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<document id="423F441937DB404C5454203388CD1BE8" ID-DOI="10.50826/bnmnszool.48.2_35" ID-ISSN="2434-091X" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="karina" IM.tables_approvedBy="karina" IM.taxonomicNames_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1718643474207" checkinUser="felipe" docAuthor="Manikandan, K, Megalaa, N, Valliappan, Subramanian, Nandini, K, Rani, Lourdu V, Dakshinamurthi, Senthil &amp; Nagappan, Nagappan" docDate="2022" docId="4B2487854212A51B392FA3FA28CBFC4C" docLanguage="en" docName="BullNatlMusNatSciSerA.48.2.35-83.pdf" docOrigin="Bulletin of the National Museum of Nature and Science. Series A, Zoology 48 (2)" docSource="http://dx.doi.org/10.4103/jpbs.jpbs_806_23" docStyle="DocumentStyle:A6FFB80BFE6276686563735C80A89639.2:BullNatlMusNatSciSerA.2021-.journal_article.english.affilindex" docStyleId="A6FFB80BFE6276686563735C80A89639" docStyleName="BullNatlMusNatSciSerA.2021-.journal_article.english.affilindex" docStyleVersion="2" docTitle="Viaderiana typica Ward 1942" docType="treatment" docVersion="2" lastPageNumber="76" masterDocId="B71DFFFD423AA5323A05A1332814FFCE" masterDocTitle="Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 3. Sahul Shelf" masterLastPageNumber="83" masterPageNumber="35" pageNumber="75" updateTime="1726962684028" updateUser="karina" zenodo-license-document="CC-BY-4.0">
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<mods:title id="00BBDD329FA2784325799BA5A8A9CB67">Crabs (Crustacea, Decapoda) from the Seas of East and Southeast Asia Collected by the RV Hakuhō Maru (KH- 72 - 1 Cruise) 3. Sahul Shelf</mods:title>
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<mods:namePart id="B5539447D6BFE2CE0EDCDDFA064C8589">Manikandan, K</mods:namePart>
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<mods:namePart id="CCA57AAB1D724DCA7716F6DFF0E3C910">Megalaa, N</mods:namePart>
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<mods:namePart id="CF7F55B596D6EA36AE9B642686F330D6">Valliappan, Subramanian</mods:namePart>
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<mods:namePart id="9590208707DC6C0B5B85A62F237D7741">Nandini, K</mods:namePart>
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<mods:namePart id="B91A3AD93F4E1F16366B5154041DBEC5">Rani, Lourdu V</mods:namePart>
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<mods:namePart id="CBA3C4C2A6D670D298301C50A82E9F57">Dakshinamurthi, Senthil</mods:namePart>
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<mods:namePart id="CBA7A7827CF96F4558D56BAB68EE78EB">Nagappan, Nagappan</mods:namePart>
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<mods:title id="5B4ECB4D381EBCD67F4FDA25CA9A8AED">Bulletin of the National Museum of Nature and Science. Series A, Zoology</mods:title>
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<mods:date id="BBB94050DD07B09475B7A6B2DCE2FE4A">2022</mods:date>
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<mods:number id="3816FA14928F792842523E1E069550B9">2022-05-20</mods:number>
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<mods:number id="144999D18EEF4C9EA079FB0E71D26D9F">48</mods:number>
