661 lines
83 KiB
XML
661 lines
83 KiB
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<document id="E4359E4F4BEDBB54D279A7EDE9CA7166" ID-CLB-Dataset="45868" ID-DOI="10.5281/zenodo.207160" ID-GBIF-Dataset="8666c9de-6e79-487e-a849-c65c91bccb90" ID-ISSN="1175-5326" ID-Zenodo-Dep="207160" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460415612743" checkinUser="plazi" docAuthor="Serrano, Artur R. M. & Aguiar, Carlos A. S." docDate="2011" docId="038587ACF720932AFF021DB617ACF93D" docLanguage="en" docName="zt03116p046.pdf" docOrigin="Zootaxa 3116" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Geocharis antheroi Serrano & Aguiar, new species" docType="treatment" docVersion="10" lastPageNumber="45" masterDocId="FFBCFFD4F7259326FF951D211619FFDD" masterDocTitle="Two new species of the genus Geocharis Ehlers, 1883 and new data on Anillina species from Portugal (Coleoptera: Carabidae)" masterLastPageNumber="46" masterPageNumber="33" pageNumber="38" updateTime="1698328697478" updateUser="ExternalLinkService">
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<mods:mods id="386553A64221E3529BE8BEC707B9DA3D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="DBE6538116F26D0BAF369306BFF1B6C6">
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<mods:title id="2FE3F212C63B22C954A5972F8ED39FD4">Two new species of the genus Geocharis Ehlers, 1883 and new data on Anillina species from Portugal (Coleoptera: Carabidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="53BF296A3DCC9EFFF3C90906A68D92DB" type="personal">
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</mods:role>
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<mods:namePart id="259D21358302608F12811476FEB13419">Serrano, Artur R. M.</mods:namePart>
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</mods:name>
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<mods:name id="98875A2031C116EF7DA792B22C04C509" type="personal">
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<mods:roleTerm id="FF93AAF428363676E1C929B22D52562C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2B9543BE20D77A5766C8C8CFC7D0370C">Aguiar, Carlos A. S.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="5F3E5AD40919249155A06BFA9768D116">text</mods:typeOfResource>
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<mods:title id="C6735EDA3D5F3EE679BB8EAD2B165ABC">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="31ED59864A3AF67F75617C6629B91172">
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<mods:date id="85830225A94D8D305B50D06346B074C9">2011</mods:date>
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<mods:detail id="0F39CA09944012E6351AC9449FD5E98B" type="volume">
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<mods:number id="17AEF7EA77FE9A7E1374C173DCCB166C">3116</mods:number>
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</mods:detail>
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<mods:start id="0A4837F8F5B4BCEBC9F4082B760967CA">33</mods:start>
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<mods:classification id="A9E8F2D6C6BDFA7F93DD6EE67E296253">journal article</mods:classification>
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<mods:identifier id="3E9DAD69BCCFC9D40E0570925DB44E53" type="CLB-Dataset">45868</mods:identifier>
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<mods:identifier id="BFAA54AC6276BCE6CA86EA632F89EB1D" type="DOI">10.5281/zenodo.207160</mods:identifier>
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<mods:identifier id="FB6685FE60B14143C93578CBF649C4C7" type="GBIF-Dataset">8666c9de-6e79-487e-a849-c65c91bccb90</mods:identifier>
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<mods:identifier id="724C015D88392A0E6FA929A2A3DF7C63" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="35C777D6C9F64B3EF1FFFD366750F43D" type="Zenodo-Dep">207160</mods:identifier>
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<treatment id="038587ACF720932AFF021DB617ACF93D" ID-DOI="http://doi.org/10.5281/zenodo.6186161" ID-GBIF-Taxon="119424899" ID-Zenodo-Dep="6186161" LSID="urn:lsid:plazi:treatment:038587ACF720932AFF021DB617ACF93D" httpUri="http://treatment.plazi.org/id/038587ACF720932AFF021DB617ACF93D" lastPageId="12" lastPageNumber="45" pageId="5" pageNumber="38">
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<subSubSection id="C3366531F7209323FF021DB617B8FF0F" pageId="5" pageNumber="38" type="nomenclature">
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<paragraph id="8B9336BAF7209323FF021DB6151EFF6C" blockId="5.[151,775,151,210]" box="[151,775,151,177]" pageId="5" pageNumber="38">
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<heading id="D0DB81D6F7209323FF021DB6151EFF6C" bold="true" box="[151,775,151,177]" fontSize="11" level="1" pageId="5" pageNumber="38" reason="1">
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<emphasis id="B958EAA8F7209323FF021DB6151EFF6C" bold="true" box="[151,775,151,177]" pageId="5" pageNumber="38">
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<taxonomicName id="4C2C4D39F7209323FF021DB61473FF6C" ID-CoL="3FNZ3" authority="Serrano & Aguiar" authorityName="Serrano & Aguiar" box="[151,618,151,177]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="38" phylum="Arthropoda" rank="species" species="antheroi" status="sp. nov.">
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<emphasis id="B958EAA8F7209323FF021DB61799FF6C" bold="true" box="[151,384,151,177]" italics="true" pageId="5" pageNumber="38">Geocharis antheroi</emphasis>
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Serrano & Aguiar
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</taxonomicName>
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,
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<taxonomicNameLabel id="A26B57D3F7209323FDE21DB6151EFF6C" box="[631,775,151,177]" pageId="5" pageNumber="38" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9336BAF7209323FF021D9B17B8FF0F" blockId="5.[151,775,151,210]" box="[151,417,186,210]" pageId="5" pageNumber="38">
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(
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<figureCitation id="13172A3FF7209323FF0A1D9B16FCFF0F" box="[159,229,186,210]" captionStart="FIGURE 4" captionStartId="6.[151,250,1916,1939]" captionTargetBox="[172,1414,183,1910]" captionTargetId="figure@6.[172,1414,183,1933]" captionTargetPageId="6" captionText="FIGURE 4. SEM photographs of structural features of Geocharis antheroi n. sp., (a), head (latero – dorsal view); (b), head (ventral view); (c), pronotum (dorsal view); (d), elytra (dorsal view); (e), right elytron (latero-dorsal view); (f), anterior half of elytra (dorsal view); (g), thorax and abdomen (male, ventral view); (h), thorax and abdomen (female, ventral view)." httpUri="https://zenodo.org/record/207164/files/figure.png" pageId="5" pageNumber="38">Figs 4</figureCitation>
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a–4h, 5c–5d, 6b)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3366531F7209323FF021C221292FEBF" pageId="5" pageNumber="38" type="materials_examined">
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<paragraph id="8B9336BAF7209323FF021C221292FEBF" blockId="5.[151,1437,258,2010]" pageId="5" pageNumber="38">
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<emphasis id="B958EAA8F7209323FF021C221751FEC6" bold="true" box="[151,328,258,283]" pageId="5" pageNumber="38">
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<typeStatus id="54978818F7209323FF021C2216CBFEC6" box="[151,210,259,283]" pageId="5" pageNumber="38">Type</typeStatus>
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material:
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</emphasis>
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<typeStatus id="54978818F7209323FEDA1C2317A3FEC7" box="[335,442,258,282]" pageId="5" pageNumber="38" type="holotype">Holotype</typeStatus>
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, 3,
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<collectingCountry id="F33B762AF7209323FE7E1C231454FEC7" box="[491,589,258,282]" name="Portugal" pageId="5" pageNumber="38">Portugal</collectingCountry>
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, Carção (Vimioso) (U.T.M. coordinate: 29TQG0010),
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<date id="FF92107AF7209323FB221C23132CFEC7" box="[1207,1333,258,282]" pageId="5" pageNumber="38" value="2010-04-22">22.IV.2010</date>
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, (A. Serrano leg.).
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<typeStatus id="54978818F7209323FE851C061799FEE2" box="[272,384,295,319]" pageId="5" pageNumber="38" type="paratype">Paratypes</typeStatus>
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, 8 3 and 12 Ƥ (23, 1Ƥ gold coated), same locality of
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<typeStatus id="54978818F7209323FC751C07125CFEE3" box="[992,1093,294,318]" pageId="5" pageNumber="38" type="holotype">holotype</typeStatus>
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,
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<date id="FF92107AF7209323FBDA1C0712D7FEE3" box="[1103,1230,294,318]" pageId="5" pageNumber="38" value="2010-04-22">22.IV.2010</date>
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, (A. Serrano leg.), 1Ƥ, Santulhão (Vimioso) (U.T.M. coordinate: 29TPG9806),
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<date id="FF92107AF7209323FCAB1C6B15A5FEBF" box="[830,956,330,354]" pageId="5" pageNumber="38" value="2010-04-22">22.IV.2010</date>
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, (A. Serrano leg.).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3366531F7209323FF501C4F1518FDCB" pageId="5" pageNumber="38" type="diagnosis">
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<paragraph id="8B9336BAF7209323FF501C4F1518FDCB" blockId="5.[151,1437,258,2010]" pageId="5" pageNumber="38">
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<emphasis id="B958EAA8F7209323FF501C4F1726FE5A" bold="true" box="[197,319,366,391]" pageId="5" pageNumber="38">Diagnosis.</emphasis>
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Anophtalmous; body slightly depressed, brown with integument microreticulated. Sparse pubescence mainly on pronotum and elytra. Elytron with vestigial striae, humeral region strongly punctured, disk with one anterior and one posterior seta. Male forelegs with the first tarsomere dilated. Males and females without a median tooth on the internal margin of the metafemora. Mesotibiae with dense pubescence in both margins.
