third palpomere longer than wide, narrower at base; fourth palpomere reduced to less than half of length of filamentous sensillae (
<figureCitation id="13222A7D3C76FFDDAF99F965FE59F902" box="[402,471,1693,1719]" captionStart="FIGURE 3" captionStartId="4.[152,255,1650,1674]" captionTargetBox="[160,1426,181,1625]" captionTargetId="figure-29@4.[160,1426,181,1625]" captionTargetPageId="4" captionText="FIGURE 3. Austrospirachtha carrijoi sp. nov., female, paratype: a, head ventral; b, mandibles (dorsal view); c, labrum (dorsal view); d, maxilla (dorsal view); e, meso-metathorax and first tergite attached to metanotum (seen through transparence as the meso-metanotum); f, elytron (dorsal view); g, elytron, close-up (dorsal view), details with arrows pointing to the three main bristles; h, tergite VIII (dorsal view); i, tergite VII (dorsal view), distal region, detail of chaetotaxy; j, sternite VIII (dorsal view); k, tergites IX and X (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.8281827" httpUri="https://zenodo.org/record/8281827/files/figure.png" pageId="1" pageNumber="425">Fig. 3</figureCitation>
).
</paragraph>
<paragraph id="8BA636F83C76FFDEAECCF939FD9FFF60" blockId="1.[151,1437,1009,2008]" lastBlockId="2.[151,1436,151,213]" lastPageId="2" lastPageNumber="426" pageId="1" pageNumber="425">
<emphasis id="B96DEAEA3C76FFDDAECCF939FEAEF96E" bold="true" box="[199,288,1729,1755]" pageId="1" pageNumber="425">Thorax</emphasis>
with considerable areas of exposed membrane between head and prothorax, and prothorax and mesometathorax (
<figureCitation id="13222A7D3C76FFDDAF2FF91DFEFCF94A" box="[292,370,1765,1791]" captionStart="FIGURE 1" captionStartId="2.[152,255,1513,1537]" captionTargetBox="[151,1435,237,1489]" captionTargetId="figure-56@2.[151,1435,237,1489]" captionTargetPageId="2" captionText="FIGURE 1. Austrospirachtha carrijoi sp. nov., female, holotype: a, habitus lateral; b, habitus dorsal. About 2.4 mm from the curvature of the abdomen to the apex of mandibula." figureDoi="http://doi.org/10.5281/zenodo.8281819" httpUri="https://zenodo.org/record/8281819/files/figure.png" pageId="1" pageNumber="425">Figs. 1</figureCitation>
;
<figureCitation id="13222A7D3C76FFDDAF8BF91DFE1FF94A" box="[384,401,1765,1791]" captionStart="FIGURE 2" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[156,1431,181,1919]" captionTargetId="figure-37@3.[156,1431,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Austrospirachtha carrijoi sp. nov., female, paratype: a, tergite II and sternite II “ring” modification by secondary sclerotization; b, habitus ventral." figureDoi="http://doi.org/10.5281/zenodo.8281823" httpUri="https://zenodo.org/record/8281823/files/figure.png" pageId="1" pageNumber="425">2</figureCitation>
).
<emphasis id="B96DEAEA3C76FFDDAFAEF91DFD90F94A" bold="true" box="[421,542,1765,1791]" pageId="1" pageNumber="425">Pronotum</emphasis>
slightly wider than long;
<emphasis id="B96DEAEA3C76FFDDAD35F91DFC45F94A" bold="true" box="[830,971,1765,1791]" pageId="1" pageNumber="425">prosternum</emphasis>
with small area of sclerotization between legs, and other areas membranous.
<emphasis id="B96DEAEA3C76FFDDAC16F8F1FD22F896" bold="true" box="[541,684,1801,1827]" pageId="1" pageNumber="425">Mesothorax</emphasis>
shorter than metathorax (
<figureCitation id="13222A7D3C76FFDDADC3F8F1FB82F896" box="[968,1036,1801,1827]" captionStart="FIGURE 3" captionStartId="4.[152,255,1650,1674]" captionTargetBox="[160,1426,181,1625]" captionTargetId="figure-29@4.[160,1426,181,1625]" captionTargetPageId="4" captionText="FIGURE 3. Austrospirachtha carrijoi sp. nov., female, paratype: a, head ventral; b, mandibles (dorsal view); c, labrum (dorsal view); d, maxilla (dorsal view); e, meso-metathorax and first tergite attached to metanotum (seen through transparence as the meso-metanotum); f, elytron (dorsal view); g, elytron, close-up (dorsal view), details with arrows pointing to the three main bristles; h, tergite VIII (dorsal view); i, tergite VII (dorsal view), distal region, detail of chaetotaxy; j, sternite VIII (dorsal view); k, tergites IX and X (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.8281827" httpUri="https://zenodo.org/record/8281827/files/figure.png" pageId="1" pageNumber="425">Fig. 3</figureCitation>
).
<emphasis id="B96DEAEA3C76FFDDAA2BF8F1FB16F896" bold="true" box="[1056,1176,1801,1827]" pageId="1" pageNumber="425">Scutellum</emphasis>
suboval.
<emphasis id="B96DEAEA3C76FFDDAB09F8F1FA12F896" bold="true" box="[1282,1436,1801,1827]" pageId="1" pageNumber="425">Endosternite</emphasis>
with base extended laterally by secondary sclerotization, where metalegs attach; between these extended nodules, no secondary sclerotization was seen; arms very long, thin and contiguous (
<figureCitation id="13222A7D3C76FFDDAA1AF8A9FBD7F8DE" box="[1041,1113,1873,1899]" captionStart="FIGURE 3" captionStartId="4.[152,255,1650,1674]" captionTargetBox="[160,1426,181,1625]" captionTargetId="figure-29@4.[160,1426,181,1625]" captionTargetPageId="4" captionText="FIGURE 3. Austrospirachtha carrijoi sp. nov., female, paratype: a, head ventral; b, mandibles (dorsal view); c, labrum (dorsal view); d, maxilla (dorsal view); e, meso-metathorax and first tergite attached to metanotum (seen through transparence as the meso-metanotum); f, elytron (dorsal view); g, elytron, close-up (dorsal view), details with arrows pointing to the three main bristles; h, tergite VIII (dorsal view); i, tergite VII (dorsal view), distal region, detail of chaetotaxy; j, sternite VIII (dorsal view); k, tergites IX and X (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.8281827" httpUri="https://zenodo.org/record/8281827/files/figure.png" pageId="1" pageNumber="425">Fig. 3</figureCitation>
).
<emphasis id="B96DEAEA3C76FFDDAA79F8A9FB43F8DE" bold="true" box="[1138,1229,1873,1899]" pageId="1" pageNumber="425">Elytron</emphasis>
subtrapezoidal in shape, with proximal and distal margins oblique to elytron length, but more strongly tapering in distal-lateral region of elytron, leaving round apex; two long bristles vertically aligned on central region, another long bristle more proximal-lateral placed; and sparser shorter bristles across elytron surface (
<figureCitation id="13222A7D3C76FFDDADDFF845FBF5F862" box="[980,1147,1981,2007]" captionStart="FIGURE 3" captionStartId="4.[152,255,1650,1674]" captionTargetBox="[160,1426,181,1625]" captionTargetId="figure-29@4.[160,1426,181,1625]" captionTargetPageId="4" captionText="FIGURE 3. Austrospirachtha carrijoi sp. nov., female, paratype: a, head ventral; b, mandibles (dorsal view); c, labrum (dorsal view); d, maxilla (dorsal view); e, meso-metathorax and first tergite attached to metanotum (seen through transparence as the meso-metanotum); f, elytron (dorsal view); g, elytron, close-up (dorsal view), details with arrows pointing to the three main bristles; h, tergite VIII (dorsal view); i, tergite VII (dorsal view), distal region, detail of chaetotaxy; j, sternite VIII (dorsal view); k, tergites IX and X (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.8281827" httpUri="https://zenodo.org/record/8281827/files/figure.png" pageId="1" pageNumber="425">Figs. 3F and G</figureCitation>
).
<emphasis id="B96DEAEA3C76FFDDAA86F845FA9DF862" bold="true" box="[1165,1299,1981,2007]" pageId="1" pageNumber="425">Hind wings</emphasis>
presumably shed off, leaving only small stubs (
<figureCitation id="13222A7D3C75FFDEAC2BFF6FFDEBFF04" box="[544,613,151,177]" captionStart="FIGURE 3" captionStartId="4.[152,255,1650,1674]" captionTargetBox="[160,1426,181,1625]" captionTargetId="figure-29@4.[160,1426,181,1625]" captionTargetPageId="4" captionText="FIGURE 3. Austrospirachtha carrijoi sp. nov., female, paratype: a, head ventral; b, mandibles (dorsal view); c, labrum (dorsal view); d, maxilla (dorsal view); e, meso-metathorax and first tergite attached to metanotum (seen through transparence as the meso-metanotum); f, elytron (dorsal view); g, elytron, close-up (dorsal view), details with arrows pointing to the three main bristles; h, tergite VIII (dorsal view); i, tergite VII (dorsal view), distal region, detail of chaetotaxy; j, sternite VIII (dorsal view); k, tergites IX and X (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.8281827" httpUri="https://zenodo.org/record/8281827/files/figure.png" pageId="2" pageNumber="426">Fig. 3</figureCitation>
). Legs developed, first tarsomere gradually increasing in length from pro to meso, and to hind legs (
<figureCitation id="13222A7D3C75FFDEAFB5FF43FD8EFF60" box="[446,512,187,213]" captionStart="FIGURE 1" captionStartId="2.[152,255,1513,1537]" captionTargetBox="[151,1435,237,1489]" captionTargetId="figure-56@2.[151,1435,237,1489]" captionTargetPageId="2" captionText="FIGURE 1. Austrospirachtha carrijoi sp. nov., female, holotype: a, habitus lateral; b, habitus dorsal. About 2.4 mm from the curvature of the abdomen to the apex of mandibula." figureDoi="http://doi.org/10.5281/zenodo.8281819" httpUri="https://zenodo.org/record/8281819/files/figure.png" pageId="2" pageNumber="426">Fig. 1</figureCitation>
).
</paragraph>
<caption id="DF6666703C75FFDEAE93FA11FD2AF990" ID-DOI="http://doi.org/10.5281/zenodo.8281819" ID-Zenodo-Dep="8281819" httpUri="https://zenodo.org/record/8281819/files/figure.png" pageId="2" pageNumber="426" startId="2.[152,255,1513,1537]" targetBox="[151,1435,237,1489]" targetPageId="2" targetType="figure">
<paragraph id="8BA636F83C75FFDEAE93FA11FD2AF990" blockId="2.[151,1436,1513,1573]" pageId="2" pageNumber="426">
<emphasis id="B96DEAEA3C75FFDEAE93FA11FE97F9B4" bold="true" box="[152,281,1513,1537]" pageId="2" pageNumber="426">FIGURE 1.</emphasis>
<taxonomicName id="4C194D7B3C75FFDEAF14FA11FDACF9B4" authority="Zilberman &amp; Pires-Silva, 2023" authorityName="Zilberman &amp; Pires-Silva" authorityYear="2023" box="[287,546,1513,1537]" class="Insecta" family="Staphylinidae" genus="Austrospirachtha" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="426" phylum="Arthropoda" rank="species" species="carrijoi" status="sp. nov.">
<emphasis id="B96DEAEA3C75FFDEAF14FA11FDACF9B4" box="[287,546,1513,1537]" italics="true" pageId="2" pageNumber="426">Austrospirachtha carrijoi</emphasis>
</taxonomicName>
<emphasis id="B96DEAEA3C75FFDEAC22FA11FDF4F9B4" bold="true" box="[553,634,1513,1537]" pageId="2" pageNumber="426">
<taxonomicNameLabel id="A25E57913C75FFDEAC22FA11FDF4F9B4" box="[553,634,1513,1537]" pageId="2" pageNumber="426" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female, holotype:
<emphasis id="B96DEAEA3C75FFDEAD4BFA11FCC3F9B4" bold="true" box="[832,845,1513,1537]" pageId="2" pageNumber="426">a</emphasis>
, habitus lateral;
<emphasis id="B96DEAEA3C75FFDEADFDFA11FB8AF9B4" bold="true" box="[1014,1028,1513,1537]" pageId="2" pageNumber="426">b</emphasis>
, habitus dorsal. About 2.4 mm from the curvature of the abdomen to the apex of mandibula.
