treatments-xml/data/03/85/87/038587BBE368FFE3FF75F9ACBA8CF86E.xml

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<document id="A0193E49CCFD514553CAF2F2B64E7EE7" ID-CLB-Dataset="31639" ID-DOI="10.11646/zootaxa.4342.1.1" ID-GBIF-Dataset="15799d2a-1968-4874-8100-452597d4e26b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1041066" ID-ZooBank="2ACD54D2-3487-432D-9323-EEC131FE2E64" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1509694849742" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2017" docId="038587BBE368FFE3FF75F9ACBA8CF86E" docLanguage="en" docName="zootaxa.4342.1.1.pdf.imf" docOrigin="Zootaxa 4342 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydraena dilutipoides Perkins 2017, new species" docType="treatment" docVersion="9" lastPageNumber="115" masterDocId="FFBCFFC3E307FF91FFE2FFA5B93CFF85" masterDocTitle="Hydraenidae of Madagascar (Insecta: Coleoptera)" masterLastPageNumber="264" masterPageNumber="1" pageNumber="112" updateTime="1698480684898" updateUser="ExternalLinkService">
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<mods:title id="21A38750E4BD9801C498736F45929266">Hydraenidae of Madagascar (Insecta: Coleoptera)</mods:title>
</mods:titleInfo>
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<mods:namePart id="8998F2EAFDD6859F6C4F8D4761E15049">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="13814E73C078F0D82340FECFE3350F43">text</mods:typeOfResource>
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<mods:title id="5D5B577EC274AC136264F890B43E9E17">Zootaxa</mods:title>
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<mods:part id="7691DC3D6090F6D0761331A0A7E93567">
<mods:date id="374C0B4E79A9E1162BD163A6DB662EFD">2017</mods:date>
<mods:detail id="1EF250566300742608FFE84B13D93037" type="pubDate">
<mods:number id="02EE5282C25D9FFF362D3F9E4C68517F">2017-11-03</mods:number>
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<mods:number id="195B82E0C3418F3445A1662352D45EE9">4342</mods:number>
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<treatment id="038587BBE368FFE3FF75F9ACBA8CF86E" ID-DOI="http://doi.org/10.5281/zenodo.5323614" ID-GBIF-Taxon="137233438" ID-Zenodo-Dep="5323614" LSID="urn:lsid:plazi:treatment:038587BBE368FFE3FF75F9ACBA8CF86E" httpUri="http://treatment.plazi.org/id/038587BBE368FFE3FF75F9ACBA8CF86E" lastPageId="114" lastPageNumber="115" pageId="111" pageNumber="112">
<subSubSection id="C3366526E368FFFEFF75F9ACBBAAF9C1" pageId="111" pageNumber="112" type="nomenclature">
<paragraph id="8B9336ADE368FFFEFF75F9ACBB7DF9A6" blockId="111.[151,662,1545,1605]" box="[151,577,1545,1571]" pageId="111" pageNumber="112">
<heading id="D0DB81C1E368FFFEFF75F9ACBB7DF9A6" bold="true" box="[151,577,1545,1571]" fontSize="11" level="1" pageId="111" pageNumber="112" reason="1">
<emphasis id="B958EABFE368FFFEFF75F9ACBB7DF9A6" bold="true" box="[151,577,1545,1571]" pageId="111" pageNumber="112">
<taxonomicName id="4C2C4D2EE368FFFEFF75F9ACB89EF9A6" ID-CoL="3N395" authority="Perkins, 2017" authorityName="Perkins" authorityYear="2017" box="[151,418,1545,1571]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="111" pageNumber="112" phylum="Arthropoda" rank="species" species="dilutipoides" status="sp. nov.">
<emphasis id="B958EABFE368FFFEFF75F9ACB89EF9A6" bold="true" box="[151,418,1545,1571]" italics="true" pageId="111" pageNumber="112">Hydraena dilutipoides</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A26B57C4E368FFFEFE53F9ACBB7DF9A6" box="[433,577,1545,1571]" pageId="111" pageNumber="112" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B9336ADE368FFFEFF75F989BBAAF9C1" blockId="111.[151,662,1545,1605]" box="[151,662,1580,1605]" pageId="111" pageNumber="112">
<figureCitation id="13172A28E368FFFEFF75F989B9CDF9C1" box="[151,241,1580,1605]" captionStart="FIGURE 99" captionStartId="112.[151,250,1445,1467]" captionText="FIGURE 99. Hydraena dilutipoides, holotype, habitus dorsal, lateral, and ventral." pageId="111" pageNumber="112">Figs. 99</figureCitation>
(habitus), 102 (aedeagus), 276 (map)
</paragraph>
</subSubSection>
<subSubSection id="C3366526E368FFFEFF75F9D1BD16F899" pageId="111" pageNumber="112" type="materials_examined">
<paragraph id="8B9336ADE368FFFEFF75F9D1BD16F899" blockId="111.[151,1436,1652,2037]" pageId="111" pageNumber="112">
<emphasis id="B958EABFE368FFFEFF75F9D1B8FCF908" bold="true" box="[151,448,1652,1677]" pageId="111" pageNumber="112">
<typeStatus id="5497880FE368FFFEFF75F9D1B9EEF908" box="[151,210,1652,1677]" pageId="111" pageNumber="112">Type</typeStatus>
Material.