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<collectingCountry id="F33B762AF7209323FAF11CFB1385FE2F" box="[1380,1436,474,498]" name="India" pageId="5" pageNumber="38">Hind</collectingCountry>
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tibiae more or less straight. Aedeagus as in
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<figureCitation id="13172A3FF7209323FDEB1CDF14DAFDCB" box="[638,707,510,534]" captionStart="FIGURE 5" captionStartId="7.[151,250,1954,1977]" captionTargetBox="[184,1380,185,1953]" captionTargetId="figure@7.[158,1441,147,1966]" captionTargetPageId="7" captionText="FIGURE 5. Geocharis barcorabelo n. sp. aedeagus, (a), median lobe and left paramere in lateral view; (b), apex of median lobe in dorsal view. Geocharis antheroi n. sp. aedeagus, (c), median lobe and left paramere in lateral view; (d), apex of median lobe in dorsal view." httpUri="https://zenodo.org/record/207165/files/figure.png" pageId="5" pageNumber="38">Figs 5</figureCitation>
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c–5d.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3366531F7209323FF501F031527F807" pageId="5" pageNumber="38" type="description">
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<paragraph id="8B9336BAF7209323FF501F031312FDE7" blockId="5.[151,1437,258,2010]" box="[197,1291,546,571]" pageId="5" pageNumber="38">
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<emphasis id="B958EAA8F7209323FF501F03174EFDE6" bold="true" box="[197,343,546,571]" pageId="5" pageNumber="38">Description.</emphasis>
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Length of
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<typeStatus id="54978818F7209323FE471F031421FDE7" box="[466,568,546,570]" pageId="5" pageNumber="38" type="holotype">holotype</typeStatus>
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:
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<quantity id="4CD49B5FF7209323FDD71F031482FDE7" box="[578,667,546,570]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="5" pageNumber="38" unit="mm" value="1.9">1.9 mm</quantity>
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. Length of
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<typeStatus id="54978818F7209323FC8E1F021592FDE6" box="[795,907,547,571]" pageId="5" pageNumber="38" type="paratype">paratypes</typeStatus>
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:
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<quantity id="4CD49B5FF7209323FC011F031202FDE7" box="[916,1051,546,570]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="1.9" metricValueMin="1.6" pageId="5" pageNumber="38" unit="mm" value="1.75" valueMax="1.9" valueMin="1.6">1.6–1.9 mm</quantity>
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(males and females).
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</paragraph>
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<paragraph id="8B9336BAF7209323FF581F6715F2FCCE" blockId="5.[151,1437,258,2010]" pageId="5" pageNumber="38">
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Head (
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<figureCitation id="13172A3FF7209323FE8D1F671744FD83" box="[280,349,582,606]" captionStart="FIGURE 4" captionStartId="6.[151,250,1916,1939]" captionTargetBox="[172,1414,183,1910]" captionTargetId="figure@6.[172,1414,183,1933]" captionTargetPageId="6" captionText="FIGURE 4. SEM photographs of structural features of Geocharis antheroi n. sp., (a), head (latero – dorsal view); (b), head (ventral view); (c), pronotum (dorsal view); (d), elytra (dorsal view); (e), right elytron (latero-dorsal view); (f), anterior half of elytra (dorsal view); (g), thorax and abdomen (male, ventral view); (h), thorax and abdomen (female, ventral view)." httpUri="https://zenodo.org/record/207164/files/figure.png" pageId="5" pageNumber="38">Fig. 4</figureCitation>
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a) more or less as long as wide [length:
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<quantity id="4CD49B5FF7209323FCB41F6715DCFD83" box="[801,965,582,606]" metricMagnitude="-4" metricUnit="m" metricValue="3.3999999999999995" metricValueMax="3.7" metricValueMin="3.1" pageId="5" pageNumber="38" unit="mm" value="0.33999999999999997" valueMax="0.37" valueMin="0.31">0.31–0.37 mm</quantity>
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(males),
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<quantity id="4CD49B5FF7209323FBBA1F6712CAFD83" box="[1071,1235,582,606]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" metricValueMax="3.7" metricValueMin="2.7" pageId="5" pageNumber="38" unit="mm" value="0.32" valueMax="0.37" valueMin="0.27">0.27–0.37 mm</quantity>
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(females); width: 0.32–0.38 (males),
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<quantity id="4CD49B5FF7209323FEE21F4B1404FD5F" box="[375,541,618,642]" metricMagnitude="-4" metricUnit="m" metricValue="3.55" metricValueMax="3.8" metricValueMin="3.3" pageId="5" pageNumber="38" unit="mm" value="0.355" valueMax="0.38" valueMin="0.33">0.33–0.38 mm</quantity>
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(females)], microsculpture distinct. Cephalic chaetotaxy (fixed setae of right side): L3+C1+F1+SA1+SP1+
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<date id="FF92107AF7209323FE721FAF141EFD7B" box="[487,519,654,678]" pageId="5" pageNumber="38">V1</date>
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+O1+P0+G2. Frontal and clypeal setae inserted in two large and two small sulci, respectively. Antennae light brown, antennomeres 1–2 longer than the others, the latter subpyriform, the third and the fourth are the shortest ones and subpyriforms, antennomeres 5–10 gradually longer and oval-shaped, the last one acuminate. Mouth-parts (
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<figureCitation id="13172A3FF7209323FE741FDB143AFCCF" box="[481,547,762,786]" captionStart="FIGURE 4" captionStartId="6.[151,250,1916,1939]" captionTargetBox="[172,1414,183,1910]" captionTargetId="figure@6.[172,1414,183,1933]" captionTargetPageId="6" captionText="FIGURE 4. SEM photographs of structural features of Geocharis antheroi n. sp., (a), head (latero – dorsal view); (b), head (ventral view); (c), pronotum (dorsal view); (d), elytra (dorsal view); (e), right elytron (latero-dorsal view); (f), anterior half of elytra (dorsal view); (g), thorax and abdomen (male, ventral view); (h), thorax and abdomen (female, ventral view)." httpUri="https://zenodo.org/record/207164/files/figure.png" pageId="5" pageNumber="38">Fig. 4</figureCitation>
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b) show the general pattern of the genus.