</paragraph>
</caption>
<paragraph id="8BA636F83C75FFDAAECCF9ABFD2AFDD4" blockId="2.[151,1437,1619,2006]" lastBlockId="6.[151,1437,150,1257]" lastPageId="6" lastPageNumber="430" pageId="2" pageNumber="426">
<emphasis id="B96DEAEA3C75FFDEAECCF9ABFEB2F9D8" bold="true" box="[199,316,1619,1645]" pageId="2" pageNumber="426">Abdomen</emphasis>
strongly physogastric and modified, recurved at segment II, with apex anteriorly positioned, reaching the head (
<figureCitation id="13222A7D3C75FFDEAF0FF98FFECBF927" box="[260,325,1655,1682]" captionStart="FIGURE 1" captionStartId="2.[152,255,1513,1537]" captionTargetBox="[151,1435,237,1489]" captionTargetId="figure-56@2.[151,1435,237,1489]" captionTargetPageId="2" captionText="FIGURE 1. Austrospirachtha carrijoi sp. nov., female, holotype: a, habitus lateral; b, habitus dorsal. About 2.4 mm from the curvature of the abdomen to the apex of mandibula." figureDoi="http://doi.org/10.5281/zenodo.8281819" httpUri="https://zenodo.org/record/8281819/files/figure.png" pageId="2" pageNumber="426">Fig. 1</figureCitation>
). Third segment modified into an egg-shaped region considerably longer than remaining segments, that follows a strong constriction separating it from remaining segments (
<figureCitation id="13222A7D3C75FFDEADD0F963FBA4F903" box="[987,1066,1691,1718]" captionStart="FIGURE 1" captionStartId="2.[152,255,1513,1537]" captionTargetBox="[151,1435,237,1489]" captionTargetId="figure-56@2.[151,1435,237,1489]" captionTargetPageId="2" captionText="FIGURE 1. Austrospirachtha carrijoi sp. nov., female, holotype: a, habitus lateral; b, habitus dorsal. About 2.4 mm from the curvature of the abdomen to the apex of mandibula." figureDoi="http://doi.org/10.5281/zenodo.8281819" httpUri="https://zenodo.org/record/8281819/files/figure.png" pageId="2" pageNumber="426">Figs. 1</figureCitation>
,
<figureCitation id="13222A7D3C75FFDEAA32F964FBC4F903" box="[1081,1098,1692,1718]" captionStart="FIGURE 4" captionStartId="5.[152,255,1904,1928]" captionTargetBox="[226,1361,191,1873]" captionTargetId="figure-37@5.[216,1372,181,1880]" captionTargetPageId="5" captionText="FIGURE 4. Austrospirachtha carrijoi sp. nov., and A. mimetes illustrations. ae, Austrospirachtha carrijoi sp. nov.: a, dorsal view of abdomen; b, sternite IV (dorsal view); c, sternite V (dorsal view); d, sterniteVI (dorsal view); e, sterniteVII (dorsal view); fj, A. mimetes: f, dorsal view of abdomen; g, sternite IV (dorsal view); h, sternite V (dorsal view); i, sternite VI (dorsal view); j, sternite VII (dorsal view). ps: primary sclerotization, se: secondary sclerotization. Blue arrows indicate pseudoappendages." figureDoi="http://doi.org/10.5281/zenodo.8281829" httpUri="https://zenodo.org/record/8281829/files/figure.png" pageId="2" pageNumber="426">4</figureCitation>
); segments VIIVIII swollen. Abdomen bearing tree pairs of
<emphasis id="B96DEAEA3C75FFDEAFE4F938FD53F96F" bold="true" box="[495,733,1728,1754]" pageId="2" pageNumber="426">pseudoappeandages</emphasis>
: first pair, at pleural region of segment IV, contiguous, strongly swollen at their ends, bearing many short bristles; second pair, at pleural region of segment V, similar to first pair, except they are smaller and seemingly less sclerotized; third pair represented by strong sclerotized, gradually swollen towards apex, distally-directed contiguous pseudoappeandages, between segments VI and VIII, centrally disposed of (
<figureCitation id="13222A7D3C75FFDEAEB4F8B7FE83F8DF" box="[191,269,1871,1898]" captionStart="FIGURE 1" captionStartId="2.[152,255,1513,1537]" captionTargetBox="[151,1435,237,1489]" captionTargetId="figure-56@2.[151,1435,237,1489]" captionTargetPageId="2" captionText="FIGURE 1. Austrospirachtha carrijoi sp. nov., female, holotype: a, habitus lateral; b, habitus dorsal. About 2.4 mm from the curvature of the abdomen to the apex of mandibula." figureDoi="http://doi.org/10.5281/zenodo.8281819" httpUri="https://zenodo.org/record/8281819/files/figure.png" pageId="2" pageNumber="426">Figs. 1</figureCitation>
,
<figureCitation id="13222A7D3C75FFDEAF16F8A8FEA0F8DF" box="[285,302,1872,1898]" captionStart="FIGURE 4" captionStartId="5.[152,255,1904,1928]" captionTargetBox="[226,1361,191,1873]" captionTargetId="figure-37@5.[216,1372,181,1880]" captionTargetPageId="5" captionText="FIGURE 4. Austrospirachtha carrijoi sp. nov., and A. mimetes illustrations. ae, Austrospirachtha carrijoi sp. nov.: a, dorsal view of abdomen; b, sternite IV (dorsal view); c, sternite V (dorsal view); d, sterniteVI (dorsal view); e, sterniteVII (dorsal view); fj, A. mimetes: f, dorsal view of abdomen; g, sternite IV (dorsal view); h, sternite V (dorsal view); i, sternite VI (dorsal view); j, sternite VII (dorsal view). ps: primary sclerotization, se: secondary sclerotization. Blue arrows indicate pseudoappendages." figureDoi="http://doi.org/10.5281/zenodo.8281829" httpUri="https://zenodo.org/record/8281829/files/figure.png" pageId="2" pageNumber="426">4</figureCitation>
).
<emphasis id="B96DEAEA3C75FFDEAF48F8B7FE25F8DF" bold="true" box="[323,427,1871,1898]" pageId="2" pageNumber="426">Tergite I</emphasis>
represented by a thin sclerotized tergite, almost straight, attached to metanotum (
<figureCitation id="13222A7D3C75FFDEAB43F8B7FA00F8DF" box="[1352,1422,1871,1898]" captionStart="FIGURE 3" captionStartId="4.[152,255,1650,1674]" captionTargetBox="[160,1426,181,1625]" captionTargetId="figure-29@4.[160,1426,181,1625]" captionTargetPageId="4" captionText="FIGURE 3. Austrospirachtha carrijoi sp. nov., female, paratype: a, head ventral; b, mandibles (dorsal view); c, labrum (dorsal view); d, maxilla (dorsal view); e, meso-metathorax and first tergite attached to metanotum (seen through transparence as the meso-metanotum); f, elytron (dorsal view); g, elytron, close-up (dorsal view), details with arrows pointing to the three main bristles; h, tergite VIII (dorsal view); i, tergite VII (dorsal view), distal region, detail of chaetotaxy; j, sternite VIII (dorsal view); k, tergites IX and X (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.8281827" httpUri="https://zenodo.org/record/8281827/files/figure.png" pageId="2" pageNumber="426">Fig. 3</figureCitation>
).
<emphasis id="B96DEAEA3C75FFDEAE9CF88BFE89F83B" bold="true" box="[151,263,1907,1934]" pageId="2" pageNumber="426">Tergite II</emphasis>
straplike, with lateral secondary sclerotization fused to secondary sclerotization of sternite II, forming one complete ring (
<figureCitation id="13222A7D3C75FFDEAF35F86FFE0CF807" box="[318,386,1943,1970]" captionStart="FIGURE 2" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[156,1431,181,1919]" captionTargetId="figure-37@3.[156,1431,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Austrospirachtha carrijoi sp. nov., female, paratype: a, tergite II and sternite II “ring” modification by secondary sclerotization; b, habitus ventral." figureDoi="http://doi.org/10.5281/zenodo.8281823" httpUri="https://zenodo.org/record/8281823/files/figure.png" pageId="2" pageNumber="426">Fig. 2</figureCitation>
).
<emphasis id="B96DEAEA3C75FFDEAF9EF86FFD81F807" bold="true" box="[405,527,1943,1970]" pageId="2" pageNumber="426">Sternite II</emphasis>
modified, with acute distally subtriangular secondary sclerotization process (
<figureCitation id="13222A7D3C75FFDEAB6FF86FFE86F863" captionStart="FIGURE 2" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[156,1431,181,1919]" captionTargetId="figure-37@3.[156,1431,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Austrospirachtha carrijoi sp. nov., female, paratype: a, tergite II and sternite II “ring” modification by secondary sclerotization; b, habitus ventral." figureDoi="http://doi.org/10.5281/zenodo.8281823" httpUri="https://zenodo.org/record/8281823/files/figure.png" pageId="2" pageNumber="426">Figs. 2A and B</figureCitation>
).
<emphasis id="B96DEAEA3C75FFDEAF14F843FE4BF863" bold="true" box="[287,453,1979,2006]" pageId="2" pageNumber="426">Tergites III-V</emphasis>
broader and more sclerotized than tergite II, mostly concealed by recurved abdomen.
<emphasis id="B96DEAEA3C71FFDAAE9CFF6EFEAFFF04" bold="true" box="[151,289,150,177]" pageId="6" pageNumber="430">Sternite III</emphasis>
modified in expanded shield-like structure on dorsal view (morphological ventral) of ovoid part of the abdomen (
<figureCitation id="13222A7D3C71FFDAAF30FF43FE04FF60" box="[315,394,187,213]" captionStart="FIGURE 1" captionStartId="2.[152,255,1513,1537]" captionTargetBox="[151,1435,237,1489]" captionTargetId="figure-56@2.[151,1435,237,1489]" captionTargetPageId="2" captionText="FIGURE 1. Austrospirachtha carrijoi sp. nov., female, holotype: a, habitus lateral; b, habitus dorsal. About 2.4 mm from the curvature of the abdomen to the apex of mandibula." figureDoi="http://doi.org/10.5281/zenodo.8281819" httpUri="https://zenodo.org/record/8281819/files/figure.png" pageId="6" pageNumber="430">Figs. 1</figureCitation>
,
<figureCitation id="13222A7D3C71FFDAAF91FF43FE24FF60" box="[410,426,187,213]" captionStart="FIGURE 2" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[156,1431,181,1919]" captionTargetId="figure-37@3.[156,1431,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Austrospirachtha carrijoi sp. nov., female, paratype: a, tergite II and sternite II “ring” modification by secondary sclerotization; b, habitus ventral." figureDoi="http://doi.org/10.5281/zenodo.8281823" httpUri="https://zenodo.org/record/8281823/files/figure.png" pageId="6" pageNumber="430">2</figureCitation>
,
<figureCitation id="13222A7D3C71FFDAAFB3FF43FE51FF60" box="[440,479,187,213]" captionStart="FIGURE 4" captionStartId="5.[152,255,1904,1928]" captionTargetBox="[226,1361,191,1873]" captionTargetId="figure-37@5.[216,1372,181,1880]" captionTargetPageId="5" captionText="FIGURE 4. Austrospirachtha carrijoi sp. nov., and A. mimetes illustrations. ae, Austrospirachtha carrijoi sp. nov.: a, dorsal view of abdomen; b, sternite IV (dorsal view); c, sternite V (dorsal view); d, sterniteVI (dorsal view); e, sterniteVII (dorsal view); fj, A. mimetes: f, dorsal view of abdomen; g, sternite IV (dorsal view); h, sternite V (dorsal view); i, sternite VI (dorsal view); j, sternite VII (dorsal view). ps: primary sclerotization, se: secondary sclerotization. Blue arrows indicate pseudoappendages." figureDoi="http://doi.org/10.5281/zenodo.8281829" httpUri="https://zenodo.org/record/8281829/files/figure.png" pageId="6" pageNumber="430">4A</figureCitation>
).