<typeStatus id="5497880FE368FFFEFEB1F9D1B8FCF908" box="[339,448,1652,1677]" pageId="111" pageNumber="112" type="holotype">Holotype</typeStatus>
</emphasis>
(male):
<collectingRegion id="49E8F84FE368FFFEFDC1F9D1BB90F908" box="[547,684,1652,1677]" country="Madagascar" name="Mahajanga" pageId="111" pageNumber="112">
<emphasis id="B958EABFE368FFFEFDC1F9D1BB90F908" bold="true" box="[547,684,1652,1677]" pageId="111" pageNumber="112">Mahajanga</emphasis>
</collectingRegion>
, Parc National Tsingy de Bemaraha,
<quantity id="4CD49B48E368FFFEFBBEF9D1BD90F908" box="[1116,1196,1652,1677]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="111" pageNumber="112" unit="km" value="2.5">2.5 km</quantity>
62° ENE Bekopaka, Ankidrodroa River, sifted litter, tropical dry forest on Tsingy, elev.
<quantity id="4CD49B48E368FFFEFC46F93DBACCF92A" box="[932,1008,1688,1712]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="111" pageNumber="112" unit="m" value="100.0">100 m</quantity>
,
<geoCoordinate id="EE18506AE368FFFEFC1DF93DBD63F935" box="[1023,1119,1688,1712]" direction="south" orientation="latitude" pageId="111" pageNumber="112" precision="925" value="-19.116667">19° 7' S</geoCoordinate>
,
<geoCoordinate id="EE18506AE368FFFEFB8DF93DBDE3F92A" box="[1135,1247,1688,1712]" direction="east" orientation="longitude" pageId="111" pageNumber="112" precision="925" value="44.8">44° 48' E</geoCoordinate>
,
<date id="FF92106DE368FFFEFB0CF932BCA4F935" box="[1262,1432,1687,1712]" pageId="111" pageNumber="112" value="2001-11-11" valueMax="2001-11-15" valueMin="2001-11-11">1115 xi 2001</date>
, Fisher, Griswold
<emphasis id="B958EABFE368FFFEFE83F918B8A1F950" box="[353,413,1724,1749]" italics="true" pageId="111" pageNumber="112">et al.</emphasis>
(BLF 4340) (
<collectionCode id="ED3DAE68E368FFFEFDA6F919BB41F951" box="[580,637,1724,1748]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="111" pageNumber="112">CAS</collectionCode>
)
<typeStatus id="5497880FE368FFFEFD73F919BA35F950" box="[657,777,1724,1749]" pageId="111" pageNumber="112" type="paratype">
<emphasis id="B958EABFE368FFFEFD73F919BA35F950" bold="true" box="[657,777,1724,1749]" pageId="111" pageNumber="112">Paratypes</emphasis>
</typeStatus>
(6): Same data as
<typeStatus id="5497880FE368FFFEFC0FF919BD73F950" box="[1005,1103,1724,1749]" pageId="111" pageNumber="112" type="holotype">holotype</typeStatus>
(4
<collectionCode id="ED3DAE68E368FFFEFB9EF919BD8BF951" box="[1148,1207,1724,1748]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="111" pageNumber="112">CAS</collectionCode>
);
<collectingRegion id="49E8F84FE368FFFEFB2DF919BC64F950" box="[1231,1368,1724,1749]" country="Madagascar" name="Mahajanga" pageId="111" pageNumber="112">
<emphasis id="B958EABFE368FFFEFB2DF919BC64F950" bold="true" box="[1231,1368,1724,1749]" pageId="111" pageNumber="112">Mahajanga</emphasis>
</collectingRegion>
, Parc National Tsingy de Bemaraha,
<quantity id="4CD49B48E368FFFEFE13F945BB03F97D" box="[497,575,1759,1784]" metricMagnitude="3" metricUnit="m" metricValue="3.4" pageId="111" pageNumber="112" unit="km" value="3.4">3.4 km</quantity>
93° E Bekopaka, Tombeau Vazimba, sifted litter, tropical dry forest, elev.