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</paragraph>
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<paragraph id="8B9336BAF7209323FF501E3E152FFBEF" blockId="5.[151,1437,258,2010]" pageId="5" pageNumber="38">
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Pronotum cordiform (
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<figureCitation id="13172A3FF7209323FE2D1E3F17E5FCEB" box="[440,508,798,822]" captionStart="FIGURE 4" captionStartId="6.[151,250,1916,1939]" captionTargetBox="[172,1414,183,1910]" captionTargetId="figure@6.[172,1414,183,1933]" captionTargetPageId="6" captionText="FIGURE 4. SEM photographs of structural features of Geocharis antheroi n. sp., (a), head (latero – dorsal view); (b), head (ventral view); (c), pronotum (dorsal view); (d), elytra (dorsal view); (e), right elytron (latero-dorsal view); (f), anterior half of elytra (dorsal view); (g), thorax and abdomen (male, ventral view); (h), thorax and abdomen (female, ventral view)." httpUri="https://zenodo.org/record/207164/files/figure.png" pageId="5" pageNumber="38">Fig. 4</figureCitation>
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c), transverse, about 1.2–1.3 times wider than long [length:
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<quantity id="4CD49B5FF7209323FB0C1E3F1322FCEB" box="[1177,1339,798,822]" metricMagnitude="-4" metricUnit="m" metricValue="3.9" metricValueMax="4.3" metricValueMin="3.5" pageId="5" pageNumber="38" unit="mm" value="0.39" valueMax="0.43" valueMin="0.35">0.35–0.43 mm</quantity>
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(males),
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<quantity id="4CD49B5FF7209323FF021E631725FC87" box="[151,316,834,858]" metricMagnitude="-4" metricUnit="m" metricValue="3.75" metricValueMax="4.1" metricValueMin="3.4" pageId="5" pageNumber="38" unit="mm" value="0.375" valueMax="0.41" valueMin="0.34">0.34–0.41 mm</quantity>
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(females); width:
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<quantity id="4CD49B5FF7209323FD9B1E6314AAFC87" box="[526,691,834,858]" metricMagnitude="-4" metricUnit="m" metricValue="4.699999999999999" metricValueMax="5.1" metricValueMin="4.3" pageId="5" pageNumber="38" unit="mm" value="0.47" valueMax="0.51" valueMin="0.43">0.43–0.51 mm</quantity>
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(males),
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|
|
<quantity id="4CD49B5FF7209323FC8A1E6315DDFC87" box="[799,964,834,858]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" metricValueMax="5.0" metricValueMin="4.2" pageId="5" pageNumber="38" unit="mm" value="0.45999999999999996" valueMax="0.5" valueMin="0.42">0.42–0.50 mm</quantity>
|
|||
|
|
(females)]; anterior angles not produced, widely rounded off, lateral channel not recurved inside of anterior angles; disk slightly convex, depressed between the two basal pits, with a superficial central sulcus which does not reach the anterior margin; anterior margin slightly straight and posterior margin slightly arcuate outwards; lateral margins with two or three slight denticles just before the posterior angles, which are right and dentate. Vestiture (pubescence): surface covered with scattered erect pubescence; one seta on the lateral margin in the broadest part of the pronotum, another one near the posterior angle; two additional setae inserted near the anterior angles.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B9336BAF7209323FF50191F141CFAAB" blockId="5.[151,1437,258,2010]" pageId="5" pageNumber="38">
|
|||
|
|
Elytra (
|
|||
|
|
<figureCitation id="13172A3FF7209323FE8C191F177FFB8B" box="[281,358,1086,1110]" captionStart="FIGURE 4" captionStartId="6.[151,250,1916,1939]" captionTargetBox="[172,1414,183,1910]" captionTargetId="figure@6.[172,1414,183,1933]" captionTargetPageId="6" captionText="FIGURE 4. SEM photographs of structural features of Geocharis antheroi n. sp., (a), head (latero – dorsal view); (b), head (ventral view); (c), pronotum (dorsal view); (d), elytra (dorsal view); (e), right elytron (latero-dorsal view); (f), anterior half of elytra (dorsal view); (g), thorax and abdomen (male, ventral view); (h), thorax and abdomen (female, ventral view)." httpUri="https://zenodo.org/record/207164/files/figure.png" pageId="5" pageNumber="38">Figs. 4</figureCitation>
|
|||
|
|
d–4e) 1.7–1.8 times longer than wide [length:
|
|||
|
|
<quantity id="4CD49B5FF7209323FCE0191F1201FB8B" box="[885,1048,1086,1110]" metricMagnitude="-3" metricUnit="m" metricValue="1.04" metricValueMax="1.12" metricValueMin="0.96" pageId="5" pageNumber="38" unit="mm" value="1.04" valueMax="1.12" valueMin="0.96">0.96–1.12 mm</quantity>
|
|||
|
|
(males),
|
|||
|
|
<quantity id="4CD49B5FF7209323FB14191F133DFB8B" box="[1153,1316,1086,1110]" metricMagnitude="-3" metricUnit="m" metricValue="1.01" metricValueMax="1.08" metricValueMin="0.94" pageId="5" pageNumber="38" unit="mm" value="1.01" valueMax="1.08" valueMin="0.94">0.94–1.08 mm</quantity>
|
|||
|
|
(females), width:
|
|||
|
|
<quantity id="4CD49B5FF7209323FF7219431792FBA7" box="[231,395,1122,1146]" metricMagnitude="-4" metricUnit="m" metricValue="5.85" metricValueMax="6.3" metricValueMin="5.4" pageId="5" pageNumber="38" unit="mm" value="0.585" valueMax="0.63" valueMin="0.54">0.54–0.63 mm</quantity>
|
|||
|
|
(males and females)], slightly convex, parallel and oval posteriorly, with vestigial traces of striae. Tegument microsculptured, disk more punctured in the shoulders, punctures sparser and disappearing to apex (
|
|||
|
|
<figureCitation id="13172A3FF7209323FF4E198B173EFB1F" box="[219,295,1194,1218]" captionStart="FIGURE 4" captionStartId="6.[151,250,1916,1939]" captionTargetBox="[172,1414,183,1910]" captionTargetId="figure@6.[172,1414,183,1933]" captionTargetPageId="6" captionText="FIGURE 4. SEM photographs of structural features of Geocharis antheroi n. sp., (a), head (latero – dorsal view); (b), head (ventral view); (c), pronotum (dorsal view); (d), elytra (dorsal view); (e), right elytron (latero-dorsal view); (f), anterior half of elytra (dorsal view); (g), thorax and abdomen (male, ventral view); (h), thorax and abdomen (female, ventral view)." httpUri="https://zenodo.org/record/207164/files/figure.png" pageId="5" pageNumber="38">Figs. 4</figureCitation>
|
|||
|
|
d–4f); lateral margin narrow, finely serrate from the humeral angles, which are rounded, to the seventh seta of the umbilicate series. Vestiture: part of the pubescence of the disk is arranged in six irregular lines, these setae are erect and slightly directed anteriad (
|
|||
|
|
<figureCitation id="13172A3FF7209323FD0719D314CEFAD7" box="[658,727,1266,1290]" captionStart="FIGURE 4" captionStartId="6.[151,250,1916,1939]" captionTargetBox="[172,1414,183,1910]" captionTargetId="figure@6.[172,1414,183,1933]" captionTargetPageId="6" captionText="FIGURE 4. SEM photographs of structural features of Geocharis antheroi n. sp., (a), head (latero – dorsal view); (b), head (ventral view); (c), pronotum (dorsal view); (d), elytra (dorsal view); (e), right elytron (latero-dorsal view); (f), anterior half of elytra (dorsal view); (g), thorax and abdomen (male, ventral view); (h), thorax and abdomen (female, ventral view)." httpUri="https://zenodo.org/record/207164/files/figure.png" pageId="5" pageNumber="38">Fig. 4</figureCitation>
|
|||
|
|
e); umbilicate series follows the pattern of the genus. The longest setae of this series are the 2nd, the 6th and the 9th with the 3rd, 5th, 7th and 8th, more slightly inserted within the elytral margin; besides these setae there are one parascutellar basad, two discal (one anterior and one posterior) and one apical seta (
|
|||
|
|
<figureCitation id="13172A3FF7209323FEEF187F17D8FAAB" box="[378,449,1374,1398]" captionStart="FIGURE 4" captionStartId="6.[151,250,1916,1939]" captionTargetBox="[172,1414,183,1910]" captionTargetId="figure@6.[172,1414,183,1933]" captionTargetPageId="6" captionText="FIGURE 4. SEM photographs of structural features of Geocharis antheroi n. sp., (a), head (latero – dorsal view); (b), head (ventral view); (c), pronotum (dorsal view); (d), elytra (dorsal view); (e), right elytron (latero-dorsal view); (f), anterior half of elytra (dorsal view); (g), thorax and abdomen (male, ventral view); (h), thorax and abdomen (female, ventral view)." httpUri="https://zenodo.org/record/207164/files/figure.png" pageId="5" pageNumber="38">Figs 4</figureCitation>
|
|||
|
|
d–4f).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B9336BAF7209323FF5018A2143EFA63" blockId="5.[151,1437,258,2010]" pageId="5" pageNumber="38">
|
|||
|
|
Last abdominal ventrite with one pair of medium sub-marginal setae in males, two pairs of medium sub-marginal setae in females (
|
|||
|
|
<figureCitation id="13172A3FF7209323FE0D188717C6FA63" box="[408,479,1446,1470]" captionStart="FIGURE 4" captionStartId="6.[151,250,1916,1939]" captionTargetBox="[172,1414,183,1910]" captionTargetId="figure@6.[172,1414,183,1933]" captionTargetPageId="6" captionText="FIGURE 4. SEM photographs of structural features of Geocharis antheroi n. sp., (a), head (latero – dorsal view); (b), head (ventral view); (c), pronotum (dorsal view); (d), elytra (dorsal view); (e), right elytron (latero-dorsal view); (f), anterior half of elytra (dorsal view); (g), thorax and abdomen (male, ventral view); (h), thorax and abdomen (female, ventral view)." httpUri="https://zenodo.org/record/207164/files/figure.png" pageId="5" pageNumber="38">
|
|||
|
|
Figs
|
|||
|
|
<quantity id="4CD49B5FF7209323FE45188717C6FA63" box="[464,479,1446,1470]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="5" pageNumber="38" unit="g" value="4.0">4</quantity>
|
|||
|
|
</figureCitation>
|
|||
|
|
g–4h).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B9336BAF7209323FF5018EB146DF9F7" blockId="5.[151,1437,258,2010]" pageId="5" pageNumber="38">
|
|||
|
|
Male legs with the protarsomere 1 dilated; tarsomere
|
|||
|
|
<quantity id="4CD49B5FF7209323FCB618EB1556FA3F" box="[803,847,1482,1506]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="5" pageNumber="38" unit="in" value="1.0">1 in</quantity>
|
|||
|
|
all legs more pigmented (light brown) than the others; mesotibiae with a strong pubescence on both margins; hind femora (males and females) without any median tooth on the internal margin (
|
|||
|
|
<figureCitation id="13172A3FF7209323FE4B1B331435F9F7" box="[478,556,1554,1578]" captionStart="FIGURE 4" captionStartId="6.[151,250,1916,1939]" captionTargetBox="[172,1414,183,1910]" captionTargetId="figure@6.[172,1414,183,1933]" captionTargetPageId="6" captionText="FIGURE 4. SEM photographs of structural features of Geocharis antheroi n. sp., (a), head (latero – dorsal view); (b), head (ventral view); (c), pronotum (dorsal view); (d), elytra (dorsal view); (e), right elytron (latero-dorsal view); (f), anterior half of elytra (dorsal view); (g), thorax and abdomen (male, ventral view); (h), thorax and abdomen (female, ventral view)." httpUri="https://zenodo.org/record/207164/files/figure.png" pageId="5" pageNumber="38">
|
|||
|
|
Figs.
