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<document id="42C2DA0AED38C29C2771BE33078B2313" ID-DOI="10.11646/zootaxa.2944.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5283420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629731250194" checkinUser="felipe" docAuthor="Perkins, Philip D." docDate="2011" docId="03B087E55B5BFFACFF79F7C8FA39FB99" docLanguage="en" docName="zt02944p417.pdf" docOrigin="Zootaxa 2944 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hydraena nanocolorata Perkins 2011, new species" docType="treatment" docVersion="5" lastPageNumber="64" masterDocId="FF89FF9D5B65FF93FFEEF22DFFB6FF96" masterDocTitle="New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944" masterLastPageNumber="417" masterPageNumber="1" pageNumber="63" updateTime="1699103370621" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="E0C4B4D3C2A053FAAEE2395E6748D1B5">New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944</mods:title>
</mods:titleInfo>
<mods:name id="311FB9733AADBAC24B46B251DBEB17C6" type="personal">
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<mods:namePart id="B8D3AA95D8488F1B1915CF29E2F675EB">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="FD9118413892C8D0BBE073B153D08CBF">text</mods:typeOfResource>
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<mods:title id="E9A62ED7E68D5E0133E4633AC4C884C7">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="40384B16197FF93E30539B6EB29D9C6B">
<mods:date id="0863BE7E8FFC1C7DBC7B42D910259811">2011</mods:date>
<mods:detail id="2688CC194AF6F12BF54AC3239E0BD84F" type="pubDate">
<mods:number id="AD08283D9667F0C5B9D497C7EA2A0C5B">2011-07-08</mods:number>
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<mods:number id="76CFBD972C8BDA4B32BDCC99F318B2ED">2944</mods:number>
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<mods:number id="5BF20A1ED507AEFB20D3290D86427F6E">1</mods:number>
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<mods:url id="0024C7B26F13E666E79B2EB92E711C8B">https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1</mods:url>
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<mods:classification id="1492D329D6E8EF34F4A65DC28147BDBD">journal article</mods:classification>
<mods:identifier id="B9391E48D9064ED87C121DC47FB0B848" type="DOI">10.11646/zootaxa.2944.1.1</mods:identifier>
<mods:identifier id="FF05A638FFA0C06E852DFB9A68D1CC79" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="2F3FC9C913DE1094AE7E74469D1A19A1" type="Zenodo-Dep">5283420</mods:identifier>
</mods:mods>
<treatment id="03B087E55B5BFFACFF79F7C8FA39FB99" ID-DOI="http://doi.org/10.5281/zenodo.5291530" ID-Zenodo-Dep="5291530" LSID="urn:lsid:plazi:treatment:03B087E55B5BFFACFF79F7C8FA39FB99" httpUri="http://treatment.plazi.org/id/03B087E55B5BFFACFF79F7C8FA39FB99" lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63">
<subSubSection id="C30365785B5BFFADFF79F7C8FDE0FA69" box="[151,598,1509,1535]" pageId="62" pageNumber="63" type="nomenclature">
<paragraph id="8BA636F35B5BFFADFF79F7C8FDE0FA69" blockId="62.[151,598,1509,1567]" box="[151,598,1509,1535]" pageId="62" pageNumber="63">
<heading id="D0EE819F5B5BFFADFF79F7C8FDE0FA69" bold="true" box="[151,598,1509,1535]" fontSize="11" level="1" pageId="62" pageNumber="63" reason="1">
<emphasis id="B96DEAE15B5BFFADFF79F7C8FDE0FA69" bold="true" box="[151,598,1509,1535]" pageId="62" pageNumber="63">
<taxonomicName id="4C194D705B5BFFADFF79F7C8FE01FA69" ID-CoL="3N3DZ" authority="Perkins, 2011" authorityName="Perkins" authorityYear="2011" box="[151,439,1509,1535]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="nanocolorata" status="sp. nov.">