<quantity id="4CD49B48E368FFFEFA62F97AB993F899" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="111" pageNumber="112" unit="m" value="50.0">50 m</quantity>
,
<geoCoordinate id="EE18506AE368FFFEFF59F8A1B828F899" box="[187,276,1796,1820]" direction="south" orientation="latitude" pageId="111" pageNumber="112" precision="925" value="-19.133333">19° 8' S</geoCoordinate>
,
<geoCoordinate id="EE18506AE368FFFEFEC2F8A1B8B6F899" box="[288,394,1796,1820]" direction="east" orientation="longitude" pageId="111" pageNumber="112" precision="925" value="44.816666">44° 49' E</geoCoordinate>
,
<date id="FF92106DE368FFFEFE74F8A1BB10F899" box="[406,556,1796,1821]" pageId="111" pageNumber="112" value="2001-11-06" valueMax="2001-11-10" valueMin="2001-11-06">610 xi 2001</date>
, Fisher, Griswold
<emphasis id="B958EABFE368FFFEFD18F8A0BA0EF898" box="[762,818,1796,1821]" italics="true" pageId="111" pageNumber="112">et al.</emphasis>
(BLF 4232) (2
<collectionCode id="ED3DAE68E368FFFEFC01F8A1BD22F899" box="[995,1054,1796,1820]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="111" pageNumber="112">CAS</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3366526E368FFFEFF25F882BC13F871" pageId="111" pageNumber="112" type="diagnosis">
<paragraph id="8B9336ADE368FFFEFF25F882BC13F871" blockId="111.[151,1436,1652,2037]" pageId="111" pageNumber="112">
<emphasis id="B958EABFE368FFFEFF25F882B8E4F8C5" bold="true" box="[199,472,1831,1856]" pageId="111" pageNumber="112">Differential Diagnosis.</emphasis>
Based on the aedeagi, closely related to
<taxonomicName id="4C2C4D2EE368FFFEFC2EF88CBD68F8C5" box="[972,1108,1831,1856]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="111" pageNumber="112" phylum="Arthropoda" rank="species" species="dilutipes">
<emphasis id="B958EABFE368FFFEFC2EF88CBD68F8C5" box="[972,1108,1831,1856]" italics="true" pageId="111" pageNumber="112">H. dilutipes</emphasis>
</taxonomicName>
, differing therefrom by the smaller size (ca.
<quantity id="4CD49B48E368FFFEFEB0F8E9B889F8E1" box="[338,437,1868,1892]" metricMagnitude="-3" metricUnit="m" metricValue="1.73" pageId="111" pageNumber="112" unit="mm" value="1.73">1.73 mm</quantity>
vs. 1.81), the less densely punctate pronotum, the larger elytral punctures, and having the elytra intervals not strongly raised and not forming unbroken lines. The plaques of
<taxonomicName id="4C2C4D2EE368FFFEFBDBF8D4BDE3F80D" authorityName="Perkins" authorityYear="2017" box="[1081,1247,1903,1928]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="111" pageNumber="112" phylum="Arthropoda" rank="species" species="dilutipoides">
<emphasis id="B958EABFE368FFFEFBDBF8D4BDE3F80D" box="[1081,1247,1903,1928]" italics="true" pageId="111" pageNumber="112">H. dilutipoides</emphasis>
</taxonomicName>
are more widely separated, especially posteriorly, and the elytral intervals are not as uniformly raised. Males of both species have large metatibial brushes. The aedeagi also show a relationship, having a large, flat distal extension of the main piece; the complex gonopore bearing distal piece differs significantly in the two species (
<figureCitation id="13172A28E368FFFEFB9FF879BC1EF871" box="[1149,1314,2012,2037]" captionStart-0="FIGURE 100" captionStart-1="FIGURES 101 102" captionStartId-0="113.[151,250,1877,1899]" captionStartId-1="114.[151,264,1648,1671]" captionTargetBox-1="[280,1304,194,1597]" captionTargetId-1="figure@114.[264,1323,179,1628]" captionTargetPageId-1="114" captionText-0="FIGURE 100. Aedeagus of Hydraena dilutipes, lectotype." captionText-1="FIGURES 101 102. Aedeagi of Hydraena dilutipes, non-type, and Hydraena dilutipoides, holotype." httpUri-1="https://zenodo.org/record/1041110/files/figure.png" pageId="111" pageNumber="112">Figs. 100102</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF536625E377FFE1FF75FA00BAE5FA3E" box="[151,985,1445,1467]" pageId="112" pageNumber="113">