|
|||
|
|
<quantity id="4CD49B5FF7209323FD881B331435F9F7" box="[541,556,1554,1578]" metricMagnitude="-3" metricUnit="kg" metricValue="4.0" pageId="5" pageNumber="38" unit="g" value="4.0">4</quantity>
|
|||
|
|
</figureCitation>
|
|||
|
|
g–4h).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B9336BAF7209323FF501B1714C5F967" blockId="5.[151,1437,258,2010]" pageId="5" pageNumber="38">
|
|||
|
|
Male genitalia (
|
|||
|
|
<figureCitation id="13172A3FF7209323FEE61B1717A1F993" box="[371,440,1590,1614]" captionStart="FIGURE 5" captionStartId="7.[151,250,1954,1977]" captionTargetBox="[184,1380,185,1953]" captionTargetId="figure@7.[158,1441,147,1966]" captionTargetPageId="7" captionText="FIGURE 5. Geocharis barcorabelo n. sp. aedeagus, (a), median lobe and left paramere in lateral view; (b), apex of median lobe in dorsal view. Geocharis antheroi n. sp. aedeagus, (c), median lobe and left paramere in lateral view; (d), apex of median lobe in dorsal view." httpUri="https://zenodo.org/record/207165/files/figure.png" pageId="5" pageNumber="38">Figs 5</figureCitation>
|
|||
|
|
c–5d) with median lobe arcuate (lateral view), apex strongly sharp and bent down (lateral view), arcuate inwards in the right side and broadly rounded in the left side (dorsal view); basal lobe with apophysis prominent, basal edge arcuate; internal sac as in figures 5c–5d; left and right parameres with 2 apical setae, left paramere with dorso-basal edge expanded (
|
|||
|
|
<figureCitation id="13172A3FF7209323FDE81B8314D8F967" box="[637,705,1698,1722]" captionStart="FIGURE 5" captionStartId="7.[151,250,1954,1977]" captionTargetBox="[184,1380,185,1953]" captionTargetId="figure@7.[158,1441,147,1966]" captionTargetPageId="7" captionText="FIGURE 5. Geocharis barcorabelo n. sp. aedeagus, (a), median lobe and left paramere in lateral view; (b), apex of median lobe in dorsal view. Geocharis antheroi n. sp. aedeagus, (c), median lobe and left paramere in lateral view; (d), apex of median lobe in dorsal view." httpUri="https://zenodo.org/record/207165/files/figure.png" pageId="5" pageNumber="38">Fig. 5</figureCitation>
|
|||
|
|
c).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B9336BAF7209323FF501BE71527F807" blockId="5.[151,1437,258,2010]" pageId="5" pageNumber="38">
|
|||
|
|
The female genitalia follows the general pattern described for the other species of the genus (e.g.,
|
|||
|
|
<bibRefCitation id="EFBD4B4BF7209323FA8B1BE7173EF8DF" author="Zaballos" pageId="5" pageNumber="38" refString="Zaballos, J. P. & Jeanne, C. (1987) Etude systematique du genre Geocharis (Col. Trechidae Anillini) et description d`une nouvelle espece. Bulletin de la Societe linneenne de Bordeaux, 15, 81 - 92." type="journal article" year="1987">Zaballos & Jeanne 1987</bibRefCitation>
|
|||
|
|
,
|
|||
|
|
<bibRefCitation id="EFBD4B4BF7209323FEA11BCB17C1F8DF" author="Zaballos" box="[308,472,1770,1794]" pageId="5" pageNumber="38" refString="Zaballos, J. P. (1998) Interesante nueva especie de Geocharis Ehlers, 1883 de Espana (Coleoptera, Caraboidea, Trechidae, Anillini). Graellsia, 54, 19 - 24." type="journal article" year="1998">Zaballos 1998</bibRefCitation>
|
|||
|
|
,
|
|||
|
|
<bibRefCitation id="EFBD4B4BF7209323FE701BCB148BF8DF" author="Zaballos" box="[485,658,1770,1794]" pageId="5" pageNumber="38" refString="Zaballos, J. P. (2005) Los Geocharis Ehlers, 1883 de Marruecos y Cadiz (Espana) (Coleoptera, Carabidae, Trechinae, Anillini). Graellsia, 61, 61 - 81." type="journal article" year="2005">Zaballos, 2005</bibRefCitation>
|
|||
|
|
). Female genitalia (
|
|||
|
|
<figureCitation id="13172A3FF7209323FCE61BCB15AEF8DF" box="[883,951,1770,1794]" captionStart="FIGURE 6" captionStartId="8.[151,250,634,657]" captionTargetBox="[172,1414,193,612]" captionTargetId="figure@8.[172,1415,193,613]" captionTargetPageId="8" captionText="FIGURE 6. SEM photographs of female genitalia in ventral view: (a) Geocharis barcorabelo n. sp.; (b) Geocharis antheroi n. sp." httpUri="https://zenodo.org/record/207166/files/figure.png" pageId="5" pageNumber="38">Fig. 6</figureCitation>
|
|||
|
|
b) shows the gonocoxite IX sickle-shaped, and has a long ensiform seta in the middle region of the external margin, one ensiform seta in the middle dorsal region and a double nematiform seta in the internal margin near the beginning of the apical third. Gonosubcoxite IX without special features; laterotergite IX with a variable number of setae (8–10). Internal genitalia (not shown) with spermathecal duct long, parallel and enlarged very slightly near the bursa copulatrix (length:
|
|||
|
|
<quantity id="4CD49B5FF7209323FA9B1A5B1360F84F" box="[1294,1401,1914,1938]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="5" pageNumber="38" unit="mm" value="0.16">0.16 mm</quantity>
|
|||
|
|
), a spherical spermatheca (width:
|
|||
|
|
<quantity id="4CD49B5FF7209323FE7B1ABF147AF86B" box="[494,611,1950,1974]" metricMagnitude="-5" metricUnit="m" metricValue="1.9" pageId="5" pageNumber="38" unit="mm" value="0.019">0.019 mm</quantity>
|
|||
|
|
), duct of spermathecal gland short and thin, gland fusiform (lenght:
|
|||
|
|
<quantity id="4CD49B5FF7209323FAC81ABF16DEF807" metricMagnitude="-5" metricUnit="m" metricValue="9.6" pageId="5" pageNumber="38" unit="mm" value="0.096">0.096 mm</quantity>
|
|||
|
|
), middle region membranous, apical portion sclerotized.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<caption id="DF536632F7239320FF021A5D1332F80D" httpUri="https://zenodo.org/record/207164/files/figure.png" pageId="6" pageNumber="39" targetBox="[172,1414,183,1910]" targetPageId="6">
|
|||
|
|
<paragraph id="8B9336BAF7239320FF021A5D1332F80D" blockId="6.[151,1436,1916,2000]" pageId="6" pageNumber="39">
|
|||
|
|
<emphasis id="B958EAA8F7239320FF021A5D170BF84F" bold="true" box="[151,274,1916,1939]" pageId="6" pageNumber="39">FIGURE 4.</emphasis>
|
|||
|
|
SEM photographs of structural features of
|
|||
|
|
<taxonomicName id="4C2C4D39F7239320FD501A5C1593F84E" box="[709,906,1917,1939]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="39" phylum="Arthropoda" rank="species" species="antheroi" status="sp. nov.">