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<mods:identifier id="684B791FFA36726F7D3134F7B0FD8116" type="DOI">10.50826/bnmnszool.48.2_35</mods:identifier>
<mods:identifier id="8C2E6C6916EFBF7B135E26DD44394FB1" type="ISSN">2434-091X</mods:identifier>
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<treatment id="4B2487854212A51B392FA3FA28CBFC4C" LSID="urn:lsid:plazi:treatment:4B2487854212A51B392FA3FA28CBFC4C" httpUri="http://treatment.plazi.org/id/4B2487854212A51B392FA3FA28CBFC4C" lastPageId="41" lastPageNumber="76" pageId="40" pageNumber="75">
<subSubSection id="8B9765184212A51A392FA3FA2C79FD2C" box="[810,1133,713,738]" pageId="40" pageNumber="75" type="nomenclature">
<paragraph id="C33236934212A51A392FA3FA2C79FD2C" blockId="40.[764,1178,681,738]" box="[810,1133,713,738]" pageId="40" pageNumber="75">
<heading id="987A81FF4212A51A392FA3FA2C79FD2C" box="[810,1133,713,738]" centered="true" fontSize="9" level="2" pageId="40" pageNumber="75" reason="4">
<taxonomicName id="048D4D104212A51A392FA3FA2C79FD2C" ID-CoL="5BBQG" authority="Ward, 1942" authorityName="Ward" authorityYear="1942" box="[810,1133,713,738]" class="Malacostraca" family="Pilumnidae" genus="Viaderiana" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="75" phylum="Arthropoda" rank="species" species="typica">
<emphasis id="F1F9EA814212A51A392FA3FA2BFFFD2C" bold="true" box="[810,1003,713,738]" italics="true" pageId="40" pageNumber="75">Viaderiana typica</emphasis>
<bibRefCitation id="A71C4B624212A51A39F4A3F92C79FD2C" author="Ward, M." box="[1009,1133,714,738]" pageId="40" pageNumber="75" pagination="49 - 108" refId="ref32408" refString="Ward, M. 1942. Notes on the Crustacea of the Desjardins Museum, Mauritius Institute with descriptions of new genera and species. The Mauritius Institute Bulletin 2: 49 - 108, pls. 5 - 6." type="journal article" year="1942">Ward, 1942</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8B9765184212A51B38D8A23128CBFC4C" lastPageId="41" lastPageNumber="76" pageId="40" pageNumber="75" type="materials_examined">
<paragraph id="C33236934212A51A38D8A2312CA5FC91" blockId="40.[707,1237,770,1799]" pageId="40" pageNumber="75">
<emphasis id="F1F9EA814212A51A38D8A2312BD3FCD4" box="[733,967,770,794]" italics="true" pageId="40" pageNumber="75">Material examined.</emphasis>
<materialsCitation id="73E53CCE4212A51A39E0A2312BFFFC91" collectionCode="RV" pageId="40" pageNumber="75" specimenCode="CB 4.1" specimenCount="1">
<collectionCode id="A59CAE564212A51A39E0A2312C1EFCD7" box="[997,1034,770,793]" country="Italy" name="Collection of Leptospira Strains" pageId="40" pageNumber="75">RV</collectionCode>
<emphasis id="F1F9EA814212A51A3E23A2312CC0FCD4" box="[1062,1236,770,794]" italics="true" pageId="40" pageNumber="75">Hakuhō Maru</emphasis>
KH-72-1 cruise, sta. 29,
<specimenCount id="D58BFD1A4212A51A39CBA2162C1FFCF3" box="[974,1035,805,829]" count="1" pageId="40" pageNumber="75" type="generic">1 juv.</specimenCount>
(
<specimenCode id="932B9EE84212A51A3E1FA2162C73FCF3" box="[1050,1127,805,829]" collectionCode="CB" country="Australia" name="The CB Rhizobium Collection" pageId="40" pageNumber="75">CB 4.1</specimenCode>
including lateral spines×CL 4.0 mm)
</materialsCitation>
, NSMT-Cr 30764.
