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123 lines
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<document id="17C3464660A2DE746502611DAFB6282F" ID-CLB-Dataset="31639" ID-DOI="10.11646/zootaxa.4342.1.1" ID-GBIF-Dataset="15799d2a-1968-4874-8100-452597d4e26b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1041066" ID-ZooBank="2ACD54D2-3487-432D-9323-EEC131FE2E64" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1509694849742" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2017" docId="038587BBE31AFF8CFF75FBE5B88AF914" docLanguage="en" docName="zootaxa.4342.1.1.pdf.imf" docOrigin="Zootaxa 4342 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydraena Monomadraena" docType="treatment" docVersion="9" lastPageNumber="30" masterDocId="FFBCFFC3E307FF91FFE2FFA5B93CFF85" masterDocTitle="Hydraenidae of Madagascar (Insecta: Coleoptera)" masterLastPageNumber="264" masterPageNumber="1" pageNumber="30" updateTime="1698480684898" updateUser="ExternalLinkService">
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<mods:mods id="AA2123270F8DA1063DEEB88DBC8DBC3D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="E39F8E1B4C6C6CDF0A88A14290B02296">
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<mods:title id="1045021DEA79D1D46B04BD718EC475C9">Hydraenidae of Madagascar (Insecta: Coleoptera)</mods:title>
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</mods:titleInfo>
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<mods:name id="B42418039520A41C23195BEBA33ED37C" type="personal">
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<mods:roleTerm id="1F25F5DC81D8F3AB6CA9CC1FD5EA1880">Author</mods:roleTerm>
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<mods:namePart id="81551A7DB8E7E55B5EC4B976BCEBF783">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="59EA0AF62D5B2632E28A8DC752C7CB51">text</mods:typeOfResource>
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<mods:titleInfo id="06056F2CDC24F4572887C8714815FC8A">
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<mods:title id="0C90496855AD14142CB63D9679659B73">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="1D8E07911A6559602796D4E9753CD776">
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<mods:date id="127CA7E1F77790AD5223C0DAF00B117E">2017</mods:date>
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<mods:detail id="9FBAE1ADAB187E7E8D3DBAA714F653DC" type="pubDate">
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<mods:number id="92AD3F9DFD3AEE0117E4F5E8B6AA143A">2017-11-03</mods:number>
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<mods:detail id="DEE11E51910B51C91E91220EFDCF1AAC" type="volume">
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<mods:number id="6804E4CDD4EE2F66CDC1A8A563F0493C">4342</mods:number>
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</mods:detail>
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<mods:detail id="90E37902BAF31DFA0FDC988411DF42AD" type="issue">
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<mods:number id="B9CC08526FEA56FD2515D73BC8FA5A4F">1</mods:number>
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<mods:classification id="8E15FA3228712141E390E5A651F32521">journal article</mods:classification>
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<mods:identifier id="C6D3F0A06C3C69E0891C241A6D5CE1B0" type="CLB-Dataset">31639</mods:identifier>
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<mods:identifier id="CB806BE6F37B0E66D46F38B7BFCE2C7A" type="DOI">10.11646/zootaxa.4342.1.1</mods:identifier>
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<mods:identifier id="BD23341B8AD40A3FC8E9FA00BBC8336F" type="GBIF-Dataset">15799d2a-1968-4874-8100-452597d4e26b</mods:identifier>
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<mods:identifier id="1B5DAD52B354B4F3BEE050275D21414A" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="ABB289487156CB42D0CE01C80E7355A2" type="Zenodo-Dep">1041066</mods:identifier>
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<mods:identifier id="33C7AAFE39B7421A2B420122004593F5" type="ZooBank">2ACD54D2-3487-432D-9323-EEC131FE2E64</mods:identifier>
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</mods:mods>
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<treatment id="038587BBE31AFF8CFF75FBE5B88AF914" ID-DOI="http://doi.org/10.5281/zenodo.5323403" ID-GBIF-Taxon="137233414" ID-Zenodo-Dep="5323403" LSID="urn:lsid:plazi:treatment:038587BBE31AFF8CFF75FBE5B88AF914" httpUri="http://treatment.plazi.org/id/038587BBE31AFF8CFF75FBE5B88AF914" lastPageNumber="30" pageId="29" pageNumber="30">
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<subSubSection id="C3366526E31AFF8CFF75FBE5BB94FBDF" box="[151,680,1088,1114]" pageId="29" pageNumber="30" type="nomenclature">
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<paragraph id="8B9336ADE31AFF8CFF75FBE5BB94FBDF" blockId="29.[151,680,1088,1114]" box="[151,680,1088,1114]" pageId="29" pageNumber="30">
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<heading id="D0DB81C1E31AFF8CFF75FBE5BB94FBDF" bold="true" box="[151,680,1088,1114]" fontSize="11" level="1" pageId="29" pageNumber="30" reason="1">