|
|||
|
|
<emphasis id="B958EAA8F7239320FD501A5C1593F84E" box="[709,906,1917,1939]" italics="true" pageId="6" pageNumber="39">Geocharis antheroi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A26B57D3F7239320FC1A1A5C15DBF84F" box="[911,962,1917,1938]" pageId="6" pageNumber="39" rank="species">
|
|||
|
|
<emphasis id="B958EAA8F7239320FC1A1A5C15DBF84F" bold="true" box="[911,962,1917,1938]" pageId="6" pageNumber="39">n.sp.</emphasis>
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
, (
|
|||
|
|
<emphasis id="B958EAA8F7239320FC431A5C15FBF84F" bold="true" box="[982,994,1917,1938]" pageId="6" pageNumber="39">a</emphasis>
|
|||
|
|
), head (latero–dorsal view); (
|
|||
|
|
<emphasis id="B958EAA8F7239320FA9B1A5C1305F84F" bold="true" box="[1294,1308,1917,1938]" pageId="6" pageNumber="39">b</emphasis>
|
|||
|
|
), head (ventral view); (
|
|||
|
|
<emphasis id="B958EAA8F7239320FE811ABD1706F86C" bold="true" box="[276,287,1948,1969]" pageId="6" pageNumber="39">c</emphasis>
|
|||
|
|
), pronotum (dorsal view); (
|
|||
|
|
<emphasis id="B958EAA8F7239320FDD11ABD144BF86C" bold="true" box="[580,594,1948,1969]" pageId="6" pageNumber="39">d</emphasis>
|
|||
|
|
), elytra (dorsal view); (
|
|||
|
|
<emphasis id="B958EAA8F7239320FCDB1ABD1540F86C" bold="true" box="[846,857,1948,1969]" pageId="6" pageNumber="39">e</emphasis>
|
|||
|
|
), right elytron (latero-dorsal view); (
|
|||
|
|
<emphasis id="B958EAA8F7239320FB4B1ABA12FFF86F" bold="true" box="[1246,1254,1947,1970]" pageId="6" pageNumber="39">f</emphasis>
|
|||
|
|
), anterior half of elytra (dorsal view); (
|
|||
|
|
<emphasis id="B958EAA8F7239320FEE61A9B1766F812" bold="true" box="[371,383,1978,1999]" pageId="6" pageNumber="39">g</emphasis>
|
|||
|
|
), thorax and abdomen (male, ventral view); (
|
|||
|
|
<emphasis id="B958EAA8F7239320FCD91A9B1543F812" bold="true" box="[844,858,1978,1999]" pageId="6" pageNumber="39">h</emphasis>
|
|||
|
|
), thorax and abdomen (female, ventral view).
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="DF536632F7229321FF021A831747F82B" httpUri="https://zenodo.org/record/207165/files/figure.png" pageId="7" pageNumber="40" targetBox="[184,1380,185,1953]" targetPageId="7">
|
|||
|
|
<paragraph id="8B9336BAF7229321FF021A831747F82B" blockId="7.[151,1436,1954,2038]" pageId="7" pageNumber="40">
|
|||
|
|
<emphasis id="B958EAA8F7229321FF021A83170DF865" bold="true" box="[151,276,1954,1977]" pageId="7" pageNumber="40">FIGURE 5.</emphasis>
|
|||
|
|
<taxonomicName id="4C2C4D39F7229321FE881A821412F864" box="[285,523,1955,1977]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="40" phylum="Arthropoda" rank="species" species="barcorabelo" status="sp. nov.">
|
|||
|
|
<emphasis id="B958EAA8F7229321FE881A821412F864" box="[285,523,1955,1977]" italics="true" pageId="7" pageNumber="40">Geocharis barcorabelo</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B958EAA8F7229321FD861A82145FF865" bold="true" box="[531,582,1955,1976]" pageId="7" pageNumber="40">
|
|||
|
|
<taxonomicNameLabel id="A26B57D3F7229321FD861A82145FF865" box="[531,582,1955,1976]" pageId="7" pageNumber="40" rank="species">n.sp.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
aedeagus, (
|
|||
|
|
<emphasis id="B958EAA8F7229321FD571A8214D6F865" bold="true" box="[706,719,1955,1976]" pageId="7" pageNumber="40">a</emphasis>
|
|||
|
|
), median lobe and left paramere in lateral view; (
|
|||
|
|
<emphasis id="B958EAA8F7229321FB421A8212FCF865" bold="true" box="[1239,1253,1955,1976]" pageId="7" pageNumber="40">b</emphasis>
|
|||
|
|
), apex of median lobe in dorsal view.
|
|||
|
|
<taxonomicName id="4C2C4D39F7229321FEF11AE01430F80A" box="[356,553,1985,2007]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="40" phylum="Arthropoda" rank="species" species="antheroi" status="sp. nov.">
|
|||
|
|
<emphasis id="B958EAA8F7229321FEF11AE01430F80A" box="[356,553,1985,2007]" italics="true" pageId="7" pageNumber="40">Geocharis antheroi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B958EAA8F7229321FDBA1AE3147BF80A" bold="true" box="[559,610,1986,2007]" pageId="7" pageNumber="40">
|
|||
|
|
<taxonomicNameLabel id="A26B57D3F7229321FDBA1AE3147BF80A" box="[559,610,1986,2007]" pageId="7" pageNumber="40" rank="species">n.sp.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
aedeagus, (
|
|||
|
|
<emphasis id="B958EAA8F7229321FD4F1AE314FCF80A" bold="true" box="[730,741,1986,2007]" pageId="7" pageNumber="40">c</emphasis>
|
|||
|
|
), median lobe and left paramere in lateral view; (
|
|||
|
|
<emphasis id="B958EAA8F7229321FB491AE312F3F80A" bold="true" box="[1244,1258,1986,2007]" pageId="7" pageNumber="40">d</emphasis>
|
|||
|
|
), apex of median lobe in dorsal view.
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<caption id="DF536632F72D932EFF021F5B16D3FD72" httpUri="https://zenodo.org/record/207166/files/figure.png" pageId="8" pageNumber="41" targetBox="[172,1414,193,612]" targetPageId="8">
|
|||
|
|
<paragraph id="8B9336BAF72D932EFF021F5B16D3FD72" blockId="8.[151,1436,634,687]" pageId="8" pageNumber="41">
|
|||
|
|
<emphasis id="B958EAA8F72D932EFF021F5B1714FD4D" bold="true" box="[151,269,634,657]" pageId="8" pageNumber="41">FIGURE 6</emphasis>
|
|||
|
|
. SEM photographs of female genitalia in ventral view: (
|
|||
|
|
<emphasis id="B958EAA8F72D932EFCCF1F5A157FFD4D" bold="true" box="[858,870,635,656]" pageId="8" pageNumber="41">a</emphasis>
|
|||
|
|
)
|
|||
|
|
<taxonomicName id="4C2C4D39F72D932EFCE21F5A127CFD4C" box="[887,1125,635,657]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="species" species="barcorabelo" status="sp. nov.">
|
|||
|
|
<emphasis id="B958EAA8F72D932EFCE21F5A127CFD4C" box="[887,1125,635,657]" italics="true" pageId="8" pageNumber="41">Geocharis barcorabelo</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A26B57D3F72D932EFBF81F5A1286FD4C" box="[1133,1183,635,657]" pageId="8" pageNumber="41" rank="species">
|
|||
|
|
<emphasis id="B958EAA8F72D932EFBF81F5A1280FD4D" bold="true" box="[1133,1177,635,656]" pageId="8" pageNumber="41">n.sp</emphasis>
|
|||
|
|
.