<emphasis id="B96DEAEA3C71FFDAAFFFFF42FDF5FF60" bold="true" box="[500,635,186,213]" pageId="6" pageNumber="430">Sternite IV</emphasis>
strap-like, with one notch in the middle, and secondary sclerotization separated by membranous area (
<figureCitation id="13222A7D3C71FFDAAFF4FF27FDC2FF4C" box="[511,588,223,249]" captionStart="FIGURE 1" captionStartId="2.[152,255,1513,1537]" captionTargetBox="[151,1435,237,1489]" captionTargetId="figure-56@2.[151,1435,237,1489]" captionTargetPageId="2" captionText="FIGURE 1. Austrospirachtha carrijoi sp. nov., female, holotype: a, habitus lateral; b, habitus dorsal. About 2.4 mm from the curvature of the abdomen to the apex of mandibula." figureDoi="http://doi.org/10.5281/zenodo.8281819" httpUri="https://zenodo.org/record/8281819/files/figure.png" pageId="6" pageNumber="430">Figs. 1</figureCitation>
,
<figureCitation id="13222A7D3C71FFDAAC51FF27FDE5FF4C" box="[602,619,223,249]" captionStart="FIGURE 4" captionStartId="5.[152,255,1904,1928]" captionTargetBox="[226,1361,191,1873]" captionTargetId="figure-37@5.[216,1372,181,1880]" captionTargetPageId="5" captionText="FIGURE 4. Austrospirachtha carrijoi sp. nov., and A. mimetes illustrations. ae, Austrospirachtha carrijoi sp. nov.: a, dorsal view of abdomen; b, sternite IV (dorsal view); c, sternite V (dorsal view); d, sterniteVI (dorsal view); e, sterniteVII (dorsal view); fj, A. mimetes: f, dorsal view of abdomen; g, sternite IV (dorsal view); h, sternite V (dorsal view); i, sternite VI (dorsal view); j, sternite VII (dorsal view). ps: primary sclerotization, se: secondary sclerotization. Blue arrows indicate pseudoappendages." figureDoi="http://doi.org/10.5281/zenodo.8281829" httpUri="https://zenodo.org/record/8281829/files/figure.png" pageId="6" pageNumber="430">4</figureCitation>
);
<emphasis id="B96DEAEA3C71FFDAAC74FF26FD7AFF4C" bold="true" box="[639,756,222,249]" pageId="6" pageNumber="430">sternite V</emphasis>
strap-like, with one notch medially, and considerable area of secondary sclerotization, lateral and thinly distal to primary sclerotized piece (
<figureCitation id="13222A7D3C71FFDAAA04FEFBFBD3FEA8" box="[1039,1117,259,285]" captionStart="FIGURE 1" captionStartId="2.[152,255,1513,1537]" captionTargetBox="[151,1435,237,1489]" captionTargetId="figure-56@2.[151,1435,237,1489]" captionTargetPageId="2" captionText="FIGURE 1. Austrospirachtha carrijoi sp. nov., female, holotype: a, habitus lateral; b, habitus dorsal. About 2.4 mm from the curvature of the abdomen to the apex of mandibula." figureDoi="http://doi.org/10.5281/zenodo.8281819" httpUri="https://zenodo.org/record/8281819/files/figure.png" pageId="6" pageNumber="430">Figs. 1</figureCitation>
,
<figureCitation id="13222A7D3C71FFDAAA67FEFBFBF2FEA8" box="[1132,1148,259,285]" captionStart="FIGURE 4" captionStartId="5.[152,255,1904,1928]" captionTargetBox="[226,1361,191,1873]" captionTargetId="figure-37@5.[216,1372,181,1880]" captionTargetPageId="5" captionText="FIGURE 4. Austrospirachtha carrijoi sp. nov., and A. mimetes illustrations. ae, Austrospirachtha carrijoi sp. nov.: a, dorsal view of abdomen; b, sternite IV (dorsal view); c, sternite V (dorsal view); d, sterniteVI (dorsal view); e, sterniteVII (dorsal view); fj, A. mimetes: f, dorsal view of abdomen; g, sternite IV (dorsal view); h, sternite V (dorsal view); i, sternite VI (dorsal view); j, sternite VII (dorsal view). ps: primary sclerotization, se: secondary sclerotization. Blue arrows indicate pseudoappendages." figureDoi="http://doi.org/10.5281/zenodo.8281829" httpUri="https://zenodo.org/record/8281829/files/figure.png" pageId="6" pageNumber="430">4</figureCitation>
,);
<emphasis id="B96DEAEA3C71FFDAAAA8FEFAFAADFEA8" bold="true" box="[1187,1315,258,285]" pageId="6" pageNumber="430">sternite VI</emphasis>
strap-like, with a great portion of distal secondary sclerotization displaying sinuous margin; pleural region of segment VI slightly inflated and with cluster of short bristles (
<figureCitation id="13222A7D3C71FFDAACCAFEB3FCBFFED0" box="[705,817,331,357]" captionStart="FIGURE 1" captionStartId="2.[152,255,1513,1537]" captionTargetBox="[151,1435,237,1489]" captionTargetId="figure-56@2.[151,1435,237,1489]" captionTargetPageId="2" captionText="FIGURE 1. Austrospirachtha carrijoi sp. nov., female, holotype: a, habitus lateral; b, habitus dorsal. About 2.4 mm from the curvature of the abdomen to the apex of mandibula." figureDoi="http://doi.org/10.5281/zenodo.8281819" httpUri="https://zenodo.org/record/8281819/files/figure.png" pageId="6" pageNumber="430">Figs. 1a,b</figureCitation>
,
<figureCitation id="13222A7D3C71FFDAAD36FEB3FCF8FED0" box="[829,886,331,357]" captionStart="FIGURE 4" captionStartId="5.[152,255,1904,1928]" captionTargetBox="[226,1361,191,1873]" captionTargetId="figure-37@5.[216,1372,181,1880]" captionTargetPageId="5" captionText="FIGURE 4. Austrospirachtha carrijoi sp. nov., and A. mimetes illustrations. ae, Austrospirachtha carrijoi sp. nov.: a, dorsal view of abdomen; b, sternite IV (dorsal view); c, sternite V (dorsal view); d, sterniteVI (dorsal view); e, sterniteVII (dorsal view); fj, A. mimetes: f, dorsal view of abdomen; g, sternite IV (dorsal view); h, sternite V (dorsal view); i, sternite VI (dorsal view); j, sternite VII (dorsal view). ps: primary sclerotization, se: secondary sclerotization. Blue arrows indicate pseudoappendages." figureDoi="http://doi.org/10.5281/zenodo.8281829" httpUri="https://zenodo.org/record/8281829/files/figure.png" pageId="6" pageNumber="430">4a, d</figureCitation>
).
<emphasis id="B96DEAEA3C71FFDAAD82FEB2FB96FED0" bold="true" box="[905,1048,330,357]" pageId="6" pageNumber="430">Sternite VII</emphasis>
with primary sclerotized area with proximal margin round, and distally disposed secondary sclerotization with one medial acute process (
<figureCitation id="13222A7D3C71FFDAAB1AFE97FAE0FE3C" box="[1297,1390,367,393]" captionStart="FIGURE 1" captionStartId="2.[152,255,1513,1537]" captionTargetBox="[151,1435,237,1489]" captionTargetId="figure-56@2.[151,1435,237,1489]" captionTargetPageId="2" captionText="FIGURE 1. Austrospirachtha carrijoi sp. nov., female, holotype: a, habitus lateral; b, habitus dorsal. About 2.4 mm from the curvature of the abdomen to the apex of mandibula." figureDoi="http://doi.org/10.5281/zenodo.8281819" httpUri="https://zenodo.org/record/8281819/files/figure.png" pageId="6" pageNumber="430">Figs. 1b</figureCitation>
,
<figureCitation id="13222A7D3C71FFDAAB70FE97FF28FE18" captionStart="FIGURE 4" captionStartId="5.[152,255,1904,1928]" captionTargetBox="[226,1361,191,1873]" captionTargetId="figure-37@5.[216,1372,181,1880]" captionTargetPageId="5" captionText="FIGURE 4. Austrospirachtha carrijoi sp. nov., and A. mimetes illustrations. ae, Austrospirachtha carrijoi sp. nov.: a, dorsal view of abdomen; b, sternite IV (dorsal view); c, sternite V (dorsal view); d, sterniteVI (dorsal view); e, sterniteVII (dorsal view); fj, A. mimetes: f, dorsal view of abdomen; g, sternite IV (dorsal view); h, sternite V (dorsal view); i, sternite VI (dorsal view); j, sternite VII (dorsal view). ps: primary sclerotization, se: secondary sclerotization. Blue arrows indicate pseudoappendages." figureDoi="http://doi.org/10.5281/zenodo.8281829" httpUri="https://zenodo.org/record/8281829/files/figure.png" pageId="6" pageNumber="430">4a, e</figureCitation>
).
<emphasis id="B96DEAEA3C71FFDAAEB0FE6AFEC5FE18" bold="true" box="[187,331,402,429]" pageId="6" pageNumber="430">Tergite VIII</emphasis>
with long apodemes and secondary sclerotization between them, forming overall one conspicuous plate; base subquadrate with one long main bristle on row A and one on row P, but all surface covered with midsized to rather long bristles (
<figureCitation id="13222A7D3C71FFDAAF91FE23FD81FE40" box="[410,527,475,501]" captionStart="FIGURE 3" captionStartId="4.[152,255,1650,1674]" captionTargetBox="[160,1426,181,1625]" captionTargetId="figure-29@4.[160,1426,181,1625]" captionTargetPageId="4" captionText="FIGURE 3. Austrospirachtha carrijoi sp. nov., female, paratype: a, head ventral; b, mandibles (dorsal view); c, labrum (dorsal view); d, maxilla (dorsal view); e, meso-metathorax and first tergite attached to metanotum (seen through transparence as the meso-metanotum); f, elytron (dorsal view); g, elytron, close-up (dorsal view), details with arrows pointing to the three main bristles; h, tergite VIII (dorsal view); i, tergite VII (dorsal view), distal region, detail of chaetotaxy; j, sternite VIII (dorsal view); k, tergites IX and X (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.8281827" httpUri="https://zenodo.org/record/8281827/files/figure.png" pageId="6" pageNumber="430">Figs. 3h, i</figureCitation>
).
<emphasis id="B96DEAEA3C71FFDAAC2DFE22FD4CFE40" bold="true" box="[550,706,474,501]" pageId="6" pageNumber="430">Sternite VIII</emphasis>
subtrapezoidal, distal margin narrower than proximal; two pairs or long bristles horizontally arranged little below the halfline of piece (
<figureCitation id="13222A7D3C71FFDAADB4FE07FB83FDAC" box="[959,1037,511,537]" captionStart="FIGURE 3" captionStartId="4.[152,255,1650,1674]" captionTargetBox="[160,1426,181,1625]" captionTargetId="figure-29@4.[160,1426,181,1625]" captionTargetPageId="4" captionText="FIGURE 3. Austrospirachtha carrijoi sp. nov., female, paratype: a, head ventral; b, mandibles (dorsal view); c, labrum (dorsal view); d, maxilla (dorsal view); e, meso-metathorax and first tergite attached to metanotum (seen through transparence as the meso-metanotum); f, elytron (dorsal view); g, elytron, close-up (dorsal view), details with arrows pointing to the three main bristles; h, tergite VIII (dorsal view); i, tergite VII (dorsal view), distal region, detail of chaetotaxy; j, sternite VIII (dorsal view); k, tergites IX and X (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.8281827" httpUri="https://zenodo.org/record/8281827/files/figure.png" pageId="6" pageNumber="430">Fig. 3j</figureCitation>
).
<emphasis id="B96DEAEA3C71FFDAAA28FE06FB75FDAC" bold="true" box="[1059,1275,510,537]" pageId="6" pageNumber="430">Tergites X and IX</emphasis>
very compact together (
<figureCitation id="13222A7D3C71FFDAAF0AFDDBFED4FD88" box="[257,346,547,573]" captionStart="FIGURE 3" captionStartId="4.[152,255,1650,1674]" captionTargetBox="[160,1426,181,1625]" captionTargetId="figure-29@4.[160,1426,181,1625]" captionTargetPageId="4" captionText="FIGURE 3. Austrospirachtha carrijoi sp. nov., female, paratype: a, head ventral; b, mandibles (dorsal view); c, labrum (dorsal view); d, maxilla (dorsal view); e, meso-metathorax and first tergite attached to metanotum (seen through transparence as the meso-metanotum); f, elytron (dorsal view); g, elytron, close-up (dorsal view), details with arrows pointing to the three main bristles; h, tergite VIII (dorsal view); i, tergite VII (dorsal view), distal region, detail of chaetotaxy; j, sternite VIII (dorsal view); k, tergites IX and X (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.8281827" httpUri="https://zenodo.org/record/8281827/files/figure.png" pageId="6" pageNumber="430">Fig. 3K</figureCitation>
); tergite X subtriangular;
<emphasis id="B96DEAEA3C71FFDAAC73FDDAFD79FD88" bold="true" box="[632,759,546,573]" pageId="6" pageNumber="430">sternite IX</emphasis>
at least in female seems absent or completely fused to lateral margins of tergite IX.