<paragraph id="8B9336ADE377FFE1FF75FA00BAE5FA3E" blockId="112.[151,985,1445,1467]" box="[151,985,1445,1467]" pageId="112" pageNumber="113">
<emphasis id="B958EABFE377FFE1FF75FA00B81CFA3F" bold="true" box="[151,288,1445,1467]" pageId="112" pageNumber="113">FIGURE 99.</emphasis>
<taxonomicName id="4C2C4D2EE377FFE1FEC7FA00BB35FA3E" authorityName="Perkins" authorityYear="2017" box="[293,521,1445,1467]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species" species="dilutipoides">
<emphasis id="B958EABFE377FFE1FEC7FA00BB35FA3E" box="[293,521,1445,1467]" italics="true" pageId="112" pageNumber="113">Hydraena dilutipoides</emphasis>
</taxonomicName>
, holotype, habitus dorsal, lateral, and ventral.
</paragraph>
</caption>
<subSubSection id="C3366526E377FFE3FF25FA47BC66F84D" lastPageId="114" lastPageNumber="115" pageId="112" pageNumber="113" type="description">
<paragraph id="8B9336ADE377FFE1FF25FA47BA74F9C6" blockId="112.[151,1436,1506,1999]" pageId="112" pageNumber="113">
<emphasis id="B958EABFE377FFE1FF25FA47B86DFA7E" bold="true" box="[199,337,1506,1531]" pageId="112" pageNumber="113">Description</emphasis>
. Size:
<typeStatus id="5497880FE377FFE1FE41FA47BB39FA7E" box="[419,517,1506,1531]" pageId="112" pageNumber="113" type="holotype">holotype</typeStatus>
(length/width, mm): body (length to elytral apices) 1.73/0.74; head width 0.44; pronotum 0.40/0.57, PA 0.45, PB 0.50; elytra 1.08/0.74. Dorsum dark brown, head darkest; legs brown; maxillary palpi brown to light brown, distal ½ of last palpomere darker.
</paragraph>
<paragraph id="8B9336ADE377FFE1FF25F9EBBA7FF956" blockId="112.[151,1436,1506,1999]" pageId="112" pageNumber="113">Head and pronotum densely moderately coarsely punctate, punctures of pronotum larger than those of head, each puncture with very short recumbent seta; interstices varying from narrow ridges anteriorly and posteriorly to ca. 2xpd on disc, shining. Mentum sparsely finely punctulate, shining; postmentum rugulose, dull. Genae raised, weakly rugulose, dull, with posterior ridge, shining medially.</paragraph>
<paragraph id="8B9336ADE377FFE1FF25F97ABDF5F8BA" blockId="112.[151,1436,1506,1999]" pageId="112" pageNumber="113">Pronotum bisinuate laterally, distinctly emarginate between anterior angle and widest part of pronotum; anterior margin straight behind eyes, emarginate behind frons; PF1 very shallow; PF2 shallow, very shallowly confluent medially forming very weak U-shaped impression; PF3 moderately deep; PF4 absent.</paragraph>
<paragraph id="8B9336ADE377FFE1FF25F8EFBAC7F84A" blockId="112.[151,1436,1506,1999]" pageId="112" pageNumber="113">Elytra weakly arcuate laterally; summit of posterior declivity near distal 2/3; lateral explanate margins moderately narrow; serial punctures round, separated by ca. 1xpd, slightly larger than pronotal punctures. Intervals very weakly raised, not forming distinct unbroken lines, shining, ca. 1xpd. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming shallow angle with one another.</paragraph>
<caption id="DF536625E376FFE0FF75F8F0BBD7F8EE" box="[151,747,1877,1899]" pageId="113" pageNumber="114">
<paragraph id="8B9336ADE376FFE0FF75F8F0BBD7F8EE" blockId="113.[151,747,1877,1899]" box="[151,747,1877,1899]" pageId="113" pageNumber="114">
<emphasis id="B958EABFE376FFE0FF75F8F0B810F8EF" bold="true" box="[151,300,1877,1899]" pageId="113" pageNumber="114">FIGURE 100.</emphasis>
Aedeagus of
<taxonomicName id="4C2C4D2EE376FFE0FE5AF8F0BB46F8EE" box="[440,634,1877,1899]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="113" pageNumber="114" phylum="Arthropoda" rank="species" species="dilutipes">
<emphasis id="B958EABFE376FFE0FE5AF8F0BB46F8EE" box="[440,634,1877,1899]" italics="true" pageId="113" pageNumber="114">Hydraena dilutipes</emphasis>
</taxonomicName>
, lectotype.