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
; (
|
|||
|
|
<emphasis id="B958EAA8F72D932EFB221F5A12DCFD4D" bold="true" box="[1207,1221,635,656]" pageId="8" pageNumber="41">b</emphasis>
|
|||
|
|
)
|
|||
|
|
<taxonomicName id="4C2C4D39F72D932EFB401F5A1385FD4C" box="[1237,1436,635,657]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="species" species="antheroi" status="sp. nov.">
|
|||
|
|
<emphasis id="B958EAA8F72D932EFB401F5A1385FD4C" box="[1237,1436,635,657]" italics="true" pageId="8" pageNumber="41">Geocharis antheroi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A26B57D3F72D932EFF021FBB16D3FD72" box="[151,202,666,687]" pageId="8" pageNumber="41" rank="species">n.sp.</taxonomicNameLabel>
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<subSubSection id="C3366531F72D932EFF501FFC15F5F941" pageId="8" pageNumber="41" type="etymology">
|
|||
|
|
<paragraph id="8B9336BAF72D932EFF501FFC1325FCC4" blockId="8.[151,1437,732,1908]" pageId="8" pageNumber="41">
|
|||
|
|
<emphasis id="B958EAA8F72D932EFF501FFC1755FD28" bold="true" box="[197,332,733,757]" pageId="8" pageNumber="41">Etymology.</emphasis>
|
|||
|
|
This new species is dedicated to the Portuguese naturalist Anthero Frederico Ferreira de Seabra, who greatly contributed to the knowledge of the
|
|||
|
|
<taxonomicName id="4C2C4D39F72D932EFD2D1E21152AFCC5" box="[696,819,768,792]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
|
|||
|
|
of
|
|||
|
|
<collectingCountry id="F33B762AF72D932EFCCD1E2115AEFCC5" box="[856,951,768,792]" name="Portugal" pageId="8" pageNumber="41">Portugal</collectingCountry>
|
|||
|
|
in the first half of the XX century.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B9336BAF72D932EFF521E0514CFFC75" blockId="8.[151,1437,732,1908]" pageId="8" pageNumber="41">
|
|||
|
|
<emphasis id="B958EAA8F72D932EFF521E0517F7FCE0" bold="true" box="[199,494,804,829]" pageId="8" pageNumber="41">Morphological affinities.</emphasis>
|
|||
|
|
The two new species share with most species of
|
|||
|
|
<taxonomicName id="4C2C4D39F72D932EFB8F1E051296FCE1" box="[1050,1167,804,828]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B958EAA8F72D932EFB8F1E051296FCE1" box="[1050,1167,804,828]" italics="true" pageId="8" pageNumber="41">Geocharis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
two setae on the elytral disk, one anterior and one posterior (Table 1). In Table 1 some other morphological characters like the presence or absence of elytral striae, a tooth on the internal margin of the hind femora and the number of setae of the left paramere are compared in all species of
|
|||
|
|
<taxonomicName id="4C2C4D39F72D932EFDCF1EB114D6FC75" box="[602,719,912,936]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B958EAA8F72D932EFDCF1EB114D6FC75" box="[602,719,912,936]" italics="true" pageId="8" pageNumber="41">Geocharis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B9336BAF72D932EFF501E951384FB31" blockId="8.[151,1437,732,1908]" pageId="8" pageNumber="41">
|
|||
|
|
Taking into account the presence on the elytral disk of two setae, the absence of a tooth on the internal margin of the hind femora of males and females, as well as the fact that the adults present traces of elytral striae, the new species are close to
|
|||
|
|
<emphasis id="B958EAA8F72D932EFEE11EDC17C8FBC9" box="[372,465,1020,1044]" italics="true" pageId="8" pageNumber="41">G. b i v a r i</emphasis>
|
|||
|
|
Serrano & Aguiar,
|
|||
|
|
<taxonomicName id="4C2C4D39F72D932EFD251EDC15ADFBC9" authority="Zaballos, G." authorityName="Zaballos, G." box="[688,948,1020,1044]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="species" species="julianae">
|
|||
|
|
<emphasis id="B958EAA8F72D932EFD251EDC1530FBC9" box="[688,809,1020,1044]" italics="true" pageId="8" pageNumber="41">G. julianae</emphasis>
|
|||
|
|
Zaballos,
|
|||
|
|
<emphasis id="B958EAA8F72D932EFC0B1EDC15ADFBC9" box="[926,948,1021,1044]" italics="true" pageId="8" pageNumber="41">G.</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
k o r b i (Ganglbauer),
|
|||
|
|
<taxonomicName id="4C2C4D39F72D932EFB341EDC1381FBC9" authority="Dieck" authorityName="Dieck" box="[1185,1432,1020,1044]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="species" species="massinissa">
|
|||
|
|
<emphasis id="B958EAA8F72D932EFB341EDC1321FBC9" box="[1185,1336,1021,1044]" italics="true" pageId="8" pageNumber="41">G. massinissa</emphasis>
|
|||
|
|
(Dieck)
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C2C4D39F72D932EFF02190017B0FBE5" authority="Zaballos" authorityName="Zaballos" box="[151,425,1056,1080]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="species" species="montecristoi">
|
|||
|
|
<emphasis id="B958EAA8F72D932EFF021900175BFBE5" box="[151,322,1057,1080]" italics="true" pageId="8" pageNumber="41">G. montecristoi</emphasis>
|
|||
|
|
Zaballos
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C2C4D39F72D932EFE751900153BFBE5" authority="Zaballos" authorityName="Zaballos" box="[480,802,1056,1080]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="species" species="testatretafoveata">
|
|||
|
|
<emphasis id="B958EAA8F72D932EFE75190014A3FBE5" box="[480,698,1056,1080]" italics="true" pageId="8" pageNumber="41">G. testatretafoveata</emphasis>
|
|||
|
|
Zaballos
|
|||
|
|
</taxonomicName>
|
|||
|
|
(all the adults of these species exhibit the above characters). Nevertheless, in
|
|||
|
|
<emphasis id="B958EAA8F72D932EFE01196417EBFB81" box="[404,498,1092,1116]" italics="true" pageId="8" pageNumber="41">G. b i v a r i</emphasis>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C2C4D39F72D932EFDB8196414B1FB81" box="[557,680,1092,1116]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="species" species="julianae">
|
|||
|
|
<emphasis id="B958EAA8F72D932EFDB8196414B1FB81" box="[557,680,1092,1116]" italics="true" pageId="8" pageNumber="41">G. julianae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
the elytral striae are stronger than in the new species (see
|
|||
|
|
<bibRefCitation id="EFBD4B4BF72D932EFAA9196516CBFB5D" author="Zaballos" pageId="8" pageNumber="41" refString="Zaballos, J. P. (1989) Un nuevo Geocharis (Coleoptera, Trechidae) de la Sierra de Gredos (Espana). Elytron, 3, 143 - 147." type="journal article" year="1989">Zaballos 1989</bibRefCitation>
|
|||
|
|
, Serrano & Aguiar 2004).
|
|||
|
|
<taxonomicName id="4C2C4D39F72D932EFD97194914D8FB5D" box="[514,705,1128,1152]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="species" species="bivari">
|
|||
|
|
<emphasis id="B958EAA8F72D932EFD97194914D8FB5D" box="[514,705,1128,1152]" italics="true" pageId="8" pageNumber="41">Geocharis bivari</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C2C4D39F72D932EFD4419481555FB5D" box="[721,844,1128,1152]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="species" species="julianae">
|
|||
|
|
<emphasis id="B958EAA8F72D932EFD4419481555FB5D" box="[721,844,1128,1152]" italics="true" pageId="8" pageNumber="41">G. julianae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and the remaining other four species are also easily segregated from the new ones by the shape of the median lobe of aedeagus plus the pattern of internal sac sclerites (
|
|||
|
|
<bibRefCitation id="EFBD4B4BF72D932EFF0B1991175CFB15" author="Zaballos" box="[158,325,1200,1224]" pageId="8" pageNumber="41" refString="Zaballos, J. P. (1989) Un nuevo Geocharis (Coleoptera, Trechidae) de la Sierra de Gredos (Espana). Elytron, 3, 143 - 147." type="journal article" year="1989">Zaballos 1989</bibRefCitation>
|
|||
|
|
,
|
|||
|
|
<bibRefCitation id="EFBD4B4BF72D932EFEC619911797FB15" author="Zaballos" box="[339,398,1200,1224]" pageId="8" pageNumber="41" refString="Zaballos, J. P. (2005) Los Geocharis Ehlers, 1883 de Marruecos y Cadiz (Espana) (Coleoptera, Carabidae, Trechinae, Anillini). Graellsia, 61, 61 - 81." type="journal article" year="2005">2005</bibRefCitation>
|
|||
|
|
,
|
|||
|
|
<bibRefCitation id="EFBD4B4BF72D932EFE0E199114A4FB15" author="Serrano" box="[411,701,1200,1224]" pageId="8" pageNumber="41" refString="Serrano, A. R. M. & Aguiar, C. A. S. (2004 a) Three new species of the genus Geocharis Ehlers, 1883 from Portugal (Coleoptera, Carabidae). Graellsia, 60, 71 - 80." type="journal article" year="2004" yearSuffix="a">Serrano & Aguiar 2004a</bibRefCitation>
|
|||
|
|
). The new species are easily separated from one another by the aedeagus conformation (
|
|||
|
|
<emphasis id="B958EAA8F72D932EFE3E19F517A6FB31" box="[427,447,1236,1260]" italics="true" pageId="8" pageNumber="41">cf</emphasis>
|
|||
|
|
.
|
|||
|
|
<figureCitation id="13172A3FF72D932EFE5819F51408FB31" box="[461,529,1236,1260]" captionStart="FIGURE 5" captionStartId="7.[151,250,1954,1977]" captionTargetBox="[184,1380,185,1953]" captionTargetId="figure@7.[158,1441,147,1966]" captionTargetPageId="7" captionText="FIGURE 5. Geocharis barcorabelo n. sp. aedeagus, (a), median lobe and left paramere in lateral view; (b), apex of median lobe in dorsal view. Geocharis antheroi n. sp. aedeagus, (c), median lobe and left paramere in lateral view; (d), apex of median lobe in dorsal view." httpUri="https://zenodo.org/record/207165/files/figure.png" pageId="8" pageNumber="41">Figs 5</figureCitation>
|
|||
|
|
a–5b and 5c–5d) though they much resemble each other on external morphology.