<emphasis id="B96DEAEA3C71FFDAAF9BFDBEFDA3FDD5" bold="true" box="[400,557,582,608]" pageId="6" pageNumber="430">Spermatheca</emphasis>
indistinct.
</paragraph>
</subSubSection>
<caption id="DF6666703C74FFDFAE93F86FFE6BF866" ID-DOI="http://doi.org/10.5281/zenodo.8281823" ID-Zenodo-Dep="8281823" httpUri="https://zenodo.org/record/8281823/files/figure.png" pageId="3" pageNumber="427" startId="3.[152,255,1943,1967]" targetBox="[156,1431,181,1919]" targetPageId="3" targetType="figure">
<paragraph id="8BA636F83C74FFDFAE93F86FFE6BF866" blockId="3.[151,1437,1943,2003]" pageId="3" pageNumber="427">
<emphasis id="B96DEAEA3C74FFDFAE93F86FFE96F81A" bold="true" box="[152,280,1943,1967]" pageId="3" pageNumber="427">FIGURE 2.</emphasis>
<taxonomicName id="4C194D7B3C74FFDFAF14F86FFDACF81A" authority="Zilberman &amp; Pires-Silva, 2023" authorityName="Zilberman &amp; Pires-Silva" authorityYear="2023" box="[287,546,1943,1967]" class="Insecta" family="Staphylinidae" genus="Austrospirachtha" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="427" phylum="Arthropoda" rank="species" species="carrijoi" status="sp. nov.">
<emphasis id="B96DEAEA3C74FFDFAF14F86FFDACF81A" box="[287,546,1943,1967]" italics="true" pageId="3" pageNumber="427">Austrospirachtha carrijoi</emphasis>
</taxonomicName>
<emphasis id="B96DEAEA3C74FFDFAC23F86FFDF7F81A" bold="true" box="[552,633,1943,1967]" pageId="3" pageNumber="427">
<taxonomicNameLabel id="A25E57913C74FFDFAC23F86FFDF7F81A" box="[552,633,1943,1967]" pageId="3" pageNumber="427" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female, paratype:
<emphasis id="B96DEAEA3C74FFDFAD36F86FFCC4F81A" bold="true" box="[829,842,1943,1967]" pageId="3" pageNumber="427">a</emphasis>
, tergite II and sternite II “ring” modification by secondary sclerotization;
<emphasis id="B96DEAEA3C74FFDFAF26F843FEB5F866" bold="true" box="[301,315,1979,2003]" pageId="3" pageNumber="427">b</emphasis>
, habitus ventral.
</paragraph>
</caption>
<caption id="DF6666703C73FFD8AE93F98AFE7BF8AF" ID-DOI="http://doi.org/10.5281/zenodo.8281827" ID-Zenodo-Dep="8281827" httpUri="https://zenodo.org/record/8281827/files/figure.png" pageId="4" pageNumber="428" startId="4.[152,255,1650,1674]" targetBox="[160,1426,181,1625]" targetPageId="4" targetType="figure">
<paragraph id="8BA636F83C73FFD8AE93F98AFE7BF8AF" blockId="4.[151,1437,1650,1818]" pageId="4" pageNumber="428">
<emphasis id="B96DEAEA3C73FFD8AE93F98AFE96F93F" bold="true" box="[152,280,1650,1674]" pageId="4" pageNumber="428">FIGURE 3.</emphasis>
<taxonomicName id="4C194D7B3C73FFD8AF16F98AFD91F93F" authority="Zilberman &amp; Pires-Silva, 2023" authorityName="Zilberman &amp; Pires-Silva" authorityYear="2023" box="[285,543,1650,1674]" class="Insecta" family="Staphylinidae" genus="Austrospirachtha" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="428" phylum="Arthropoda" rank="species" species="carrijoi" status="sp. nov.">
<emphasis id="B96DEAEA3C73FFD8AF16F98AFD91F93F" box="[285,543,1650,1674]" italics="true" pageId="4" pageNumber="428">Austrospirachtha carrijoi</emphasis>
</taxonomicName>
<emphasis id="B96DEAEA3C73FFD8AC2EF98AFDFBF93F" bold="true" box="[549,629,1650,1674]" pageId="4" pageNumber="428">
<taxonomicNameLabel id="A25E57913C73FFD8AC2EF98AFDFBF93F" box="[549,629,1650,1674]" pageId="4" pageNumber="428" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female, paratype:
<emphasis id="B96DEAEA3C73FFD8AD3DF98AFCCDF93F" bold="true" box="[822,835,1650,1674]" pageId="4" pageNumber="428">a</emphasis>
, head ventral;
<emphasis id="B96DEAEA3C73FFD8ADDCF98AFC6BF93F" bold="true" box="[983,997,1650,1674]" pageId="4" pageNumber="428">b</emphasis>
, mandibles (dorsal view);
<emphasis id="B96DEAEA3C73FFD8AAFAF98AFB72F93F" bold="true" box="[1265,1276,1650,1674]" pageId="4" pageNumber="428">c</emphasis>
, labrum (dorsal view);
<emphasis id="B96DEAEA3C73FFD8AED4F96EFF63F91B" bold="true" box="[223,237,1686,1710]" pageId="4" pageNumber="428">d</emphasis>
, maxilla (dorsal view);
<emphasis id="B96DEAEA3C73FFD8AFEFF96EFE61F91B" bold="true" box="[484,495,1686,1710]" pageId="4" pageNumber="428">e</emphasis>
, meso-metathorax and first tergite attached to metanotum (seen through transparence as the meso-metanotum);
<emphasis id="B96DEAEA3C73FFD8AF54F942FEE9F967" bold="true" box="[351,359,1722,1746]" pageId="4" pageNumber="428">f</emphasis>
, elytron (dorsal view);
<emphasis id="B96DEAEA3C73FFD8AC56F942FDE4F967" bold="true" box="[605,618,1722,1746]" pageId="4" pageNumber="428">g</emphasis>
, elytron, close-up (dorsal view), details with arrows pointing to the three main bristles;
<emphasis id="B96DEAEA3C73FFD8AEE6F926FF75F943" bold="true" box="[237,251,1758,1782]" pageId="4" pageNumber="428">h</emphasis>
, tergite VIII (dorsal view);
<emphasis id="B96DEAEA3C73FFD8AC04F926FD98F943" bold="true" box="[527,534,1758,1782]" pageId="4" pageNumber="428">i</emphasis>
, tergite VII (dorsal view), distal region, detail of chaetotaxy;
<emphasis id="B96DEAEA3C73FFD8AA70F926FB0DF943" bold="true" box="[1147,1155,1758,1782]" pageId="4" pageNumber="428">j</emphasis>
, sternite VIII (dorsal view);
<emphasis id="B96DEAEA3C73FFD8AE9CF8FAFF2BF8AF" bold="true" box="[151,165,1794,1818]" pageId="4" pageNumber="428">k</emphasis>
, tergites IX and X (dorsal view).
</paragraph>
</caption>
<caption id="DF6666703C72FFD9AE93F888FA13F841" ID-DOI="http://doi.org/10.5281/zenodo.8281829" ID-Zenodo-Dep="8281829" httpUri="https://zenodo.org/record/8281829/files/figure.png" pageId="5" pageNumber="429" startId="5.[152,255,1904,1928]" targetBox="[226,1361,191,1873]" targetPageId="5" targetType="figure">
<paragraph id="8BA636F83C72FFD9AE93F888FA13F841" blockId="5.[151,1437,1904,2036]" pageId="5" pageNumber="429">
<emphasis id="B96DEAEA3C72FFD9AE93F888FE96F83D" bold="true" box="[152,280,1904,1928]" pageId="5" pageNumber="429">FIGURE 4.</emphasis>
<taxonomicName id="4C194D7B3C72FFD9AF14F888FDAFF83D" authority="Zilberman &amp; Pires-Silva, 2023" authorityName="Zilberman &amp; Pires-Silva" authorityYear="2023" box="[287,545,1904,1928]" class="Insecta" family="Staphylinidae" genus="Austrospirachtha" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="429" phylum="Arthropoda" rank="species" species="carrijoi" status="sp. nov.">
<emphasis id="B96DEAEA3C72FFD9AF14F888FDAFF83D" box="[287,545,1904,1928]" italics="true" pageId="5" pageNumber="429">Austrospirachtha carrijoi</emphasis>
</taxonomicName>
<emphasis id="B96DEAEA3C72FFD9AC23F888FDF7F83D" bold="true" box="[552,633,1904,1928]" pageId="5" pageNumber="429">
<taxonomicNameLabel id="A25E57913C72FFD9AC23F888FDF7F83D" box="[552,633,1904,1928]" pageId="5" pageNumber="429" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="4C194D7B3C72FFD9ACBAF888FCAFF83D" authorityName="Watson" authorityYear="1973" box="[689,801,1904,1928]" class="Insecta" family="Staphylinidae" genus="Austrospirachtha" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="429" phylum="Arthropoda" rank="species" species="mimetes">
<emphasis id="B96DEAEA3C72FFD9ACBAF888FCAFF83D" box="[689,801,1904,1928]" italics="true" pageId="5" pageNumber="429">A. mimetes</emphasis>
</taxonomicName>
illustrations.
<emphasis id="B96DEAEA3C72FFD9ADA0F888FC5EF83D" bold="true" box="[939,976,1904,1928]" pageId="5" pageNumber="429">ae</emphasis>
,
<taxonomicName id="4C194D7B3C72FFD9ADD6F888FB51F83D" authority="Zilberman &amp; Pires-Silva, 2023" authorityName="Zilberman &amp; Pires-Silva" authorityYear="2023" box="[989,1247,1904,1928]" class="Insecta" family="Staphylinidae" genus="Austrospirachtha" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="429" phylum="Arthropoda" rank="species" species="carrijoi" status="sp. nov.">
<emphasis id="B96DEAEA3C72FFD9ADD6F888FB51F83D" box="[989,1247,1904,1928]" italics="true" pageId="5" pageNumber="429">Austrospirachtha carrijoi</emphasis>
</taxonomicName>
<emphasis id="B96DEAEA3C72FFD9AAEDF888FAB9F83D" bold="true" box="[1254,1335,1904,1928]" pageId="5" pageNumber="429">
<taxonomicNameLabel id="A25E57913C72FFD9AAEDF888FAB9F83D" box="[1254,1335,1904,1928]" pageId="5" pageNumber="429" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
:
<emphasis id="B96DEAEA3C72FFD9AB4EF888FADCF83D" bold="true" box="[1349,1362,1904,1928]" pageId="5" pageNumber="429">a</emphasis>
, dorsal view of abdomen;
<emphasis id="B96DEAEA3C72FFD9AF45F86CFED2F819" bold="true" box="[334,348,1940,1964]" pageId="5" pageNumber="429">b</emphasis>
, sternite IV (dorsal view);
<emphasis id="B96DEAEA3C72FFD9AC6EF86CFDFEF819" bold="true" box="[613,624,1940,1964]" pageId="5" pageNumber="429">c</emphasis>
, sternite V (dorsal view);
<emphasis id="B96DEAEA3C72FFD9AD64F86CFCF3F819" bold="true" box="[879,893,1940,1964]" pageId="5" pageNumber="429">d</emphasis>
, sternite VI (dorsal view);
<emphasis id="B96DEAEA3C72FFD9AA8EF86CFB1EF819" bold="true" box="[1157,1168,1940,1964]" pageId="5" pageNumber="429">e</emphasis>
, sternite VII (dorsal view);
<emphasis id="B96DEAEA3C72FFD9AE9CF840FF3AF865" bold="true" box="[151,180,1976,2000]" pageId="5" pageNumber="429">fj</emphasis>
,
<taxonomicName id="4C194D7B3C72FFD9AECAF840FEBEF865" authorityName="Watson" authorityYear="1973" box="[193,304,1976,2000]" class="Insecta" family="Staphylinidae" genus="Austrospirachtha" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="429" phylum="Arthropoda" rank="species" species="mimetes">
<emphasis id="B96DEAEA3C72FFD9AECAF840FEBEF865" box="[193,304,1976,2000]" italics="true" pageId="5" pageNumber="429">A. mimetes</emphasis>
</taxonomicName>
:
<emphasis id="B96DEAEA3C72FFD9AF35F840FEC8F865" bold="true" box="[318,326,1976,2000]" pageId="5" pageNumber="429">f</emphasis>
, dorsal view of abdomen;
<emphasis id="B96DEAEA3C72FFD9AC5FF840FDEFF865" bold="true" box="[596,609,1976,2000]" pageId="5" pageNumber="429">g</emphasis>
, sternite IV (dorsal view);
<emphasis id="B96DEAEA3C72FFD9AD78F840FC0FF865" bold="true" box="[883,897,1976,2000]" pageId="5" pageNumber="429">h</emphasis>
, sternite V (dorsal view);
<emphasis id="B96DEAEA3C72FFD9AA81F840FB1FF865" bold="true" box="[1162,1169,1976,2000]" pageId="5" pageNumber="429">i</emphasis>
, sternite VI (dorsal view);
<emphasis id="B96DEAEA3C72FFD9AE9CF824FF11F841" bold="true" box="[151,159,2012,2036]" pageId="5" pageNumber="429">j</emphasis>
, sternite VII (dorsal view).