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<emphasis id="B958EABFE31AFF8CFF75FBE5BB94FBDF" bold="true" box="[151,680,1088,1114]" pageId="29" pageNumber="30">
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<taxonomicName id="4C2C4D2EE31AFF8CFF75FBE5B8D0FBDF" ID-CoL="8VRR8" authority="(Monomadraena)" authorityName="Monomadraena" box="[151,492,1088,1114]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
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<emphasis id="B958EABFE31AFF8CFF75FBE5B82CFBDF" bold="true" box="[151,272,1088,1114]" italics="true" pageId="29" pageNumber="30">Hydraena</emphasis>
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(
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<emphasis id="B958EABFE31AFF8CFEC2FBE5B8DEFBDF" bold="true" box="[288,482,1088,1114]" italics="true" pageId="29" pageNumber="30">Monomadraena</emphasis>
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)
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</taxonomicName>
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, new subgenus
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3366526E31AFF8CFF75FB22BB60FAF4" pageId="29" pageNumber="30" type="materials_examined">
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<paragraph id="8B9336ADE31AFF8CFF75FB22BA77FB19" blockId="29.[151,843,1158,1180]" box="[151,843,1158,1180]" pageId="29" pageNumber="30">
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Type Species:
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<taxonomicName id="4C2C4D2EE31AFF8CFEC8FB23BBF7FB19" authority="Perkins, 2017" authorityName="Perkins" authorityYear="2017" box="[298,715,1158,1180]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="tibiodentipes" status="sp. nov." subGenus="Monomadraena">
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<emphasis id="B958EABFE31AFF8CFEC8FB23B8B3FB19" box="[298,399,1158,1180]" italics="true" pageId="29" pageNumber="30">Hydraena</emphasis>
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(
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<emphasis id="B958EABFE31AFF8CFE7CFB23BB7CFB19" box="[414,576,1158,1180]" italics="true" pageId="29" pageNumber="30">Monomadraena</emphasis>
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)
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<emphasis id="B958EABFE31AFF8CFDAAFB23BBF7FB19" box="[584,715,1158,1180]" italics="true" pageId="29" pageNumber="30">tibiodentipes</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A26B57C4E31AFF8CFD33FB22BA77FB19" box="[721,843,1158,1180]" pageId="29" pageNumber="30" rank="species">new species</taxonomicNameLabel>
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</paragraph>
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<paragraph id="8B9336ADE31AFF8CFF75FB6CBB60FAF4" blockId="29.[151,1437,1225,1682]" pageId="29" pageNumber="30">
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This new subgenus comprises 45 species, which are placed in six species groups. Two of these species,
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<taxonomicName id="4C2C4D2EE31AFF8CFA63FB6FB80EFA83" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="impressicollis">
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<emphasis id="B958EABFE31AFF8CFA63FB6FB80EFA83" italics="true" pageId="29" pageNumber="30">H. impressicollis</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C2C4D2EE31AFF8CFE8EFB4BBB33FA83" authorityName="Perkins" authorityYear="2017" box="[364,527,1261,1286]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="pentarubra">
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<emphasis id="B958EABFE31AFF8CFE8EFB4BBB33FA83" box="[364,527,1261,1286]" italics="true" pageId="29" pageNumber="30">H. pentarubra</emphasis>
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</taxonomicName>
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, were included in the molecular phylogeny of
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<bibRefCitation id="EFBD4B5CE31AFF8CFBCCFB48BC19FA83" box="[1070,1317,1261,1286]" pageId="29" pageNumber="30" refString="Trizzino, M., Jach, M. A., Audisio, P., Alonso, R. & Ribera, I. (2012) A molecular phylogeny of the cosmopolitan hyperdiverse genus Hydraena (Coleoptera, Hydraenidae). Systematic Entomology, 38 (1), 192 - 208. https: // doi. org / 10.1111 / j. 1365 - 3113.2012.00654. x" type="journal article">
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Trizzino
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<emphasis id="B958EABFE31AFF8CFB77FB4BBDECFA83" box="[1173,1232,1261,1286]" italics="true" pageId="29" pageNumber="30">et al.</emphasis>
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(2012)
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</bibRefCitation>
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, and were found to be located near the base of
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<taxonomicName id="4C2C4D2EE31AFF8CFDD1FAB4BB9FFAAF" box="[563,675,1297,1322]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