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B9336BAF72D932EFF5019D915F5F941" blockId="8.[151,1437,732,1908]" pageId="8" pageNumber="41">
|
|||
|
|
The growing knowledge of the genus
|
|||
|
|
<taxonomicName id="4C2C4D39F72D932EFDFC19D914C7FACD" box="[617,734,1272,1296]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B958EAA8F72D932EFDFC19D914C7FACD" box="[617,734,1272,1296]" italics="true" pageId="8" pageNumber="41">Geocharis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in terms of number of species and morphological peculiarities, corroborates former conclusions (e.g.
|
|||
|
|
<bibRefCitation id="EFBD4B4BF72D932EFDAA183D14F9FAE9" author="Zaballos" box="[575,736,1308,1332]" pageId="8" pageNumber="41" refString="Zaballos, J. P. (2005) Los Geocharis Ehlers, 1883 de Marruecos y Cadiz (Espana) (Coleoptera, Carabidae, Trechinae, Anillini). Graellsia, 61, 61 - 81." type="journal article" year="2005">Zaballos 2005</bibRefCitation>
|
|||
|
|
) about the difficulty of identifying its species based only in the external morphology, with the exception of some taxa (see Table 1). Characters that are not mentioned in Table 1 like ovulate elytra and the presence of a vestigial sutural stria (
|
|||
|
|
<taxonomicName id="4C2C4D39F72D932EFCC0184415D2FAA1" box="[853,971,1380,1404]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="species" species="coiffaiti">
|
|||
|
|
<emphasis id="B958EAA8F72D932EFCC0184415D2FAA1" box="[853,971,1380,1404]" italics="true" pageId="8" pageNumber="41">G. coiffaiti</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C2C4D39F72D932EFB9418441293FAA1" box="[1025,1162,1380,1404]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="species" species="femoralis">
|
|||
|
|
<emphasis id="B958EAA8F72D932EFB9418441293FAA1" box="[1025,1162,1380,1404]" italics="true" pageId="8" pageNumber="41">G. femoralis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
), left paramere features, the pronotum shape or even hind tibiae shape are useful for identifying particular
|
|||
|
|
<taxonomicName id="4C2C4D39F72D932EFBB918A912B8FA7D" box="[1068,1185,1416,1440]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B958EAA8F72D932EFBB918A912B8FA7D" box="[1068,1185,1416,1440]" italics="true" pageId="8" pageNumber="41">Geocharis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
species. This happens with adults of
|
|||
|
|
<taxonomicName id="4C2C4D39F72D932EFEAF188C17E3FA19" authority="Zaballos" authorityName="Zaballos" box="[314,506,1452,1476]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="species" species="leoni">
|
|||
|
|
<emphasis id="B958EAA8F72D932EFEAF188C178BFA19" box="[314,402,1452,1476]" italics="true" pageId="8" pageNumber="41">G. leoni</emphasis>
|
|||
|
|
Zaballos
|
|||
|
|
</taxonomicName>
|
|||
|
|
which present the left paramere with lamellar and membranous scales in the apex instead of setae, the adults of
|
|||
|
|
<taxonomicName id="4C2C4D39F72D932EFE4A18F01487FA35" box="[479,670,1488,1512]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="species" species="portalegrensis">
|
|||
|
|
<emphasis id="B958EAA8F72D932EFE4A18F01487FA35" box="[479,670,1488,1512]" italics="true" pageId="8" pageNumber="41">G. portalegrensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
which exhibit a pronotal disk strongly flattened, the adults of
|
|||
|
|
<taxonomicName id="4C2C4D39F72D932EFAC418F01759F9D1" authority="Zaballos" authorityName="Zaballos" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="species" species="iborensis">
|
|||
|
|
<emphasis id="B958EAA8F72D932EFAC418F016C1F9D1" italics="true" pageId="8" pageNumber="41">G. iborensis</emphasis>
|
|||
|
|
Zaballos
|
|||
|
|
</taxonomicName>
|
|||
|
|
which present the hind tibiae arcuate inwards instead of the general straight pattern, and the adults of
|
|||
|
|
<taxonomicName id="4C2C4D39F72D932EFF231B38172BF9ED" box="[182,306,1560,1584]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="species" species="julianae">
|
|||
|
|
<emphasis id="B958EAA8F72D932EFF231B38172BF9ED" box="[182,306,1560,1584]" italics="true" pageId="8" pageNumber="41">G. julianae</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
which present a fold in the internal margin of this structure (
|
|||
|
|
<bibRefCitation id="EFBD4B4BF72D932EFC721B391294F9ED" author="Zaballos" box="[999,1165,1560,1584]" pageId="8" pageNumber="41" refString="Zaballos, J. P. (1989) Un nuevo Geocharis (Coleoptera, Trechidae) de la Sierra de Gredos (Espana). Elytron, 3, 143 - 147." type="journal article" year="1989">Zaballos 1989</bibRefCitation>
|
|||
|
|
, 1990, 1998,
|
|||
|
|
<bibRefCitation id="EFBD4B4BF72D932EFAB21B391733F989" author="Serrano" pageId="8" pageNumber="41" refString="Serrano, A. R. M. & Aguiar, C. A. S. (2000) Two new Geocharis Ehlers, 1883 from Portugal (Coleoptera, Carabidae). Nouvelle Revue d`Entomologie (N. S.), 17, 329 - 335." type="journal article" year="2000">Serrano & Aguiar 2000</bibRefCitation>
|
|||
|
|
). Indeed, the best characters to identify and segregate the
|
|||
|
|
<taxonomicName id="4C2C4D39F72D932EFC571B1D122EF989" box="[962,1079,1596,1620]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B958EAA8F72D932EFC571B1D122EF989" box="[962,1079,1596,1620]" italics="true" pageId="8" pageNumber="41">Geocharis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
species, besides the ones mentioned above, are found in the aedeagus, as are the general conformation of median lobe (lateral view), the apex of the same structure (dorsal view) and the armature pattern of the internal sac.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3366531F72D932AFF521B89143DFE71" lastPageId="12" lastPageNumber="45" pageId="8" pageNumber="41" type="materials_examined">
|
|||
|
|
<paragraph id="8B9336BAF72D932EFF521B891388F8A9" blockId="8.[151,1437,732,1908]" pageId="8" pageNumber="41">
|
|||
|
|
<emphasis id="B958EAA8F72D932EFF521B89121EF91C" bold="true" box="[199,1031,1704,1729]" pageId="8" pageNumber="41">
|
|||
|
|
Faunistic data on other
|
|||
|
|
<taxonomicName id="4C2C4D39F72D932EFE641B89147EF91C" box="[497,615,1704,1729]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B958EAA8F72D932EFE641B89147EF91C" bold="true" box="[497,615,1704,1729]" italics="true" pageId="8" pageNumber="41">Geocharis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
taxa and
|
|||
|
|
<emphasis id="B958EAA8F72D932EFD721B89121EF91C" box="[743,1031,1704,1729]" italics="true" pageId="8" pageNumber="41">
|
|||
|
|
<taxonomicName id="4C2C4D39F72D932EFD721B89121AF91C" box="[743,1027,1704,1729]" class="Insecta" family="Carabidae" genus="Hypotyphlus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="species" species="lusitanicus">Hypotyphlus lusitanicus</taxonomicName>
|
|||
|
|
.