<emphasis id="B96DEAE15B5BFFADFF79F7C8FE01FA69" bold="true" box="[151,439,1509,1535]" italics="true" pageId="62" pageNumber="63">Hydraena nanocolorata</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25E579A5B5BFFADFE28F7C8FDE0FA69" box="[454,598,1509,1535]" pageId="62" pageNumber="63" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30365785B5BFFADFF79F42AFEC2F989" box="[151,372,1543,1567]" pageId="62" pageNumber="63" type="description">
<paragraph id="8BA636F35B5BFFADFF79F42AFEC2F989" blockId="62.[151,598,1509,1567]" box="[151,372,1543,1567]" pageId="62" pageNumber="63">
(
<figureCitation id="13222A765B5BFFADFF71F42AFF4BF989" box="[159,253,1543,1567]" captionStart="FIGURE 98" captionStartId="218.[151,250,1753,1776]" captionTargetBox="[244,1366,222,1732]" captionTargetId="figure-29@218.[198,1389,193,1732]" captionTargetPageId="218" captionText="FIGURE 98. Hydraena nanocolorata, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283603" httpUri="https://zenodo.org/record/5283603/files/figure.png" pageId="62" pageNumber="63">Figs. 98</figureCitation>
,
<figureCitation id="13222A765B5BFFADFEE6F42AFE82F989" box="[264,308,1543,1567]" captionStart="FIGURES 100–101" captionStartId="220.[151,264,1650,1673]" captionTargetBox="[309,1276,224,1579]" captionTargetId="figure-32@220.[264,1323,193,1629]" captionTargetPageId="220" captionText="FIGURES 100–101. Aedeagi of holotypes. 100. H. nanocolorata. 101. H. clarinis." figureDoi="http://doi.org/10.5281/zenodo.5283609" httpUri="https://zenodo.org/record/5283609/files/figure.png" pageId="62" pageNumber="63">100</figureCitation>
,
<figureCitation id="13222A765B5BFFADFEAEF42AFEDDF989" box="[320,363,1543,1567]" captionStart="FIGURES 475–478" captionStartId="394.[151,264,1650,1673]" captionTargetBox="[165,1421,259,1607]" captionTargetId="figure-23@394.[165,1421,193,1629]" captionTargetPageId="394" captionText="FIGURES 475–478. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284093" httpUri="https://zenodo.org/record/5284093/files/figure.png" pageId="62" pageNumber="63">475</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30365785B5BFFADFF79F47DFE61F88A" pageId="62" pageNumber="63" type="materials_examined">
<paragraph id="8BA636F35B5BFFADFF79F47DFE61F88A" blockId="62.[151,1436,1615,2036]" pageId="62" pageNumber="63">
Type Material.
<materialsCitation id="3B713CAE5B5BFFADFEBEF47DFAACF939" collectingDate="1988-07-01" collectingDateMax="1988-08-30" collectingDateMin="1988-07-01" collectionCode="NMW" collectorName="D. Dudgeon" elevation="160" latitude="-5.55" location="Koje Creek" longLatPrecision="1305" longitude="145.38333" municipality="Ramu Basin" pageId="62" pageNumber="63" specimenCount="1" stateProvince="Madang" typeStatus="holotype">
<typeStatus id="54A288515B5BFFADFEBEF47DFE0BF9FE" box="[336,445,1616,1640]" pageId="62" pageNumber="63" type="holotype">
<emphasis id="B96DEAE15B5BFFADFEBEF47DFE0BF9FE" bold="true" box="[336,445,1616,1640]" pageId="62" pageNumber="63">Holotype</emphasis>
</typeStatus>
(male):
<emphasis id="B96DEAE15B5BFFADFDCBF47DFCB5F9FE" bold="true" box="[549,771,1615,1640]" pageId="62" pageNumber="63">
<collectingRegion id="49DDF8115B5BFFADFDCBF47DFD48F9FE" box="[549,766,1615,1640]" country="Papua New Guinea" name="Madang" pageId="62" pageNumber="63">Madang Province</collectingRegion>
:
</emphasis>
<collectingRegion id="49DDF8115B5BFFADFCFAF462FCE5F9F1" box="[788,851,1615,1639]" country="Papua New Guinea" name="West Sepik" pageId="62" pageNumber="63">Sepik</collectingRegion>
<collectingMunicipality id="6BC2AC895B5BFFADFCB5F47DFC5FF9F1" box="[859,1001,1615,1639]" pageId="62" pageNumber="63">Ramu Basin</collectingMunicipality>
,