<emphasis id="B96DEAEA3C72FFD9AFCBF825FE59F841" box="[448,471,2013,2036]" italics="true" pageId="5" pageNumber="429">ps</emphasis>
: primary sclerotization,
<emphasis id="B96DEAEA3C72FFD9ACD8F825FD66F841" box="[723,744,2013,2036]" italics="true" pageId="5" pageNumber="429">se</emphasis>
: secondary sclerotization. Blue arrows indicate pseudoappendages.
</paragraph>
</caption>
<subSubSection id="C30365733C71FFDAAECCFD93FCBAFD1C" pageId="6" pageNumber="430" type="biology_ecology">
<paragraph id="8BA636F83C71FFDAAECCFD93FCBAFD1C" blockId="6.[151,1437,150,1257]" pageId="6" pageNumber="430">
<emphasis id="B96DEAEA3C71FFDAAECCFD93FE1BFD31" bold="true" box="[199,405,618,645]" pageId="6" pageNumber="430">Host relationship</emphasis>
. Two specimens were collected in a small (~
<quantity id="4CE19B1D3C71FFDAAD8CFD93FC41FD30" box="[903,975,619,645]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="430" unit="cm" value="20.0">20 cm</quantity>
) eroded mound nest, with
<taxonomicName id="4C194D7B3C71FFDAAAFCFD93FF39FD1C" class="Insecta" family="Termitidae" genus="Australitermes" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="430" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B96DEAEA3C71FFDAAAFCFD93FA12FD30" box="[1271,1436,619,645]" italics="true" pageId="6" pageNumber="430">Australitermes</emphasis>
sp.
</taxonomicName>
(
<figureCitation id="13222A7D3C71FFDAAECDFD77FEFFFD1C" box="[198,369,655,681]" captionStart="FIGURE 6" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[200,1387,181,1919]" captionTargetId="figure-37@7.[200,1387,181,1919]" captionTargetPageId="7" captionText="FIGURE 6. Termite species from the nest from which Austrospirachtha carrijoi sp. nov. individuals were collected. Australitermes sp.: a, lateral view; c, dorsal view of the head. Nasutitermes sp.: b, lateral view; d, dorsal view of the head." figureDoi="http://doi.org/10.5281/zenodo.8281835" httpUri="https://zenodo.org/record/8281835/files/figure.png" pageId="6" pageNumber="430">Figs. 6A and C</figureCitation>
) and
<taxonomicName id="4C194D7B3C71FFDAAFA4FD77FDE4FD1C" box="[431,618,655,681]" class="Insecta" family="Termitidae" genus="Nasutitermes" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="430" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B96DEAEA3C71FFDAAFA4FD77FDCDFD1D" box="[431,579,655,680]" italics="true" pageId="6" pageNumber="430">Nasutitermes</emphasis>
sp.
</taxonomicName>
(
<figureCitation id="13222A7D3C71FFDAAC72FD77FCA6FD1C" box="[633,808,655,681]" captionStart="FIGURE 6" captionStartId="7.[152,255,1943,1967]" captionTargetBox="[200,1387,181,1919]" captionTargetId="figure-37@7.[200,1387,181,1919]" captionTargetPageId="7" captionText="FIGURE 6. Termite species from the nest from which Austrospirachtha carrijoi sp. nov. individuals were collected. Australitermes sp.: a, lateral view; c, dorsal view of the head. Nasutitermes sp.: b, lateral view; d, dorsal view of the head." figureDoi="http://doi.org/10.5281/zenodo.8281835" httpUri="https://zenodo.org/record/8281835/files/figure.png" pageId="6" pageNumber="430">Figs. 6B and D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30365733C71FFDAAECCFD4BFEB3FD44" pageId="6" pageNumber="430" type="etymology">
<paragraph id="8BA636F83C71FFDAAECCFD4BFEB3FD44" blockId="6.[151,1437,150,1257]" pageId="6" pageNumber="430">
<emphasis id="B96DEAEA3C71FFDAAECCFD4BFEC6FD78" bold="true" box="[199,328,691,717]" pageId="6" pageNumber="430">Etymology</emphasis>
. The specific name is dedicated to Dr. Tiago F. Carrijo, the termitologist and colleague who collected the specimens.
</paragraph>
</subSubSection>
<subSubSection id="C30365733C71FFDAAECCFD03FB75FCE8" pageId="6" pageNumber="430" type="materials_examined">
<paragraph id="8BA636F83C71FFDAAECCFD03FB75FCE8" blockId="6.[151,1437,150,1257]" pageId="6" pageNumber="430">
<emphasis id="B96DEAEA3C71FFDAAECCFD03FEE3FCA1" bold="true" box="[199,365,762,789]" pageId="6" pageNumber="430">Type material</emphasis>
.
<materialsCitation id="3B713CA53C71FFDAAF71FD03FD81FCE8" collectingDate="2014-07-08" collectingDateMax="2014-07-10" collectingDateMin="2014-07-08" collectionCode="MZSP" collectorName="T. Carrijo" country="Australia" latitude="-13.04759" location="Litchfield National Park" longLatPrecision="5" longitude="130.9106" municipality="Litchfield" pageId="6" pageNumber="430" specimenCode="MZSP 21193" specimenCount="1" stateProvince="Northern Territory" typeStatus="holotype">
<typeStatus id="54A2885A3C71FFDAAF71FD03FE6CFCA0" box="[378,482,763,789]" pageId="6" pageNumber="430">Holotype</typeStatus>
(#F).
<emphasis id="B96DEAEA3C71FFDAAC2EFD02FD1BFCA1" bold="true" box="[549,661,762,788]" pageId="6" pageNumber="430">
<collectingCountry id="F30E76683C71FFDAAC2EFD02FD1BFCA1" box="[549,661,762,788]" name="Australia" pageId="6" pageNumber="430">Australia</collectingCountry>
</emphasis>
:
<collectingRegion id="49DDF81A3C71FFDAACA9FD03FCFBFCA0" box="[674,885,763,789]" country="Australia" name="Northern Territory" pageId="6" pageNumber="430">Northern Territory</collectingRegion>
:
<collectingMunicipality id="6BC2AC823C71FFDAAD76FD03FC60FCA0" box="[893,1006,763,789]" pageId="6" pageNumber="430">Litchfield</collectingMunicipality>
,
<location id="8EC660233C71FFDAADF2FD03FA87FCA0" LSID="urn:lsid:plazi:treatment:03B087EE3C76FFD4AE9CFC51FDC2FDD4:8EC660233C71FFDAADF2FD03FA87FCA0" box="[1017,1289,763,789]" country="Australia" latitude="-13.04759" longLatPrecision="5" longitude="130.9106" municipality="Litchfield" name="Litchfield National Park" pageId="6" pageNumber="430" stateProvince="Northern Territory">Litchfield National Park</location>
, -
<geoCoordinate id="EE2D503F3C71FFDAAB10FD02FA16FCA0" box="[1307,1432,762,789]" degrees="13.04759" direction="south" orientation="latitude" pageId="6" pageNumber="430" precision="1" value="-13.04759">13.04759S</geoCoordinate>
,
<geoCoordinate id="EE2D503F3C71FFDAAE9CFCE7FE9AFC8C" box="[151,276,799,825]" degrees="130.9106" direction="east" orientation="longitude" pageId="6" pageNumber="430" precision="5" value="130.9106">130.9106E</geoCoordinate>
,
<date id="FFA710383C71FFDAAF14FCE7FE4DFC8C" box="[287,451,799,825]" pageId="6" pageNumber="430" value="2014-07-08" valueMax="2014-07-10" valueMin="2014-07-08">
<collectingDate id="EFE3E9D03C71FFDAAF14FCE7FE4DFC8C" box="[287,451,799,825]" pageId="6" pageNumber="430" value="2014-07-08" valueMax="2014-07-10" valueMin="2014-07-08">8-10/VII/2014</collectingDate>
</date>
,
<collectorName id="26EC532E3C71FFDAAFC6FCE7FDB6FC8C" box="[461,568,799,825]" pageId="6" pageNumber="430">T. Carrijo</collectorName>
col., with
<taxonomicName id="4C194D7B3C71FFDAACA5FCE7FCF7FC8C" box="[686,889,799,825]" class="Insecta" family="Termitidae" genus="Australitermes" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="430" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B96DEAEA3C71FFDAACA5FCE7FCDDFC8C" box="[686,851,799,825]" italics="true" pageId="6" pageNumber="430">Australitermes</emphasis>
sp.
</taxonomicName>
(MZUSP 23728), and
<taxonomicName id="4C194D7B3C71FFDAAA7DFCE7FABEFC8C" box="[1142,1328,799,825]" class="Insecta" family="Termitidae" genus="Nasutitermes" kingdom="Animalia" order="Blattodea" pageId="6" pageNumber="430" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B96DEAEA3C71FFDAAA7DFCE7FA84FC8D" box="[1142,1290,799,824]" italics="true" pageId="6" pageNumber="430">Nasutitermes</emphasis>
sp.
</taxonomicName>
(MZUSP 26578) beetle code
<specimenCode id="DBBF9E833C71FFDAAF78FCBBFD85FCE8" box="[371,523,835,861]" pageId="6" pageNumber="430">MZSP 21193</specimenCode>
.
</materialsCitation>
<materialsCitation id="3B713CA53C71FFDAAC1EFCBBFB75FCE8" box="[533,1275,835,861]" pageId="6" pageNumber="430" specimenCount="1" typeStatus="paratype">
<typeStatus id="54A2885A3C71FFDAAC1EFCBBFDF9FCE8" box="[533,631,835,861]" pageId="6" pageNumber="430">Paratype</typeStatus>
(#F). 1, same data and collected along with the holotype.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30365733C71FFD4AECCFC9EFDC2FDD4" lastPageId="8" lastPageNumber="432" pageId="6" pageNumber="430" type="discussion">
<paragraph id="8BA636F83C71FFDAAECCFC9EFDB3FB5C" blockId="6.[151,1437,150,1257]" pageId="6" pageNumber="430">
<emphasis id="B96DEAEA3C71FFDAAECCFC9EFE5AFC34" bold="true" box="[199,468,870,897]" pageId="6" pageNumber="430">Comparative remarks</emphasis>
.