</paragraph>
<paragraph id="C33236934212A51A38D8A2592BD3FB77" blockId="40.[707,1237,770,1799]" pageId="40" pageNumber="75">
<emphasis id="F1F9EA814212A51A38D8A2592B2EFC4C" box="[733,826,874,898]" italics="true" pageId="40" pageNumber="75">Remarks</emphasis>
. In the genus
<taxonomicName id="048D4D104212A51A39EDA2592C4AFC4C" authorityName="Ward" authorityYear="1942" box="[1000,1118,874,898]" class="Malacostraca" family="Pilumnidae" genus="Viaderiana" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="75" phylum="Arthropoda" rank="genus">
<emphasis id="F1F9EA814212A51A39EDA2592C4AFC4C" box="[1000,1118,874,898]" italics="true" pageId="40" pageNumber="75">Viaderiana</emphasis>
</taxonomicName>
, the carapace dorsal surface is typically covered with long, rather sparse, silky setae, and shallowly divided into regions; the carapace anterolateral margin is armed with two spiniform tubercles and a third vestigial tooth or granule behind the external orbital angle. The specimen at hand is unfortunately lacking all the chelipeds and ambulatory legs, while the generic affiliation is without doubt in
<taxonomicName id="048D4D104212A51A394FA5912BD4FB74" authorityName="Ward" authorityYear="1942" box="[842,960,1186,1210]" class="Malacostraca" family="Pilumnidae" genus="Viaderiana" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="75" phylum="Arthropoda" rank="genus">
<emphasis id="F1F9EA814212A51A394FA5912BD4FB74" box="[842,960,1186,1210]" italics="true" pageId="40" pageNumber="75">Viaderiana</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="C33236934212A51B38D8A5F62A65FD84" blockId="40.[707,1237,770,1799]" lastBlockId="41.[140,669,215,898]" lastPageId="41" lastPageNumber="76" pageId="40" pageNumber="75">
The carapace is narrow and distinctly quadrate with the almost longitudinal posterolateral margins, and the posterior of the two anterolateral spines is slightly larger than the first. The latter character is of specific value in
<taxonomicName id="048D4D104212A51A3E1DA47C2C9AFAA9" authorityName="Ward" authorityYear="1942" box="[1048,1166,1359,1383]" class="Malacostraca" family="Pilumnidae" genus="Viaderiana" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="75" phylum="Arthropoda" rank="genus">
<emphasis id="F1F9EA814212A51A3E1DA47C2C9AFAA9" box="[1048,1166,1359,1383]" italics="true" pageId="40" pageNumber="75">Viaderiana</emphasis>
</taxonomicName>
, but it is also affected by maturity, and the present specimen is a juvenile. As for the narrowly quadrate carapace, this specimen is close to
<taxonomicName id="048D4D104212A51A3E6AA48B2B5EFA3C" authority="Ward, 1942" authorityName="Ward" authorityYear="1942" class="Malacostraca" family="Pilumnidae" genus="Viaderiana" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="75" phylum="Arthropoda" rank="species" species="typica">
<emphasis id="F1F9EA814212A51A3E6AA48B2CC0FA01" box="[1135,1236,1464,1487]" italics="true" pageId="40" pageNumber="75">V. typica</emphasis>
<bibRefCitation id="A71C4B624212A51A38C6A4E92B5EFA3C" author="Ward, M." box="[707,842,1498,1522]" pageId="40" pageNumber="75" pagination="49 - 108" refId="ref32408" refString="Ward, M. 1942. Notes on the Crustacea of the Desjardins Museum, Mauritius Institute with descriptions of new genera and species. The Mauritius Institute Bulletin 2: 49 - 108, pls. 5 - 6." type="journal article" year="1942">Ward, 1942</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="048D4D104212A51A3990A4E82B10F9DB" authority="(Zehntner, 1894)" baseAuthorityName="Zehntner" baseAuthorityYear="1894" class="Malacostraca" family="Pilumnidae" genus="Viaderiana" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="75" phylum="Arthropoda" rank="species" species="quadrispinosa">
<emphasis id="F1F9EA814212A51A3990A4E82C47FA3C" box="[917,1107,1498,1522]" italics="true" pageId="40" pageNumber="75">V. quadrispinosa</emphasis>
(
<bibRefCitation id="A71C4B624212A51A3E69A4E92AE9F9DB" author="Zehntner, L." pageId="40" pageNumber="75" pagination="135 - 214" refId="ref32914" refString="Zehntner, L. 1894. Crustaces de l`Archipel Malais. Voyage de MM. M. Bedot et C. Pictes dans l`Archipel Malais. Revue Suisse de Zoologie 2: 135 - 214, pls. 7 - 9." type="journal article" year="1894">Zehntner, 1894</bibRefCitation>
)
</taxonomicName>