<location id="8EC660285B5BFFADFC1BF462FBCFF9F1" LSID="urn:lsid:plazi:treatment:03B087E55B5BFFACFF79F7C8FA39FB99:8EC660285B5BFFADFC1BF462FBCFF9F1" box="[1013,1145,1615,1639]" latitude="-5.55" longLatPrecision="1305" longitude="145.38333" municipality="Ramu Basin" name="Koje Creek" pageId="62" pageNumber="63" stateProvince="Madang">Kojé Creek</location>
, unshaded creek, small to medium cobbles, gravel and fine sand (the latter in areas of slack water), filamentous algae on stony substrates, small accumulations of leaf litter,
<quantity id="4CE19B165B5BFFADFDFBF4BAFDE8F939" box="[533,606,1687,1711]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="62" pageNumber="63" unit="m" value="160.0">
<elevation id="0034D1C05B5BFFADFDFBF4BAFDE8F939" box="[533,606,1687,1711]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="62" pageNumber="63" unit="m" value="160.0">160 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345B5BFFADFD85F4BAFD72F939" box="[619,708,1687,1711]" degrees="5" direction="south" minutes="33" orientation="latitude" pageId="62" pageNumber="63" precision="925" value="-5.55">5° 33' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345B5BFFADFD3FF4BAFCFFF939" box="[721,841,1687,1711]" degrees="145" direction="east" minutes="23" orientation="longitude" pageId="62" pageNumber="63" precision="925" value="145.38333">145° 23' E</geoCoordinate>
,
<date id="FFA710335B5BFFADFCBBF4BAFBA9F939" box="[853,1055,1687,1711]" pageId="62" pageNumber="63" value="1988-07-01" valueMax="1988-08-30" valueMin="1988-07-01">
<collectingDate id="EFE3E9DB5B5BFFADFCBBF4BAFBA9F939" box="[853,1055,1687,1711]" pageId="62" pageNumber="63" value="1988-07-01" valueMax="1988-08-30" valueMin="1988-07-01">1 vii–30 viii 1988</collectingDate>
</date>
,
<collectorName id="26EC53255B5BFFADFBC5F4B5FB02F939" box="[1067,1204,1687,1711]" pageId="62" pageNumber="63">D. Dudgeon</collectorName>
(
<collectionCode id="ED08AE365B5BFFADFB2AF4BAFAA4F939" box="[1220,1298,1687,1711]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="62" pageNumber="63">NMW</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B713CAE5B5BFFADFACAF4B5FCD1F942" collectingDate="1988-07-01" collectingDateMax="1988-08-30" collectingDateMin="1988-07-01" collectionCode="NMW" collectorName="D. Dudgeon" elevation="160" latitude="-5.55" location="Same" longLatPrecision="1305" longitude="145.38333" municipality="Ramu Basin" pageId="62" pageNumber="63" specimenCount="1" stateProvince="Madang" typeStatus="paratype">
<emphasis id="B96DEAE15B5BFFADFACAF4B5FA2AF926" bold="true" box="[1316,1436,1688,1712]" pageId="62" pageNumber="63">
<typeStatus id="54A288515B5BFFADFACAF4B5FA2AF926" box="[1316,1436,1688,1712]" pageId="62" pageNumber="63" type="paratype">Paratypes</typeStatus>
</emphasis>
(101):
<emphasis id="B96DEAE15B5BFFADFF02F491FE7CF942" bold="true" box="[236,458,1723,1748]" pageId="62" pageNumber="63">
<collectingRegion id="49DDF8115B5BFFADFF02F491FE73F942" box="[236,453,1723,1748]" country="Papua New Guinea" name="Madang" pageId="62" pageNumber="63">Madang Province</collectingRegion>
:
</emphasis>
<location id="8EC660285B5BFFADFE32F491FDACF942" LSID="urn:lsid:plazi:treatment:03B087E55B5BFFACFF79F7C8FA39FB99:8EC660285B5BFFADFE32F491FDACF942" box="[476,538,1724,1748]" latitude="-5.55" longLatPrecision="1305" longitude="145.38333" municipality="Ramu Basin" name="Same" pageId="62" pageNumber="63" stateProvince="Madang">Same</location>
data as holotype (84
<collectionCode id="ED08AE365B5BFFADFCFCF491FCE8F942" box="[786,862,1724,1748]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="62" pageNumber="63">NMW</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B713CAE5B5BFFADFC9AF491FE65F88A" collectingDate="1965-01-27" collectingDateMax="1965-01-30" collectingDateMin="1965-01-27" collectionCode="NHM" collectorName="M. E. Bacchus" elevation="305" latitude="-5.9833336" location="Gusap" longLatPrecision="1305" longitude="146.08333" municipality="Markham Valley" pageId="62" pageNumber="63" specimenCount="1" stateProvince="Morobe" typeStatus="paratype">