</paragraph>
</caption>
<caption id="DF536625E375FFE3FF75F9D5BDA6F902" box="[151,1178,1648,1671]" httpUri="https://zenodo.org/record/1041110/files/figure.png" pageId="114" pageNumber="115" targetBox="[280,1304,194,1597]" targetPageId="114">
<paragraph id="8B9336ADE375FFE3FF75F9D5BDA6F902" blockId="114.[151,1178,1648,1671]" box="[151,1178,1648,1671]" pageId="114" pageNumber="115">
<emphasis id="B958EABFE375FFE3FF75F9D5B851F903" bold="true" box="[151,365,1648,1671]" pageId="114" pageNumber="115">FIGURES 101102.</emphasis>
Aedeagi of
<taxonomicName id="4C2C4D2EE375FFE3FE0BF9D4BB97F902" box="[489,683,1649,1671]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="dilutipes">
<emphasis id="B958EABFE375FFE3FE0BF9D4BB97F902" box="[489,683,1649,1671]" italics="true" pageId="114" pageNumber="115">Hydraena dilutipes</emphasis>
</taxonomicName>
, non-type, and
<taxonomicName id="4C2C4D2EE375FFE3FCA8F9D4BD10F902" authorityName="Perkins" authorityYear="2017" box="[842,1068,1649,1671]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="dilutipoides">
<emphasis id="B958EABFE375FFE3FCA8F9D4BD10F902" box="[842,1068,1649,1671]" italics="true" pageId="114" pageNumber="115">Hydraena dilutipoides</emphasis>
</taxonomicName>
, holotype.
</paragraph>
</caption>
<paragraph id="8B9336ADE375FFE3FF25F916BC66F84D" blockId="114.[151,1437,1715,2028]" pageId="114" pageNumber="115">
Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/1/2/1.3. P1 laminate; median carina sinuate in profile. P2 l/w ca. 2/1, sides parallel or slightly converging to blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques large, widest at anterior 1/3 anteriorly and posteriorly rounded, more sharply so posteriorly, converging slightly anteriorly, not raised, located at sides of median depression. Metaventrite with very short longitudinal ridge on each side, extended posteriorly from margin of each mesocoxal cavity. AIS width at slightly arcuate posterior margin ca.
<date id="FF92106DE375FFE3FE39F8CDB8CBF805" box="[475,503,1896,1920]" pageId="114" pageNumber="115">2x</date>
P2. Protibia moderately wide, slightly arcuate on medial margin, slightly more so on lateral margin. Mesotibia straight. Metatibia slightly arcuate, distal ½ of medial margin emarginate and with large brush of setae. Abdominal apex symmetrical, last tergite with deep and rather wide apicomedian notch.
</paragraph>
</subSubSection>
<subSubSection id="C3366526E375FFE3FF25F876BA8CF86E" box="[199,944,2003,2028]" pageId="114" pageNumber="115" type="etymology">
<paragraph id="8B9336ADE375FFE3FF25F876BA8CF86E" blockId="114.[151,1437,1715,2028]" box="[199,944,2003,2028]" pageId="114" pageNumber="115">
<emphasis id="B958EABFE375FFE3FF25F876B872F869" bold="true" box="[199,334,2003,2028]" pageId="114" pageNumber="115">Etymology.</emphasis>
Named in reference to the relationship to
<taxonomicName id="4C2C4D2EE375FFE3FCC4F870BA94F869" box="[806,936,2003,2028]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="dilutipes">
<emphasis id="B958EABFE375FFE3FCC4F870BA94F869" box="[806,936,2003,2028]" italics="true" pageId="114" pageNumber="115">H. dilutipes</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>