. The latter species is well known by its characteristic chocolate brown color pattern, although in the specimen examined, the color is completely faded, without trace of color pattern. In adult
<taxonomicName id="048D4D104212A51A3927A7BB2B95F951" authorityName="Ward" authorityYear="1942" box="[802,897,1672,1695]" class="Malacostraca" family="Pilumnidae" genus="Viaderiana" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="75" phylum="Arthropoda" rank="species" species="typica">
<emphasis id="F1F9EA814212A51A3927A7BB2B95F951" box="[802,897,1672,1695]" italics="true" pageId="40" pageNumber="75">V. typica</emphasis>
</taxonomicName>
, the carapace has the coarsely areolated dorsal surface and stronger anterolateral teeth. Recently,
<bibRefCitation id="A71C4B624212A51A39A0A7FE2C65F92B" author="Maenosono, T." box="[933,1137,1741,1765]" pageId="40" pageNumber="75" pagination="15 - 36" refId="ref28489" refString="Maenosono, T. 2020. Notes on some species of the genus Viaderiana Ward, 1942 (Crustacea: Decapoda: Brachyura: Pilumnidae) from the Ryukyu Islands, southern Japan. Fauna Ryukyuana 58: 15 - 36 (In Japanese with English summary)." type="journal article" year="2020">Maenosono (2020)</bibRefCitation>
reported on four species of
<taxonomicName id="048D4D104212A51A3992A7DC2C19F8C9" authorityName="Ward" authorityYear="1942" box="[919,1037,1775,1799]" class="Malacostraca" family="Pilumnidae" genus="Viaderiana" kingdom="Animalia" order="Decapoda" pageId="40" pageNumber="75" phylum="Arthropoda" rank="genus">
<emphasis id="F1F9EA814212A51A3992A7DC2C19F8C9" box="[919,1037,1775,1799]" italics="true" pageId="40" pageNumber="75">Viaderiana</emphasis>
</taxonomicName>
from the Ryukyu Islands, namely,
<taxonomicName id="048D4D104213A51B3B41A1EB28D9FEDC" authority="(Takeda and Miyake, 1969)" baseAuthorityName="Takeda and Miyake" baseAuthorityYear="1969" class="Malacostraca" family="Pilumnidae" genus="Viaderiana" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="76" phylum="Arthropoda" rank="species" species="incerta">
<emphasis id="F1F9EA814213A51B3B41A1EB29B9FF21" box="[324,429,216,239]" italics="true" pageId="41" pageNumber="76">V. incerta</emphasis>
(Takeda and Miyake, 1969)
</taxonomicName>
,
<taxonomicName id="048D4D104213A51B3ADBA1C829C6FEDC" authority="Ward, 1942" authorityName="Ward" authorityYear="1942" box="[222,466,250,274]" class="Malacostraca" family="Pilumnidae" genus="Viaderiana" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="76" phylum="Arthropoda" rank="species" species="typica">
<emphasis id="F1F9EA814213A51B3ADBA1C82954FEDC" box="[222,320,251,274]" italics="true" pageId="41" pageNumber="76">V. typica</emphasis>
<bibRefCitation id="A71C4B624213A51B3B48A1C929C6FEDC" author="Ward, M." box="[333,466,250,274]" pageId="41" pageNumber="76" pagination="49 - 108" refId="ref32408" refString="Ward, M. 1942. Notes on the Crustacea of the Desjardins Museum, Mauritius Institute with descriptions of new genera and species. The Mauritius Institute Bulletin 2: 49 - 108, pls. 5 - 6." type="journal article" year="1942">Ward, 1942</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="048D4D104213A51B3BE6A1C8295EFEFB" authority="(Zehntner, 1894)" baseAuthorityName="Zehntner" baseAuthorityYear="1894" class="Malacostraca" family="Pilumnidae" genus="Viaderiana" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="76" phylum="Arthropoda" rank="species" species="quadrispinosa">
<emphasis id="F1F9EA814213A51B3BE6A1C82A89FEDC" box="[483,669,250,274]" italics="true" pageId="41" pageNumber="76">V. quadrispinosa</emphasis>
(
<bibRefCitation id="A71C4B624213A51B3A96A02E2956FEFB" author="Zehntner, L." box="[147,322,285,309]" pageId="41" pageNumber="76" pagination="135 - 214" refId="ref32914" refString="Zehntner, L. 1894. Crustaces de l`Archipel Malais. Voyage de MM. M. Bedot et C. Pictes dans l`Archipel Malais. Revue Suisse de Zoologie 2: 135 - 214, pls. 7 - 9." type="journal article" year="1894">Zehntner, 1894</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="048D4D104213A51B3B94A02E292EFE99" authority="(A. Milne-Edwards, 1873)" baseAuthorityName="A. Milne-Edwards" baseAuthorityYear="1873" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="76" phylum="Arthropoda" rank="species" species="longipes">
<emphasis id="F1F9EA814213A51B3B94A02E2A04FEFB" box="[401,528,285,309]" italics="true" pageId="41" pageNumber="76">V. longipes</emphasis>
(A.