<emphasis id="B96DEAE15B5BFFADFC9AF491FBFBF942" bold="true" box="[884,1101,1723,1748]" pageId="62" pageNumber="63">
<collectingRegion id="49DDF8115B5BFFADFC9AF491FBFEF942" box="[884,1096,1723,1748]" country="Papua New Guinea" name="Morobe" pageId="62" pageNumber="63">Morobe Province</collectingRegion>
:
</emphasis>
<collectingMunicipality id="6BC2AC895B5BFFADFBB0F491FAABF942" box="[1118,1309,1724,1748]" pageId="62" pageNumber="63">Markham Valley</collectingMunicipality>
,
<location id="8EC660285B5BFFADFAC7F491FAC2F942" LSID="urn:lsid:plazi:treatment:03B087E55B5BFFACFF79F7C8FA39FB99:8EC660285B5BFFADFAC7F491FAC2F942" box="[1321,1396,1724,1748]" latitude="-5.9833336" longLatPrecision="1305" longitude="146.08333" municipality="Markham Valley" name="Gusap" pageId="62" pageNumber="63" stateProvince="Morobe">Gusap</location>
,
<locationDeviation id="9A142FD15B5BFFADFA6EF491FEEAF961" location="Lae" pageId="62" pageNumber="63">
<quantity id="4CE19B165B5BFFADFA6EF491FF62F961" metricMagnitude="5" metricUnit="m" metricValue="1.4484096" pageId="62" pageNumber="63" unit="mi" value="90.0">90 miles</quantity>
NW of Lae
</locationDeviation>
, ex dead leaves on mud wetted by seepage,
<quantity id="4CE19B165B5BFFADFCA5F4F2FC23F961" box="[843,917,1759,1783]" metricMagnitude="2" metricUnit="m" metricValue="3.05" pageId="62" pageNumber="63" unit="m" value="305.0">
<elevation id="0034D1C05B5BFFADFCA5F4F2FC23F961" box="[843,917,1759,1783]" metricMagnitude="2" metricUnit="m" metricValue="3.05" pageId="62" pageNumber="63" unit="m" value="305.0">305 m</elevation>
</quantity>
,
<geoCoordinate id="EE2D50345B5BFFADFC4FF4F2FC4DF961" box="[929,1019,1759,1783]" degrees="5" direction="south" minutes="59" orientation="latitude" pageId="62" pageNumber="63" precision="925" value="-5.9833336">5° 59' S</geoCoordinate>
,
<geoCoordinate id="EE2D50345B5BFFADFBE9F4F2FBC4F961" box="[1031,1138,1759,1783]" degrees="146" direction="east" minutes="5" orientation="longitude" pageId="62" pageNumber="63" precision="925" value="146.08333">146° 5' E</geoCoordinate>
,
<date id="FFA710335B5BFFADFB90F4F2FAA2F961" box="[1150,1300,1759,1783]" pageId="62" pageNumber="63" value="1965-01-27" valueMax="1965-01-30" valueMin="1965-01-27">
<collectingDate id="EFE3E9DB5B5BFFADFB90F4F2FAA2F961" box="[1150,1300,1759,1783]" pageId="62" pageNumber="63" value="1965-01-27" valueMax="1965-01-30" valueMin="1965-01-27">27–30 i 1965</collectingDate>
</date>
,
<collectorName id="26EC53255B5BFFADFACEF4CDFF7CF88A" pageId="62" pageNumber="63">M. E. Bacchus</collectorName>
(MEB 163) (17
<collectionCode id="ED08AE365B5BFFADFE6BF529FE7DF88A" box="[389,459,1796,1820]" pageId="62" pageNumber="63">NHM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30365785B5BFFADFF29F50AFE3EF862" pageId="62" pageNumber="63" type="diagnosis">
<paragraph id="8BA636F35B5BFFADFF29F50AFE3EF862" blockId="62.[151,1436,1615,2036]" pageId="62" pageNumber="63">
<emphasis id="B96DEAE15B5BFFADFF29F50AFE64F8D6" bold="true" box="[199,466,1831,1856]" pageId="62" pageNumber="63">Differential Diagnosis.</emphasis>
Among members of the
<emphasis id="B96DEAE15B5BFFADFD02F505FCE7F8D6" box="[748,849,1832,1856]" italics="true" pageId="62" pageNumber="63">Colorata</emphasis>
group similar to
<taxonomicName id="4C194D705B5BFFADFBFAF504FB20F8D6" authorityName="Perkins" authorityYear="2011" box="[1044,1174,1832,1856]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="colorata">
<emphasis id="B96DEAE15B5BFFADFBFAF504FB9EF8D6" box="[1044,1064,1833,1856]" italics="true" pageId="62" pageNumber="63">H</emphasis>
.