<taxonomicName id="4C194D7B3C71FFDAAFEDFC9FFC87FC35" authority="Zilberman &amp; Pires-Silva, 2023" authorityName="Zilberman &amp; Pires-Silva" authorityYear="2023" box="[486,777,871,897]" class="Insecta" family="Staphylinidae" genus="Austrospirachtha" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="430" phylum="Arthropoda" rank="species" species="carrijoi" status="sp. nov.">
<emphasis id="B96DEAEA3C71FFDAAFEDFC9FFC87FC35" box="[486,777,871,897]" italics="true" pageId="6" pageNumber="430">Austrospirachtha carrijoi</emphasis>
</taxonomicName>
<emphasis id="B96DEAEA3C71FFDAAD18FC9FFCFFFC34" bold="true" box="[787,881,871,897]" pageId="6" pageNumber="430">
<taxonomicNameLabel id="A25E57913C71FFDAAD18FC9FFCFFFC34" box="[787,881,871,897]" pageId="6" pageNumber="430" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar in overall appearance to
<taxonomicName id="4C194D7B3C71FFDAAB1DFC9FFA1BFC35" authorityName="Watson" authorityYear="1973" box="[1302,1429,871,897]" class="Insecta" family="Staphylinidae" genus="Austrospirachtha" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="430" phylum="Arthropoda" rank="species" species="mimetes">
<emphasis id="B96DEAEA3C71FFDAAB1DFC9FFA1BFC35" box="[1302,1429,871,897]" italics="true" pageId="6" pageNumber="430">A. mimetes</emphasis>
</taxonomicName>
. However, there are several distinct differences between the two species. These include the shape of the abdomen, secondary sclerotization on sternites, shape of the last pseudoappendages (
<figureCitation id="13222A7D3C71FFDAADDEFC57FB97FC7C" box="[981,1049,943,969]" captionStart="FIGURE 4" captionStartId="5.[152,255,1904,1928]" captionTargetBox="[226,1361,191,1873]" captionTargetId="figure-37@5.[216,1372,181,1880]" captionTargetPageId="5" captionText="FIGURE 4. Austrospirachtha carrijoi sp. nov., and A. mimetes illustrations. ae, Austrospirachtha carrijoi sp. nov.: a, dorsal view of abdomen; b, sternite IV (dorsal view); c, sternite V (dorsal view); d, sterniteVI (dorsal view); e, sterniteVII (dorsal view); fj, A. mimetes: f, dorsal view of abdomen; g, sternite IV (dorsal view); h, sternite V (dorsal view); i, sternite VI (dorsal view); j, sternite VII (dorsal view). ps: primary sclerotization, se: secondary sclerotization. Blue arrows indicate pseudoappendages." figureDoi="http://doi.org/10.5281/zenodo.8281829" httpUri="https://zenodo.org/record/8281829/files/figure.png" pageId="6" pageNumber="430">Fig. 4</figureCitation>
), and a less pronounced difference in the maxilla (with
<taxonomicName id="4C194D7B3C71FFDAAF8FFC2BFD8CFC59" authorityName="Watson" authorityYear="1973" box="[388,514,979,1005]" class="Insecta" family="Staphylinidae" genus="Austrospirachtha" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="430" phylum="Arthropoda" rank="species" species="mimetes">
<emphasis id="B96DEAEA3C71FFDAAF8FFC2BFE1BFC58" box="[388,405,979,1005]" italics="true" pageId="6" pageNumber="430">A</emphasis>
.
<emphasis id="B96DEAEA3C71FFDAAFADFC2BFD8CFC59" box="[422,514,979,1004]" italics="true" pageId="6" pageNumber="430">mimetes</emphasis>
</taxonomicName>
in parentheses): The first maxillary palpomere is distinct in
<taxonomicName id="4C194D7B3C71FFDAAAB4FC2BFAB9FC59" authority="Zilberman &amp; Pires-Silva, 2023" authorityName="Zilberman &amp; Pires-Silva" authorityYear="2023" box="[1215,1335,979,1005]" class="Insecta" family="Staphylinidae" genus="Austrospirachtha" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="430" phylum="Arthropoda" rank="species" species="carrijoi" status="sp. nov.">
<emphasis id="B96DEAEA3C71FFDAAAB4FC2BFB5EFC58" box="[1215,1232,979,1005]" italics="true" pageId="6" pageNumber="430">A</emphasis>
.
<emphasis id="B96DEAEA3C71FFDAAAEAFC2BFAB9FC59" box="[1249,1335,979,1004]" italics="true" pageId="6" pageNumber="430">carrijoi</emphasis>
</taxonomicName>
<emphasis id="B96DEAEA3C71FFDAAB4BFC2BFA12FC58" bold="true" box="[1344,1436,979,1005]" pageId="6" pageNumber="430">
<taxonomicNameLabel id="A25E57913C71FFDAAB4BFC2BFA12FC58" box="[1344,1436,979,1005]" pageId="6" pageNumber="430" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(Indistinct in
<taxonomicName id="4C194D7B3C71FFDAAF3AFC0FFE20FBA5" authorityName="Watson" authorityYear="1973" box="[305,430,1015,1041]" class="Insecta" family="Staphylinidae" genus="Austrospirachtha" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="430" phylum="Arthropoda" rank="species" species="mimetes">
<emphasis id="B96DEAEA3C71FFDAAF3AFC0FFECCFBA4" box="[305,322,1015,1041]" italics="true" pageId="6" pageNumber="430">A</emphasis>
.
<emphasis id="B96DEAEA3C71FFDAAF59FC0FFE20FBA5" box="[338,430,1015,1040]" italics="true" pageId="6" pageNumber="430">mimetes</emphasis>
</taxonomicName>
, see
<bibRefCitation id="EF884B093C71FFDAAFE2FC0FFD0EFBA4" author="Watson, J. A. L." box="[489,640,1015,1041]" pageId="6" pageNumber="430" pagination="307 - 310" refId="ref3125" refString="Watson, J. A. L. (1973) Austrospirachtha mimetes, A New Termitophilous Corotocine From Northern Australia (Coleoptera: Staphylinidae), Journal of the Australian Entomological Society, 12, 307 - 310. https: // doi. org / 10.1111 / j. 1440 - 6055.1973. tb 01678. x" type="journal article" year="1973">Watson 1973</bibRefCitation>
, fig. 5); the modified, egg-like part of the abdomen is longer than the subsequent segments combined in
<taxonomicName id="4C194D7B3C71FFDAAC28FBE3FD17FB81" authority="Zilberman &amp; Pires-Silva, 2023" authorityName="Zilberman &amp; Pires-Silva" authorityYear="2023" box="[547,665,1051,1077]" class="Insecta" family="Staphylinidae" genus="Austrospirachtha" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="430" phylum="Arthropoda" rank="species" species="carrijoi" status="sp. nov.">
<emphasis id="B96DEAEA3C71FFDAAC28FBE3FDBAFB80" box="[547,564,1051,1077]" italics="true" pageId="6" pageNumber="430">A</emphasis>
.
<emphasis id="B96DEAEA3C71FFDAAC48FBE3FD17FB81" box="[579,665,1051,1076]" italics="true" pageId="6" pageNumber="430">carrijoi</emphasis>
</taxonomicName>
<emphasis id="B96DEAEA3C71FFDAACA9FBE3FD73FB80" bold="true" box="[674,765,1051,1077]" pageId="6" pageNumber="430">
<taxonomicNameLabel id="A25E57913C71FFDAACA9FBE3FD73FB80" box="[674,765,1051,1077]" pageId="6" pageNumber="430" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13222A7D3C71FFDAAD05FBE3FCE9FB80" box="[782,871,1051,1077]" captionStart="FIGURE 4" captionStartId="5.[152,255,1904,1928]" captionTargetBox="[226,1361,191,1873]" captionTargetId="figure-37@5.[216,1372,181,1880]" captionTargetPageId="5" captionText="FIGURE 4. Austrospirachtha carrijoi sp. nov., and A. mimetes illustrations. ae, Austrospirachtha carrijoi sp. nov.: a, dorsal view of abdomen; b, sternite IV (dorsal view); c, sternite V (dorsal view); d, sterniteVI (dorsal view); e, sterniteVII (dorsal view); fj, A. mimetes: f, dorsal view of abdomen; g, sternite IV (dorsal view); h, sternite V (dorsal view); i, sternite VI (dorsal view); j, sternite VII (dorsal view). ps: primary sclerotization, se: secondary sclerotization. Blue arrows indicate pseudoappendages." figureDoi="http://doi.org/10.5281/zenodo.8281829" httpUri="https://zenodo.org/record/8281829/files/figure.png" pageId="6" pageNumber="430">Fig. 4A</figureCitation>
) (shorter in
<taxonomicName id="4C194D7B3C71FFDAADFDFBE3FBFDFB81" authorityName="Watson" authorityYear="1973" box="[1014,1139,1051,1077]" class="Insecta" family="Staphylinidae" genus="Austrospirachtha" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="430" phylum="Arthropoda" rank="species" species="mimetes">
<emphasis id="B96DEAEA3C71FFDAADFDFBE3FB89FB80" box="[1014,1031,1051,1077]" italics="true" pageId="6" pageNumber="430">A</emphasis>
.
<emphasis id="B96DEAEA3C71FFDAAA1CFBE3FBFDFB81" box="[1047,1139,1051,1076]" italics="true" pageId="6" pageNumber="430">mimetes</emphasis>
</taxonomicName>
,
<figureCitation id="13222A7D3C71FFDAAA89FBE3FB57FB80" box="[1154,1241,1051,1077]" captionStart="FIGURE 4" captionStartId="5.[152,255,1904,1928]" captionTargetBox="[226,1361,191,1873]" captionTargetId="figure-37@5.[216,1372,181,1880]" captionTargetPageId="5" captionText="FIGURE 4. Austrospirachtha carrijoi sp. nov., and A. mimetes illustrations. ae, Austrospirachtha carrijoi sp. nov.: a, dorsal view of abdomen; b, sternite IV (dorsal view); c, sternite V (dorsal view); d, sterniteVI (dorsal view); e, sterniteVII (dorsal view); fj, A. mimetes: f, dorsal view of abdomen; g, sternite IV (dorsal view); h, sternite V (dorsal view); i, sternite VI (dorsal view); j, sternite VII (dorsal view). ps: primary sclerotization, se: secondary sclerotization. Blue arrows indicate pseudoappendages." figureDoi="http://doi.org/10.5281/zenodo.8281829" httpUri="https://zenodo.org/record/8281829/files/figure.png" pageId="6" pageNumber="430">Fig. 4F</figureCitation>
); sternite
<collectionCode id="ED08AE3D3C71FFDAAB42FBE3FAD3FB80" box="[1353,1373,1051,1077]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="430" type="Museum">V</collectionCode>
in
<taxonomicName id="4C194D7B3C71FFDAAB8FFBE3FF63FBED" authority="Zilberman &amp; Pires-Silva, 2023" authorityName="Zilberman &amp; Pires-Silva" authorityYear="2023" class="Insecta" family="Staphylinidae" genus="Austrospirachtha" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="430" phylum="Arthropoda" rank="species" species="carrijoi" status="sp. nov.">
<emphasis id="B96DEAEA3C71FFDAAB8FFBE3FA1BFB80" box="[1412,1429,1051,1077]" italics="true" pageId="6" pageNumber="430">A</emphasis>
.
<emphasis id="B96DEAEA3C71FFDAAE9CFBC7FF63FBED" box="[151,237,1087,1112]" italics="true" pageId="6" pageNumber="430">carrijoi</emphasis>
</taxonomicName>
<emphasis id="B96DEAEA3C71FFDAAEFEFBC7FEDEFBEC" bold="true" box="[245,336,1087,1113]" pageId="6" pageNumber="430">
<taxonomicNameLabel id="A25E57913C71FFDAAEFEFBC7FEDEFBEC" box="[245,336,1087,1113]" pageId="6" pageNumber="430" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a thinly distal area of secondary sclerotization in addition to the lateral ones (
<figureCitation id="13222A7D3C71FFDAAAF2FBC7FADEFBEC" box="[1273,1360,1087,1113]" captionStart="FIGURE 4" captionStartId="5.[152,255,1904,1928]" captionTargetBox="[226,1361,191,1873]" captionTargetId="figure-37@5.[216,1372,181,1880]" captionTargetPageId="5" captionText="FIGURE 4. Austrospirachtha carrijoi sp. nov., and A. mimetes illustrations. ae, Austrospirachtha carrijoi sp. nov.: a, dorsal view of abdomen; b, sternite IV (dorsal view); c, sternite V (dorsal view); d, sterniteVI (dorsal view); e, sterniteVII (dorsal view); fj, A. mimetes: f, dorsal view of abdomen; g, sternite IV (dorsal view); h, sternite V (dorsal view); i, sternite VI (dorsal view); j, sternite VII (dorsal view). ps: primary sclerotization, se: secondary sclerotization. Blue arrows indicate pseudoappendages." figureDoi="http://doi.org/10.5281/zenodo.8281829" httpUri="https://zenodo.org/record/8281829/files/figure.png" pageId="6" pageNumber="430">Fig. 4C</figureCitation>
) (only lateral areas have secondary sclerotization in
<taxonomicName id="4C194D7B3C71FFDAAC96FB9BFC94FBC9" authorityName="Watson" authorityYear="1973" box="[669,794,1123,1149]" class="Insecta" family="Staphylinidae" genus="Austrospirachtha" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="430" phylum="Arthropoda" rank="species" species="mimetes">
<emphasis id="B96DEAEA3C71FFDAAC96FB9BFD20FBC8" box="[669,686,1123,1149]" italics="true" pageId="6" pageNumber="430">A</emphasis>
.