<bibRefCitation id="A71C4B624213A51B3851A02E2926FE99" author="Milne-Edwards, A." pageId="41" pageNumber="76" pagination="155 - 332" refId="ref29278" refString="Milne-Edwards, A. 1873. Recherches sur la faune carcinologique de la Nouvelle-Caledonie. Deuxieme Partie. Nouvelles Archives du Museum d`Histoire Naturelle, Paris 8: 155 - 332, pls. 4 - 18." type="journal article" year="1873">Milne-Edwards, 1873</bibRefCitation>
)
</taxonomicName>
, and discussed the identification challenges in detail, with fine photographs. Following the details elucidated by
<bibRefCitation id="A71C4B624213A51B3824A0B628C2FE71" author="Maenosono, T." pageId="41" pageNumber="76" pagination="15 - 36" refId="ref28489" refString="Maenosono, T. 2020. Notes on some species of the genus Viaderiana Ward, 1942 (Crustacea: Decapoda: Brachyura: Pilumnidae) from the Ryukyu Islands, southern Japan. Fauna Ryukyuana 58: 15 - 36 (In Japanese with English summary)." type="journal article" year="2020">Maenosono (2020)</bibRefCitation>
, the young specimen at hand, having no chelipeds and ambulatory legs, is reasonably, but tentatively identified as
<taxonomicName id="048D4D104213A51B3B95A0DE29F8FDCA" authorityName="Ward" authorityYear="1942" box="[400,492,493,516]" class="Malacostraca" family="Pilumnidae" genus="Viaderiana" kingdom="Animalia" order="Decapoda" pageId="41" pageNumber="76" phylum="Arthropoda" rank="species" species="typica">
<emphasis id="F1F9EA814213A51B3B95A0DE29F8FDCA" box="[400,492,493,516]" italics="true" pageId="41" pageNumber="76">V. typica</emphasis>
</taxonomicName>
among the congeners from the Ryukyu Islands as well as the other, about 14, Indo-West Pacific congeners.