<emphasis id="B96DEAE15B5BFFADFBD9F505FB20F8D6" box="[1079,1174,1832,1856]" italics="true" pageId="62" pageNumber="63">colorata</emphasis>
</taxonomicName>
in having shallow pronotal foveae PF2 and a distinctive pronotal fascia (
<figureCitation id="13222A765B5BFFADFD32F561FC8DF8F2" box="[732,827,1868,1892]" captionStart="FIGURE 87" captionStartId="210.[151,250,1753,1776]" captionTargetBox="[222,1378,219,1722]" captionTargetId="figure-29@210.[195,1391,193,1732]" captionTargetPageId="210" captionText="FIGURE 87. Hydraena colorata, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283581" httpUri="https://zenodo.org/record/5283581/files/figure.png" pageId="62" pageNumber="63">Figs. 87</figureCitation>
,
<figureCitation id="13222A765B5BFFADFCA7F561FCDEF8F2" box="[841,872,1868,1892]" captionStart="FIGURE 98" captionStartId="218.[151,250,1753,1776]" captionTargetBox="[244,1366,222,1732]" captionTargetId="figure-29@218.[198,1389,193,1732]" captionTargetPageId="218" captionText="FIGURE 98. Hydraena nanocolorata, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283603" httpUri="https://zenodo.org/record/5283603/files/figure.png" pageId="62" pageNumber="63">98</figureCitation>
); distinguished therefrom by the smaller size (ca. 1.15 vs.
<quantity id="4CE19B165B5BFFADFF18F542FEE8F811" box="[246,350,1903,1927]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="62" pageNumber="63" unit="mm" value="1.25">1.25 mm</quantity>
), the differently shaped pronotum (PA slightly greater than PB), the proportionally shorter elytra, and the slightly sparser dorsal punctation. The pronotal fasciae also differ in the two species, the color band ratios being ca. 6/9/
<quantity id="4CE19B165B5BFFADFEC0F595FEEEF846" box="[302,344,1976,2000]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="62" pageNumber="63" unit="in" value="4.0">4 in</quantity>
<taxonomicName id="4C194D705B5BFFADFEB1F594FDA1F846" authorityName="Perkins" authorityYear="2011" box="[351,535,1976,2000]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="nanocolorata">
<emphasis id="B96DEAE15B5BFFADFEB1F594FEC5F846" box="[351,371,1977,2000]" italics="true" pageId="62" pageNumber="63">H</emphasis>
.
<emphasis id="B96DEAE15B5BFFADFE6EF595FDA1F846" box="[384,535,1976,2000]" italics="true" pageId="62" pageNumber="63">nanocolorata</emphasis>
</taxonomicName>
and ca. 7/10/
<quantity id="4CE19B165B5BFFADFD40F595FD6EF846" box="[686,728,1976,2000]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="62" pageNumber="63" unit="in" value="5.0">5 in</quantity>
<taxonomicName id="4C194D705B5BFFADFD30F594FCE9F846" authorityName="Perkins" authorityYear="2011" box="[734,863,1976,2000]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="colorata">
<emphasis id="B96DEAE15B5BFFADFD30F594FD44F846" box="[734,754,1977,2000]" italics="true" pageId="62" pageNumber="63">H</emphasis>
.