<emphasis id="B96DEAEA3C71FFDAACB5FB9BFC94FBC9" box="[702,794,1123,1148]" italics="true" pageId="6" pageNumber="430">mimetes</emphasis>
</taxonomicName>
,
<figureCitation id="13222A7D3C71FFDAAD21FB9BFC0BFBC8" box="[810,901,1123,1149]" captionStart="FIGURE 4" captionStartId="5.[152,255,1904,1928]" captionTargetBox="[226,1361,191,1873]" captionTargetId="figure-37@5.[216,1372,181,1880]" captionTargetPageId="5" captionText="FIGURE 4. Austrospirachtha carrijoi sp. nov., and A. mimetes illustrations. ae, Austrospirachtha carrijoi sp. nov.: a, dorsal view of abdomen; b, sternite IV (dorsal view); c, sternite V (dorsal view); d, sterniteVI (dorsal view); e, sterniteVII (dorsal view); fj, A. mimetes: f, dorsal view of abdomen; g, sternite IV (dorsal view); h, sternite V (dorsal view); i, sternite VI (dorsal view); j, sternite VII (dorsal view). ps: primary sclerotization, se: secondary sclerotization. Blue arrows indicate pseudoappendages." figureDoi="http://doi.org/10.5281/zenodo.8281829" httpUri="https://zenodo.org/record/8281829/files/figure.png" pageId="6" pageNumber="430">Fig. 4H</figureCitation>
); sternite
<collectionCode id="ED08AE3D3C71FFDAADFEFB9BFB9DFBC8" box="[1013,1043,1123,1149]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="6" pageNumber="430">VI</collectionCode>
in
<taxonomicName id="4C194D7B3C71FFDAAA31FB9BFB3FFBC9" authority="Zilberman &amp; Pires-Silva, 2023" authorityName="Zilberman &amp; Pires-Silva" authorityYear="2023" box="[1082,1201,1123,1149]" class="Insecta" family="Staphylinidae" genus="Austrospirachtha" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="430" phylum="Arthropoda" rank="species" species="carrijoi" status="sp. nov.">
<emphasis id="B96DEAEA3C71FFDAAA31FB9BFBC5FBC8" box="[1082,1099,1123,1149]" italics="true" pageId="6" pageNumber="430">A</emphasis>
.
<emphasis id="B96DEAEA3C71FFDAAA50FB9BFB3FFBC9" box="[1115,1201,1123,1148]" italics="true" pageId="6" pageNumber="430">carrijoi</emphasis>
</taxonomicName>
<emphasis id="B96DEAEA3C71FFDAAAB2FB9BFA9AFBC8" bold="true" box="[1209,1300,1123,1149]" pageId="6" pageNumber="430">
<taxonomicNameLabel id="A25E57913C71FFDAAAB2FB9BFA9AFBC8" box="[1209,1300,1123,1149]" pageId="6" pageNumber="430" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a distal secondary sclerotization with a sinuous margin (
<figureCitation id="13222A7D3C71FFDAACBBFB7FFC89FB14" box="[688,775,1159,1185]" captionStart="FIGURE 4" captionStartId="5.[152,255,1904,1928]" captionTargetBox="[226,1361,191,1873]" captionTargetId="figure-37@5.[216,1372,181,1880]" captionTargetPageId="5" captionText="FIGURE 4. Austrospirachtha carrijoi sp. nov., and A. mimetes illustrations. ae, Austrospirachtha carrijoi sp. nov.: a, dorsal view of abdomen; b, sternite IV (dorsal view); c, sternite V (dorsal view); d, sterniteVI (dorsal view); e, sterniteVII (dorsal view); fj, A. mimetes: f, dorsal view of abdomen; g, sternite IV (dorsal view); h, sternite V (dorsal view); i, sternite VI (dorsal view); j, sternite VII (dorsal view). ps: primary sclerotization, se: secondary sclerotization. Blue arrows indicate pseudoappendages." figureDoi="http://doi.org/10.5281/zenodo.8281829" httpUri="https://zenodo.org/record/8281829/files/figure.png" pageId="6" pageNumber="430">Fig. 4D</figureCitation>
) (straight margin in
<taxonomicName id="4C194D7B3C71FFDAADE1FB7FFBEAFB15" authorityName="Watson" authorityYear="1973" box="[1002,1124,1159,1185]" class="Insecta" family="Staphylinidae" genus="Austrospirachtha" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="430" phylum="Arthropoda" rank="species" species="mimetes">
<emphasis id="B96DEAEA3C71FFDAADE1FB7FFC75FB14" box="[1002,1019,1159,1185]" italics="true" pageId="6" pageNumber="430">A</emphasis>
.
<emphasis id="B96DEAEA3C71FFDAAA03FB7FFBEAFB15" box="[1032,1124,1159,1184]" italics="true" pageId="6" pageNumber="430">mimetes</emphasis>
</taxonomicName>
,
<figureCitation id="13222A7D3C71FFDAAA7AFB7FFB30FB14" box="[1137,1214,1159,1185]" captionStart="FIGURE 4" captionStartId="5.[152,255,1904,1928]" captionTargetBox="[226,1361,191,1873]" captionTargetId="figure-37@5.[216,1372,181,1880]" captionTargetPageId="5" captionText="FIGURE 4. Austrospirachtha carrijoi sp. nov., and A. mimetes illustrations. ae, Austrospirachtha carrijoi sp. nov.: a, dorsal view of abdomen; b, sternite IV (dorsal view); c, sternite V (dorsal view); d, sterniteVI (dorsal view); e, sterniteVII (dorsal view); fj, A. mimetes: f, dorsal view of abdomen; g, sternite IV (dorsal view); h, sternite V (dorsal view); i, sternite VI (dorsal view); j, sternite VII (dorsal view). ps: primary sclerotization, se: secondary sclerotization. Blue arrows indicate pseudoappendages." figureDoi="http://doi.org/10.5281/zenodo.8281829" httpUri="https://zenodo.org/record/8281829/files/figure.png" pageId="6" pageNumber="430">Fig. 4I</figureCitation>
); and sternite VII in
<taxonomicName id="4C194D7B3C71FFDAAE9CFB53FE83FB71" authority="Zilberman &amp; Pires-Silva, 2023" authorityName="Zilberman &amp; Pires-Silva" authorityYear="2023" box="[151,269,1195,1221]" class="Insecta" family="Staphylinidae" genus="Austrospirachtha" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="430" phylum="Arthropoda" rank="species" species="carrijoi" status="sp. nov.">
<emphasis id="B96DEAEA3C71FFDAAE9CFB53FF26FB70" box="[151,168,1195,1221]" italics="true" pageId="6" pageNumber="430">A</emphasis>
.
<emphasis id="B96DEAEA3C71FFDAAEBCFB53FE83FB71" box="[183,269,1195,1220]" italics="true" pageId="6" pageNumber="430">carrijoi</emphasis>
</taxonomicName>
<emphasis id="B96DEAEA3C71FFDAAF1FFB53FEE0FB70" bold="true" box="[276,366,1195,1221]" pageId="6" pageNumber="430">
<taxonomicNameLabel id="A25E57913C71FFDAAF1FFB53FEE0FB70" box="[276,366,1195,1221]" pageId="6" pageNumber="430" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has a distally disposed secondary sclerotization with a medial acute process (
<figureCitation id="13222A7D3C71FFDAAAD2FB53FAA0FB70" box="[1241,1326,1195,1221]" captionStart="FIGURE 4" captionStartId="5.[152,255,1904,1928]" captionTargetBox="[226,1361,191,1873]" captionTargetId="figure-37@5.[216,1372,181,1880]" captionTargetPageId="5" captionText="FIGURE 4. Austrospirachtha carrijoi sp. nov., and A. mimetes illustrations. ae, Austrospirachtha carrijoi sp. nov.: a, dorsal view of abdomen; b, sternite IV (dorsal view); c, sternite V (dorsal view); d, sterniteVI (dorsal view); e, sterniteVII (dorsal view); fj, A. mimetes: f, dorsal view of abdomen; g, sternite IV (dorsal view); h, sternite V (dorsal view); i, sternite VI (dorsal view); j, sternite VII (dorsal view). ps: primary sclerotization, se: secondary sclerotization. Blue arrows indicate pseudoappendages." figureDoi="http://doi.org/10.5281/zenodo.8281829" httpUri="https://zenodo.org/record/8281829/files/figure.png" pageId="6" pageNumber="430">Fig. 4E</figureCitation>
) (without acute process in
<taxonomicName id="4C194D7B3C71FFDAAF5AFB37FE42FB5D" authorityName="Watson" authorityYear="1973" box="[337,460,1231,1257]" class="Insecta" family="Staphylinidae" genus="Austrospirachtha" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="430" phylum="Arthropoda" rank="species" species="mimetes">
<emphasis id="B96DEAEA3C71FFDAAF5AFB37FEECFB5C" box="[337,354,1231,1257]" italics="true" pageId="6" pageNumber="430">A</emphasis>
.
<emphasis id="B96DEAEA3C71FFDAAF7BFB37FE42FB5D" box="[368,460,1231,1256]" italics="true" pageId="6" pageNumber="430">mimetes</emphasis>
</taxonomicName>
,
<figureCitation id="13222A7D3C71FFDAAFD1FB37FDA9FB5C" box="[474,551,1231,1257]" captionStart="FIGURE 4" captionStartId="5.[152,255,1904,1928]" captionTargetBox="[226,1361,191,1873]" captionTargetId="figure-37@5.[216,1372,181,1880]" captionTargetPageId="5" captionText="FIGURE 4. Austrospirachtha carrijoi sp. nov., and A. mimetes illustrations. ae, Austrospirachtha carrijoi sp. nov.: a, dorsal view of abdomen; b, sternite IV (dorsal view); c, sternite V (dorsal view); d, sterniteVI (dorsal view); e, sterniteVII (dorsal view); fj, A. mimetes: f, dorsal view of abdomen; g, sternite IV (dorsal view); h, sternite V (dorsal view); i, sternite VI (dorsal view); j, sternite VII (dorsal view). ps: primary sclerotization, se: secondary sclerotization. Blue arrows indicate pseudoappendages." figureDoi="http://doi.org/10.5281/zenodo.8281829" httpUri="https://zenodo.org/record/8281829/files/figure.png" pageId="6" pageNumber="430">Fig. 4J</figureCitation>
).
</paragraph>
<caption id="DF6666703C71FFDAAE93F820FB18F845" ID-DOI="http://doi.org/10.5281/zenodo.8281833" ID-Zenodo-Dep="8281833" box="[152,1174,2008,2032]" httpUri="https://zenodo.org/record/8281833/files/figure.png" pageId="6" pageNumber="430" startId="6.[152,255,2008,2032]" targetBox="[221,1366,1287,1980]" targetPageId="6" targetType="figure">
<paragraph id="8BA636F83C71FFDAAE93F820FB18F845" blockId="6.[152,1174,2008,2032]" box="[152,1174,2008,2032]" pageId="6" pageNumber="430">
<emphasis id="B96DEAEA3C71FFDAAE93F820FE96F845" bold="true" box="[152,280,2008,2032]" pageId="6" pageNumber="430">FIGURE 5.</emphasis>
Chaetotaxy of mentum-submentum.