</paragraph>
<paragraph id="C33236934213A51B3AA2A36628CBFC4C" blockId="41.[140,669,215,898]" pageId="41" pageNumber="76">
<emphasis id="F1F9EA814213A51B3AA2A366293DFDA3" box="[167,297,597,621]" italics="true" pageId="41" pageNumber="76">Distribution</emphasis>
. West Pacific (Ryukyu Islands,
<collectingCountry id="BB9A76034213A51B3A89A34428D3FD41" box="[140,199,631,655]" name="Palau" pageId="41" pageNumber="76">Palau</collectingCountry>
Islands,
<collectingCountry id="BB9A76034213A51B3B33A34429B4FD41" box="[310,416,631,655]" name="Indonesia" pageId="41" pageNumber="76">Indonesia</collectingCountry>
,
<collectingCountry id="BB9A76034213A51B3BB7A3442A35FD41" box="[434,545,631,655]" name="Vietnam" pageId="41" pageNumber="76">Viet Nam</collectingCountry>
and
<collectingCountry id="BB9A76034213A51B3866A34428AAFD7C" name="Thailand" pageId="41" pageNumber="76">Thailand</collectingCountry>
), Indian Ocean (
<collectingCountry id="BB9A76034213A51B3B71A3A929F5FD7C" box="[372,481,666,690]" name="Mauritius" pageId="41" pageNumber="76">Mauritius</collectingCountry>
), and now northern
<collectingCountry id="BB9A76034213A51B3AB0A38E290EFD1B" box="[181,282,701,725]" name="Australia" pageId="41" pageNumber="76">Australia</collectingCountry>
, as referred to
<bibRefCitation id="A71C4B624213A51B3BB8A38E2A5CFD1B" author="Ward, M." box="[445,584,701,725]" pageId="41" pageNumber="76" pagination="49 - 108" refId="ref32408" refString="Ward, M. 1942. Notes on the Crustacea of the Desjardins Museum, Mauritius Institute with descriptions of new genera and species. The Mauritius Institute Bulletin 2: 49 - 108, pls. 5 - 6." type="journal article" year="1942">Ward (1942)</bibRefCitation>
,
<bibRefCitation id="A71C4B624213A51B3851A38E28D8FD39" author="Serene, R." pageId="41" pageNumber="76" pagination="903 - 918" refId="ref30964" refString="Serene, R. 1971. Observations preliminaires sur les brachyoures nouveaux ou mal connus du sud-east Asi- atique (Crustacea Decapoda). Bulletin du Museum national d`Histoire naturelle, Paris (2) 32: 903 - 918." type="journal article" year="1971">Serène (1971</bibRefCitation>
,
<bibRefCitation id="A71C4B624213A51B3AE5A3EC290DFD39" author="Serene, R." box="[224,281,735,759]" pageId="41" pageNumber="76" pagination="711 - 722" refId="ref31005" refString="Serene, R. 1980. Notes sur quelques crustaces brachyoures provenant principalement l`ile Maurice et conserves au Museum d`Histoire Naturelle de Geneve. Description d`Etisus zehntneri sp. nov. Revue Suisse de Zoologie 87: 711 - 722." type="journal article" year="1980">1980</bibRefCitation>
),
<bibRefCitation id="A71C4B624213A51B3B36A3EC29CCFD39" author="Takeda, M." box="[307,472,735,759]" pageId="41" pageNumber="76" pagination="185 - 213" refId="ref31646" refString="Takeda, M. 1972. New and rare crabs from the Palau Islands. Micronesica 7: 185 - 213." type="journal article" year="1972">Takeda (1972)</bibRefCitation>
, and
<bibRefCitation id="A71C4B624213A51B3824A3D328C3FCD4" author="Maenosono, T." pageId="41" pageNumber="76" pagination="15 - 36" refId="ref28489" refString="Maenosono, T. 2020. Notes on some species of the genus Viaderiana Ward, 1942 (Crustacea: Decapoda: Brachyura: Pilumnidae) from the Ryukyu Islands, southern Japan. Fauna Ryukyuana 58: 15 - 36 (In Japanese with English summary)." type="journal article" year="2020">Maenosono (2020)</bibRefCitation>
.
<bibRefCitation id="A71C4B624213A51B3AE3A23129A1FCD4" author="Maenosono, T." box="[230,437,770,794]" pageId="41" pageNumber="76" pagination="15 - 36" refId="ref28489" refString="Maenosono, T. 2020. Notes on some species of the genus Viaderiana Ward, 1942 (Crustacea: Decapoda: Brachyura: Pilumnidae) from the Ryukyu Islands, southern Japan. Fauna Ryukyuana 58: 15 - 36 (In Japanese with English summary)." type="journal article" year="2020">Maenosono (2020)</bibRefCitation>
recorded
<specimenCount id="D58BFD1A4213A51B382EA2312A89FCD4" box="[555,669,770,794]" count="2" pageId="41" pageNumber="76" type="male">two males</specimenCount>
collected from crevices of a dead coral block at Urasoe, Okinawa-jima in the central Ryukyu Islands.
</paragraph>
</subSubSection>
</treatment>
</document>
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