<emphasis id="B96DEAE15B5BFFADFCEEF595FCE9F846" box="[768,863,1976,2000]" italics="true" pageId="62" pageNumber="63">colorata</emphasis>
</taxonomicName>
. The male genitalia of the two species distinctively differ (
<figureCitation id="13222A765B5BFFADFF0AF5F1FEF7F862" box="[228,321,2012,2036]" captionStart="FIGURES 88–89" captionStartId="211.[151,264,1561,1584]" captionTargetBox="[249,1332,217,1516]" captionTargetId="figure-29@211.[226,1360,193,1540]" captionTargetPageId="211" captionText="FIGURES 88–89. Aedeagi of holotypes. 88. H. koma. 89. H. colorata." figureDoi="http://doi.org/10.5281/zenodo.5283585" httpUri="https://zenodo.org/record/5283585/files/figure.png" pageId="62" pageNumber="63">Figs. 89</figureCitation>
,
<figureCitation id="13222A765B5BFFADFEA3F5F1FECDF862" box="[333,379,2012,2036]" captionStart="FIGURES 100–101" captionStartId="220.[151,264,1650,1673]" captionTargetBox="[309,1276,224,1579]" captionTargetId="figure-32@220.[264,1323,193,1629]" captionTargetPageId="220" captionText="FIGURES 100–101. Aedeagi of holotypes. 100. H. nanocolorata. 101. H. clarinis." figureDoi="http://doi.org/10.5281/zenodo.5283609" httpUri="https://zenodo.org/record/5283609/files/figure.png" pageId="62" pageNumber="63">100</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30365785B5AFFACFF29F2BAFDAFFC51" pageId="63" pageNumber="64" type="description">
<paragraph id="8BA636F35B5AFFACFF29F2BAFCA8FE8D" blockId="63.[151,1437,151,1040]" pageId="63" pageNumber="64">
<emphasis id="B96DEAE15B5AFFACFF29F2BAFEEEFF26" bold="true" box="[199,344,151,176]" inLineHeading="true" pageId="63" pageNumber="64" reason="1">Description.</emphasis>
Size:
<typeStatus id="54A288515B5AFFACFE42F2BAFDB8FF39" box="[428,526,151,175]" pageId="63" pageNumber="64" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.15/0.52; head 0.17/0.29; pronotum 0.30/0.39, PA 0.33, PB 0.32; elytra 0.69/0.52. Dorsum of head piceous; pronotum testaceous to light brown in front of and behind piceous fascia, ratios of color bands, as measured in midline, ca. 6/9/4; elytra brown; legs light brown to testaceous; maxillary palpi testaceous.
</paragraph>
<paragraph id="8BA636F35B5AFFACFF29F305FCC5FE62" blockId="63.[151,1437,151,1040]" pageId="63" pageNumber="64">Frons punctures ca. 1xef; interstices shining, 2–3xpd. Clypeus microreticulate laterally, very finely sparsely punctate medially. Mentum and postmentum very sparsely and very finely punctulate, strongly shining. Genae very slightly raised, shining, without posterior ridge. Pronotum cordiform, anterior margin emarginate; punctures on disc ca. 1xpd those of frons, interstices shining, 2–4xpd, punctures slightly larger and denser at anterior and posterior; PF1 united to form shallow transverse impression across anterior 1/3 of pronotal disc; PF2 shallow, oval, oblique, shallowly confluent; PF3 moderately deep; PF4 shallow.</paragraph>
<paragraph id="8BA636F35B5AFFACFF29F3D2FE2BFD31" blockId="63.[151,1437,151,1040]" pageId="63" pageNumber="64">Elytra weakly convex, posterior declivity gradual, summit of posterior declivity slightly before midlength; lateral explanate margins narrow; on basal 1/3 punctures ca. 1xpd largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, shining, on disc ca. 1–2xpd, as are interstices between punctures of a row. Apices in dorsal aspect weakly separately rounded, in posterior aspect margins forming shallow angle with one another.</paragraph>
<paragraph id="8BA636F35B5AFFACFF29F099FDAFFC51" blockId="63.[151,1437,151,1040]" pageId="63" pageNumber="64">