<emphasis id="B96DEAEA3C71FFDAAC99F820FD11F845" bold="true" box="[658,671,2008,2032]" pageId="6" pageNumber="430">a</emphasis>
,
<taxonomicName id="4C194D7B3C71FFDAACA7F820FC2FF845" authority="Zilberman &amp; Pires-Silva, 2023" authorityName="Zilberman &amp; Pires-Silva" authorityYear="2023" box="[684,929,2008,2032]" class="Insecta" family="Staphylinidae" genus="Austrospirachta" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="430" phylum="Arthropoda" rank="species" species="carrijoi" status="sp. nov.">
<emphasis id="B96DEAEA3C71FFDAACA7F820FC2FF845" box="[684,929,2008,2032]" italics="true" pageId="6" pageNumber="430">Austrospirachta carrijoi</emphasis>
</taxonomicName>
<emphasis id="B96DEAEA3C71FFDAADA3F820FC76F845" bold="true" box="[936,1016,2008,2032]" pageId="6" pageNumber="430">
<taxonomicNameLabel id="A25E57913C71FFDAADA3F820FC76F845" box="[936,1016,2008,2032]" pageId="6" pageNumber="430" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="B96DEAEA3C71FFDAAA0DF820FB9AF845" bold="true" box="[1030,1044,2008,2032]" pageId="6" pageNumber="430">b</emphasis>
,
<taxonomicName id="4C194D7B3C71FFDAAA2AF820FB1EF845" authorityName="Watson" authorityYear="1973" box="[1057,1168,2008,2032]" class="Insecta" family="Staphylinidae" genus="Austrospirachtha" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="430" phylum="Arthropoda" rank="species" species="mimetes">
<emphasis id="B96DEAEA3C71FFDAAA2AF820FB1EF845" box="[1057,1168,2008,2032]" italics="true" pageId="6" pageNumber="430">A. mimetes</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF6666703C70FFDBAE93F86FFAE1F861" ID-DOI="http://doi.org/10.5281/zenodo.8281835" ID-Zenodo-Dep="8281835" httpUri="https://zenodo.org/record/8281835/files/figure.png" pageId="7" pageNumber="431" startId="7.[152,255,1943,1967]" targetBox="[200,1387,181,1919]" targetPageId="7" targetType="figure">
<paragraph id="8BA636F83C70FFDBAE93F86FFAE1F861" blockId="7.[151,1437,1943,2004]" pageId="7" pageNumber="431">
<emphasis id="B96DEAEA3C70FFDBAE93F86FFE91F81A" bold="true" box="[152,287,1943,1967]" pageId="7" pageNumber="431">FIGURE 6.</emphasis>
Termite species from the nest from which
<taxonomicName id="4C194D7B3C70FFDBAD0EF860FB80F81A" authority="Zilberman &amp; Pires-Silva, 2023" authorityName="Zilberman &amp; Pires-Silva" authorityYear="2023" box="[773,1038,1944,1968]" class="Insecta" family="Staphylinidae" genus="Austrospirachtha" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="431" phylum="Arthropoda" rank="species" species="carrijoi" status="sp. nov.">
<emphasis id="B96DEAEA3C70FFDBAD0EF860FB80F81A" box="[773,1038,1944,1968]" italics="true" pageId="7" pageNumber="431">Austrospirachtha carrijoi</emphasis>
</taxonomicName>
<emphasis id="B96DEAEA3C70FFDBAA10F860FBFCF805" bold="true" box="[1051,1138,1944,1968]" pageId="7" pageNumber="431">
<taxonomicNameLabel id="A25E57913C70FFDBAA10F860FBFCF805" box="[1051,1138,1944,1968]" pageId="7" pageNumber="431" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
individuals were collected.
<taxonomicName id="4C194D7B3C70FFDBAE9CF844FEDDF866" box="[151,339,1980,2004]" class="Insecta" family="Termitidae" genus="Australitermes" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="431" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B96DEAEA3C70FFDBAE9CF844FEA2F861" box="[151,300,1980,2004]" italics="true" pageId="7" pageNumber="431">Australitermes</emphasis>
sp.
</taxonomicName>
:
<emphasis id="B96DEAEA3C70FFDBAF56F844FEE4F861" bold="true" box="[349,362,1980,2004]" pageId="7" pageNumber="431">a</emphasis>
, lateral view;
<emphasis id="B96DEAEA3C70FFDBAFF1F844FD8BF861" bold="true" box="[506,517,1980,2004]" pageId="7" pageNumber="431">c</emphasis>
, dorsal view of the head.
<taxonomicName id="4C194D7B3C70FFDBAD00F844FC36F866" box="[779,952,1980,2003]" class="Insecta" family="Termitidae" genus="Nasutitermes" kingdom="Animalia" order="Blattodea" pageId="7" pageNumber="431" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B96DEAEA3C70FFDBAD00F844FC1FF866" box="[779,913,1980,2003]" italics="true" pageId="7" pageNumber="431">Nasutitermes</emphasis>
sp.
</taxonomicName>
:
<emphasis id="B96DEAEA3C70FFDBADC9F844FC5EF861" bold="true" box="[962,976,1980,2004]" pageId="7" pageNumber="431">b</emphasis>
, lateral view;
<emphasis id="B96DEAEA3C70FFDBAA6AF844FBE1F861" bold="true" box="[1121,1135,1980,2004]" pageId="7" pageNumber="431">d</emphasis>
, dorsal view of the head.
</paragraph>
</caption>
<paragraph id="8BA636F83C7FFFD4AECCFF6FFC15FE40" blockId="8.[151,1437,151,609]" pageId="8" pageNumber="432">
Upon reading
<bibRefCitation id="EF884B093C7FFFD4AF61FF6FFDADFF04" author="Watson, J. A. L." box="[362,547,151,177]" pageId="8" pageNumber="432" pagination="307 - 310" refId="ref3125" refString="Watson, J. A. L. (1973) Austrospirachtha mimetes, A New Termitophilous Corotocine From Northern Australia (Coleoptera: Staphylinidae), Journal of the Australian Entomological Society, 12, 307 - 310. https: // doi. org / 10.1111 / j. 1440 - 6055.1973. tb 01678. x" type="journal article" year="1973">Watsons (1973)</bibRefCitation>
work and examining and interpreting this new species, it is worth noting that there is some confusion regarding what constitutes sternite III. This confusion is understandable, as the modifications to the membranous area and secondary sclerotization can disrupt the typical morphology of these beetles. Our main disagreement with Watson pertains to the modified, inflated, egg-like part of the abdomen, where we consider the sclerotized plate to be a modification of sternite III, whereas Watson views it simply as secondary sclerotization of the intersegmental membrane (
<figureCitation id="13222A7D3C7FFFD4AFE5FEB3FDB4FED0" box="[494,570,331,357]" captionStart="FIGURE 1" captionStartId="2.[152,255,1513,1537]" captionTargetBox="[151,1435,237,1489]" captionTargetId="figure-56@2.[151,1435,237,1489]" captionTargetPageId="2" captionText="FIGURE 1. Austrospirachtha carrijoi sp. nov., female, holotype: a, habitus lateral; b, habitus dorsal. About 2.4 mm from the curvature of the abdomen to the apex of mandibula." figureDoi="http://doi.org/10.5281/zenodo.8281819" httpUri="https://zenodo.org/record/8281819/files/figure.png" pageId="8" pageNumber="432">Figs. 1</figureCitation>
,
<figureCitation id="13222A7D3C7FFFD4AC4CFEB3FDD9FED0" box="[583,599,331,357]" captionStart="FIGURE 2" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[156,1431,181,1919]" captionTargetId="figure-37@3.[156,1431,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Austrospirachtha carrijoi sp. nov., female, paratype: a, tergite II and sternite II “ring” modification by secondary sclerotization; b, habitus ventral." figureDoi="http://doi.org/10.5281/zenodo.8281823" httpUri="https://zenodo.org/record/8281823/files/figure.png" pageId="8" pageNumber="432">2</figureCitation>
,
<figureCitation id="13222A7D3C7FFFD4AC69FEB3FD18FED0" box="[610,662,331,357]" captionStart="FIGURE 4" captionStartId="5.[152,255,1904,1928]" captionTargetBox="[226,1361,191,1873]" captionTargetId="figure-37@5.[216,1372,181,1880]" captionTargetPageId="5" captionText="FIGURE 4. Austrospirachtha carrijoi sp. nov., and A. mimetes illustrations. ae, Austrospirachtha carrijoi sp. nov.: a, dorsal view of abdomen; b, sternite IV (dorsal view); c, sternite V (dorsal view); d, sterniteVI (dorsal view); e, sterniteVII (dorsal view); fj, A. mimetes: f, dorsal view of abdomen; g, sternite IV (dorsal view); h, sternite V (dorsal view); i, sternite VI (dorsal view); j, sternite VII (dorsal view). ps: primary sclerotization, se: secondary sclerotization. Blue arrows indicate pseudoappendages." figureDoi="http://doi.org/10.5281/zenodo.8281829" httpUri="https://zenodo.org/record/8281829/files/figure.png" pageId="8" pageNumber="432">4a, f</figureCitation>
). What Watson identifies as the true third sternite, we regard as part of segment IV, with its secondary sclerotization (
<figureCitation id="13222A7D3C7FFFD4AC97FE97FD76FE3C" box="[668,760,367,393]" captionStart="FIGURE 1" captionStartId="2.[152,255,1513,1537]" captionTargetBox="[151,1435,237,1489]" captionTargetId="figure-56@2.[151,1435,237,1489]" captionTargetPageId="2" captionText="FIGURE 1. Austrospirachtha carrijoi sp. nov., female, holotype: a, habitus lateral; b, habitus dorsal. About 2.4 mm from the curvature of the abdomen to the apex of mandibula." figureDoi="http://doi.org/10.5281/zenodo.8281819" httpUri="https://zenodo.org/record/8281819/files/figure.png" pageId="8" pageNumber="432">Figs. 1b</figureCitation>
,
<figureCitation id="13222A7D3C7FFFD4AD0EFE97FCFFFE3C" box="[773,881,367,393]" captionStart="FIGURE 4" captionStartId="5.[152,255,1904,1928]" captionTargetBox="[226,1361,191,1873]" captionTargetId="figure-37@5.[216,1372,181,1880]" captionTargetPageId="5" captionText="FIGURE 4. Austrospirachtha carrijoi sp. nov., and A. mimetes illustrations. ae, Austrospirachtha carrijoi sp. nov.: a, dorsal view of abdomen; b, sternite IV (dorsal view); c, sternite V (dorsal view); d, sterniteVI (dorsal view); e, sterniteVII (dorsal view); fj, A. mimetes: f, dorsal view of abdomen; g, sternite IV (dorsal view); h, sternite V (dorsal view); i, sternite VI (dorsal view); j, sternite VII (dorsal view). ps: primary sclerotization, se: secondary sclerotization. Blue arrows indicate pseudoappendages." figureDoi="http://doi.org/10.5281/zenodo.8281829" httpUri="https://zenodo.org/record/8281829/files/figure.png" pageId="8" pageNumber="432">4a, b, f, g</figureCitation>
). We could not find evidence to support Watsons claim, as there is no clear segmental delimitation between these two pieces, we consider it belong to segment IV. Furthermore, while it is not a strict rule, in
<taxonomicName id="4C194D7B3C7FFFD4ACB7FE4FFCB9FE64" box="[700,823,439,465]" class="Insecta" family="Staphylinidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="432" phylum="Arthropoda" rank="tribe" tribe="Corotocini">Corotocini</taxonomicName>
, secondary sclerotization usually arises from primary sclerotized pieces rather than membranes far from the main sclerites.
</paragraph>
<paragraph id="8BA636F83C7FFFD4AECCFE07FDC2FDD4" blockId="8.[151,1437,151,609]" pageId="8" pageNumber="432">
As for Watsons observations, it is noteworthy that we also havent found the labial palpi, the labium being overall extremely reduced; but in
<bibRefCitation id="EF884B093C7FFFD4AC19FDDBFC87FD88" author="Jacobson, H. R. &amp; Kistner, D. H. &amp; Pasteels, J. M." box="[530,777,547,573]" pageId="8" pageNumber="432" pagination="1 - 245" refId="ref3045" refString="Jacobson, H. R., Kistner, D. H. &amp; Pasteels, J. M. (1986) Generic revision, phylogenetic classification, and phylogeny of the termitophilous tribe Corotocini (Coleoptera: Staphylinidae). Sociobiology, 12, 1 - 245." type="journal article" year="1986">
Jacobson
<emphasis id="B96DEAEA3C7FFFD4AC8BFDDBFD3FFD88" box="[640,689,547,573]" italics="true" pageId="8" pageNumber="432">et al</emphasis>
. (1986)
</bibRefCitation>
the authors stated at least for
<taxonomicName id="4C194D7B3C7FFFD4AA52FDDBFB5AFD89" authorityName="Watson" authorityYear="1973" box="[1113,1236,547,573]" class="Insecta" family="Staphylinidae" genus="Austrospirachtha" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="432" phylum="Arthropoda" rank="species" species="mimetes">
<emphasis id="B96DEAEA3C7FFFD4AA52FDDBFBE4FD88" box="[1113,1130,547,573]" italics="true" pageId="8" pageNumber="432">A</emphasis>
.
<emphasis id="B96DEAEA3C7FFFD4AA73FDDBFB5AFD89" box="[1144,1236,547,572]" italics="true" pageId="8" pageNumber="432">mimetes</emphasis>
</taxonomicName>
that, even though it is small, the labial palpi is 3-articled.
</paragraph>
</subSubSection>
</treatment>
</document>