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<emphasis id="B958EABFE31AFF8CFDD1FAB4BB9FFAAF" box="[563,675,1297,1322]" italics="true" pageId="29" pageNumber="30">Hydraena</emphasis>
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</taxonomicName>
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(see
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<figureCitation id="13172A28E31AFF8CFD02FAB4BA7FFAAF" box="[736,835,1297,1322]" captionStart="FIGURES 344 – 345. 344" captionStartId="261.[151,264,1715,1738]" captionTargetBox="[266,1317,241,1662]" captionTargetId="figure@261.[226,1360,193,1694]" captionTargetPageId="261" captionText="FIGURES 344 – 345. 344. Modified basal portion of DNA sequence data phylogeny of Hydraena, showing identity of Madagascar species, and new subgenera. Used by permission of Ignacio Ribera. 345. Three examples of aedeagi having very short parameres that originate near the basal orifice, and have only one or two setae." httpUri="https://zenodo.org/record/1041202/files/figure.png" pageId="29" pageNumber="30">Fig. 344</figureCitation>
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). Paratypes of several of the species described herein have been retained in 100% ethanol, so future DNA sequence studies will be possible, to give a more complete picture of the basal groups of
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<taxonomicName id="4C2C4D2EE31AFF8CFE07FAFCBB69FAF7" box="[485,597,1369,1394]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
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<emphasis id="B958EABFE31AFF8CFE07FAFCBB69FAF7" box="[485,597,1369,1394]" italics="true" pageId="29" pageNumber="30">Hydraena</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3366526E31AFF8CFF25FAD8B9D4F9CF" pageId="29" pageNumber="30" type="diagnosis">
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<paragraph id="8B9336ADE31AFF8CFF25FAD8B9D4F9CF" blockId="29.[151,1437,1225,1682]" pageId="29" pageNumber="30">
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<emphasis id="B958EABFE31AFF8CFF25FAD8B87DFA13" bold="true" box="[199,321,1405,1430]" pageId="29" pageNumber="30">Diagnosis.</emphasis>
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In addition to the usual diagnostic characters of
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<taxonomicName id="4C2C4D2EE31AFF8CFC6AFAD8BAC4FA13" box="[904,1016,1405,1430]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
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<emphasis id="B958EABFE31AFF8CFC6AFAD8BAC4FA13" box="[904,1016,1405,1430]" italics="true" pageId="29" pageNumber="30">Hydraena</emphasis>
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</taxonomicName>
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, with the following combination of characters: head with genae raised, with a posterior ridge, usually interrupted in midline; metaventrite with an anteromedian carina that attains the anterior margin of the plaques, frequently joining the plaques, together forming an inverted Y-shape; males with modified mesotibiae, having a row of strong spines along the inner margin (e.g.,
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<figureCitation id="13172A28E31AFF8CFF75F9A8B9D7F9A3" box="[151,235,1549,1574]" captionStart="FIGURE 57" captionStartId="68.[151,250,1215,1237]" captionText="FIGURE 57. Hydraena tibiodentipes, holotype, habitus dorsal, lateral, and ventral." pageId="29" pageNumber="30">Fig. 57</figureCitation>
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); and the aedeagus having only one paramere, on the right side, which is long and originates near the basal orifice.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3366526E31AFF8CFF25F9F0B88AF914" pageId="29" pageNumber="30" type="etymology">
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<paragraph id="8B9336ADE31AFF8CFF25F9F0B88AF914" blockId="29.[151,1437,1225,1682]" pageId="29" pageNumber="30">
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<emphasis id="B958EABFE31AFF8CFF25F9F0B872F9EB" bold="true" box="[199,334,1621,1646]" pageId="29" pageNumber="30">Etymology.</emphasis>
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Named in reference to the single paramere, the geographical distribution (
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<collectingCountry id="F33B763DE31AFF8CFB2CF9F0BC67F9EB" box="[1230,1371,1621,1646]" name="Madagascar" pageId="29" pageNumber="30">Madagascar</collectingCountry>
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), and relationship to
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<taxonomicName id="4C2C4D2EE31AFF8CFEDDF9DCB893F917" box="[319,431,1657,1682]" class="Insecta" family="Hydraenidae" genus="Hydraena" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
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<emphasis id="B958EABFE31AFF8CFEDDF9DCB893F917" box="[319,431,1657,1682]" italics="true" pageId="29" pageNumber="30">Hydraena</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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