Ratios of P2 width and plaque shape (P2/w/l/s) ca. 3/1/4/4. P1 laminate; median carina sinuate in profile. P2 very slightly raised, l/w ca. 2/1, sides slightly converging toward apex. Plaques short, carinate, located at sides of deep, inverted U-shaped median depression. Metaventrite with very low midlongitudinal ridge between P2 and apex of median depression. AIS slightly concave, width at straight posterior margin ca. 1.5x P2. Profemur (male) with small, pointed tubercle next to trochanter; protibia slightly arcuate, medial margin in distal 1/2 distinctively thickened and bearing row of very short, peg-like spines. Mesotibia straight. Metatibia straight, medial margin in distal 1/4 with few recumbent hair-like setae. Abdominal apex symmetrical; last tergite (male) deeply notched. Aedeagus as illustrated (
<figureCitation id="13222A765B5AFFACFE44F182FDBAFC51" box="[426,524,943,967]" captionStart="FIGURES 100–101" captionStartId="220.[151,264,1650,1673]" captionTargetBox="[309,1276,224,1579]" captionTargetId="figure-32@220.[264,1323,193,1629]" captionTargetPageId="220" captionText="FIGURES 100–101. Aedeagi of holotypes. 100. H. nanocolorata. 101. H. clarinis." figureDoi="http://doi.org/10.5281/zenodo.5283609" httpUri="https://zenodo.org/record/5283609/files/figure.png" pageId="63" pageNumber="64">Fig. 100</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30365785B5AFFACFF29F1F9FBC6FC7D" box="[199,1136,980,1004]" pageId="63" pageNumber="64" type="etymology">
<paragraph id="8BA636F35B5AFFACFF29F1F9FBC6FC7D" blockId="63.[151,1437,151,1040]" box="[199,1136,980,1004]" pageId="63" pageNumber="64">
<emphasis id="B96DEAE15B5AFFACFF29F1F9FEFBFC7A" bold="true" box="[199,333,980,1004]" inLineHeading="true" pageId="63" pageNumber="64" reason="1">Etymology.</emphasis>
Named in reference to the small size and relationship with
<taxonomicName id="4C194D705B5AFFACFC06F1F8FBDFFC7A" authorityName="Perkins" authorityYear="2011" box="[1000,1129,980,1004]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="colorata">
<emphasis id="B96DEAE15B5AFFACFC06F1F8FC4AFC7A" box="[1000,1020,981,1004]" italics="true" pageId="63" pageNumber="64">H</emphasis>
.
<emphasis id="B96DEAE15B5AFFACFBE4F1F9FBDFFC7A" box="[1034,1129,980,1004]" italics="true" pageId="63" pageNumber="64">colorata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C30365785B5AFFACFF29F1DAFA39FB99" box="[199,1423,1015,1040]" pageId="63" pageNumber="64" type="distribution">
<paragraph id="8BA636F35B5AFFACFF29F1DAFA39FB99" blockId="63.[151,1437,151,1040]" box="[199,1423,1015,1040]" pageId="63" pageNumber="64">
<emphasis id="B96DEAE15B5AFFACFF29F1DAFED6FB86" bold="true" box="[199,352,1015,1040]" inLineHeading="true" pageId="63" pageNumber="64" reason="1">Distribution.</emphasis>
Currently known from two localities in eastern Area 3; elevations
<quantity id="4CE19B165B5AFFACFBA3F1DAFB22FB99" box="[1101,1172,1015,1039]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="63" pageNumber="64" unit="m" value="160.0">160 m</quantity>
and
<quantity id="4CE19B165B5AFFACFB24F1DAFAA7FB99" box="[1226,1297,1015,1039]" metricMagnitude="2" metricUnit="m" metricValue="3.05" pageId="63" pageNumber="64" unit="m" value="305.0">305 m</quantity>
(
<figureCitation id="13222A765B5AFFACFACEF1DAFA34FB99" box="[1312,1410,1015,1039]" captionStart="FIGURES 475–478" captionStartId="394.[151,264,1650,1673]" captionTargetBox="[165,1421,259,1607]" captionTargetId="figure-23@394.[165,1421,193,1629]" captionTargetPageId="394" captionText="FIGURES 475–478. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284093" httpUri="https://zenodo.org/record/5284093/files/figure.png" pageId="63" pageNumber="64">Fig. 475</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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