|
|||
|
|
</emphasis>
|
|||
|
|
</emphasis>
|
|||
|
|
<taxonomicName id="4C2C4D39F72D932EFB871B8916CBF939" authority="Schatzmayr, 1937" authorityName="Schatzmayr" authorityYear="1937" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="species" species="olisipensis">
|
|||
|
|
Geocharis olisipensis
|
|||
|
|
<bibRefCitation id="EFBD4B4BF72D932EFA871B8916CBF939" author="Schatzmayr" pageId="8" pageNumber="41" refString="Schatzmayr, A. (1937) Due nuovi Bembidiini anoftalmi della Regione Paleartica. Pubblicasione del Museo Entomologico " Pietro Rossi Duino, 1, 327 - 328." type="journal article" year="1937">Schatzmayr, 1937</bibRefCitation>
|
|||
|
|
</taxonomicName>
|
|||
|
|
. Material examined: Bucelas (U.T.M. coordinates: 29SMD9305),
|
|||
|
|
<date id="FF92107AF72D932EFC511BED122FF939" box="[964,1078,1740,1764]" pageId="8" pageNumber="41" value="2011-03-03">3.III.2011</date>
|
|||
|
|
, 53, 4 Ƥ,
|
|||
|
|
<date id="FF92107AF72D932EFB271BED132BF939" box="[1202,1330,1740,1764]" pageId="8" pageNumber="41" value="2011-03-22">22.III.2011</date>
|
|||
|
|
, 13, 2Ƥ,
|
|||
|
|
<date id="FF92107AF72D932EFF021BD1170FF8D5" box="[151,278,1776,1800]" pageId="8" pageNumber="41" value="2011-03-30">30.III.2011</date>
|
|||
|
|
, 23,
|
|||
|
|
<specimenCount id="9D2AFD33F72D932EFEC71BD1179BF8D5" box="[338,386,1776,1800]" pageId="8" pageNumber="41" type="generic">1 Ƥ.</specimenCount>
|
|||
|
|
Within the genus
|
|||
|
|
<taxonomicName id="4C2C4D39F72D932EFDD91BD114D8F8D5" box="[588,705,1776,1800]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="41" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B958EAA8F72D932EFDD91BD114D8F8D5" box="[588,705,1776,1800]" italics="true" pageId="8" pageNumber="41">Geocharis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
this species was the first one described for
|
|||
|
|
<collectingCountry id="F33B762AF72D932EFB311BD1131AF8D5" box="[1188,1283,1776,1800]" name="Portugal" pageId="8" pageNumber="41">Portugal</collectingCountry>
|
|||
|
|
based on two specimens collected near Lisbon (
|
|||
|
|
<bibRefCitation id="EFBD4B4BF72D932EFD891A3514F1F8F1" author="Schatzmayr" box="[540,744,1812,1836]" pageId="8" pageNumber="41" refString="Schatzmayr, A. (1937) Due nuovi Bembidiini anoftalmi della Regione Paleartica. Pubblicasione del Museo Entomologico " Pietro Rossi Duino, 1, 327 - 328." type="journal article" year="1937">Schatzmayr 1937</bibRefCitation>
|
|||
|
|
).
|
|||
|
|
<bibRefCitation id="EFBD4B4BF72D932EFD681A351235F8F1" author="Serrano" box="[765,1068,1812,1836]" pageId="8" pageNumber="41" refString="Serrano, A. R. M. & Aguiar, C. A. S. (2004 a) Three new species of the genus Geocharis Ehlers, 1883 from Portugal (Coleoptera, Carabidae). Graellsia, 60, 71 - 80." type="journal article" year="2004" yearSuffix="a">Serrano & Aguiar (2004a)</bibRefCitation>
|
|||
|
|
after some efforts to locate this species found it in the outskirts of Lisbon (Valejas and Fanhões) and more recently in Serra de Montejunto (almost
|
|||
|
|
<quantity id="4CD49B5FF72D932EFF021A7D16C7F8A9" box="[151,222,1884,1908]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="41" unit="km" value="50.0">50 km</quantity>
|
|||
|
|
north of Lisbon) (
|
|||
|
|
<bibRefCitation id="EFBD4B4BF72D932EFE301A7D14B4F8A9" author="Serrano" box="[421,685,1884,1908]" pageId="8" pageNumber="41" refString="Serrano, A. R. M. & Aguiar, C. A. S. (2008) A new species of the genus Geocharis Ehlers, 1883 and some faunistic data on endogean carabids from Portugal (Coleoptera: Carabidae). Revue Suisse de Zoologie, 115, 779 - 788." type="journal article" year="2008">Serrano & Aguiar 2008</bibRefCitation>
|
|||
|
|
). The new locality near Bucelas is close to the former ones (
|
|||
|
|
<figureCitation id="13172A3FF72D932EFAAA1A7D139AF8A9" box="[1343,1411,1884,1908]" captionStart="FIGURE 7" captionStartId="11.[151,250,1975,1998]" captionTargetBox="[359,1229,221,1922]" captionTargetId="figure@11.[337,1249,193,1954]" captionTargetPageId="11" captionText="FIGURE 7. Geographical distribution in Portugal (U. T. M. 10 x 10 km coordinates) of G. barcorabelo n. sp. (B), G. a n t h e ro i n. sp. (A), G. olisipensis (O), G. quartaui (Q), and H. lusitanicus (L) (red colour capitals)." httpUri="https://zenodo.org/record/207167/files/figure.png" pageId="8" pageNumber="41">Fig. 7</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<caption id="DF536632F72E932DFF021A961238F82C" httpUri="https://zenodo.org/record/207167/files/figure.png" pageId="11" pageNumber="44" targetBox="[359,1229,221,1922]" targetPageId="11">
|
|||
|
|
<paragraph id="8B9336BAF72E932DFF021A961238F82C" blockId="11.[151,1433,1975,2033]" pageId="11" pageNumber="44">
|
|||
|
|
<emphasis id="B958EAA8F72E932DFF021A96170CF810" bold="true" box="[151,277,1975,1998]" pageId="11" pageNumber="44">FIGURE 7.</emphasis>
|
|||
|
|
Geographical distribution in Portugal (U.T.M. 10x10 km coordinates) of
|
|||
|
|
<taxonomicName id="4C2C4D39F72E932DFB861A9812B4F813" box="[1043,1197,1976,1998]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="44" phylum="Arthropoda" rank="species" species="barcorabelo" status="sp. nov.">
|
|||
|
|
<emphasis id="B958EAA8F72E932DFB861A9812B4F813" box="[1043,1197,1976,1998]" italics="true" pageId="11" pageNumber="44">G. barcorabelo</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A26B57D3F72E932DFB201A9912E9F813" box="[1205,1264,1976,1998]" pageId="11" pageNumber="44" rank="species">
|
|||
|
|
<emphasis id="B958EAA8F72E932DFB201A9912F3F810" bold="true" box="[1205,1258,1976,1997]" pageId="11" pageNumber="44">n. sp</emphasis>
|
|||
|
|
.
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
(
|
|||
|
|
<emphasis id="B958EAA8F72E932DFA941A99130BF810" bold="true" box="[1281,1298,1976,1997]" pageId="11" pageNumber="44">B</emphasis>
|
|||
|
|
),
|
|||
|
|
<emphasis id="B958EAA8F72E932DFABC1A981380F813" box="[1321,1433,1976,1998]" italics="true" pageId="11" pageNumber="44">G. a n t h e ro i</emphasis>
|
|||
|
|
<emphasis id="B958EAA8F72E932DFF021AFA16D3F82D" bold="true" box="[151,202,2011,2032]" pageId="11" pageNumber="44">n.sp.</emphasis>
|
|||
|
|
(
|
|||
|
|
<emphasis id="B958EAA8F72E932DFF4D1AFA16F3F82D" bold="true" box="[216,234,2011,2032]" pageId="11" pageNumber="44">A</emphasis>
|
|||
|
|
),
|
|||
|
|
<taxonomicName id="4C2C4D39F72E932DFF6A1AFD179FF82C" box="[255,390,2011,2033]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="44" phylum="Arthropoda" rank="species" species="olisipensis">
|
|||
|
|
<emphasis id="B958EAA8F72E932DFF6A1AFD179FF82C" box="[255,390,2011,2033]" italics="true" pageId="11" pageNumber="44">G. olisipensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<emphasis id="B958EAA8F72E932DFE001AFB17B0F82C" bold="true" box="[405,425,2010,2033]" pageId="11" pageNumber="44">O</emphasis>
|
|||
|
|
),
|
|||
|
|
<taxonomicName id="4C2C4D39F72E932DFE281AFD1429F82C" box="[445,560,2012,2033]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="44" phylum="Arthropoda" rank="species" species="quartaui">
|
|||
|
|
<emphasis id="B958EAA8F72E932DFE281AFD1429F82C" box="[445,560,2012,2033]" italics="true" pageId="11" pageNumber="44">G. quartaui</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<emphasis id="B958EAA8F72E932DFDAA1AFB144AF82C" bold="true" box="[575,595,2010,2033]" pageId="11" pageNumber="44">Q</emphasis>
|
|||
|
|
), and
|
|||
|
|
<taxonomicName id="4C2C4D39F72E932DFD061AFD1539F82C" box="[659,800,2011,2033]" class="Insecta" family="Carabidae" genus="Hypotyphlus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="44" phylum="Arthropoda" rank="species" species="lusitanicus">
|
|||
|
|
<emphasis id="B958EAA8F72E932DFD061AFD1539F82C" box="[659,800,2011,2033]" italics="true" pageId="11" pageNumber="44">H. lusitanicus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(
|
|||
|
|
<emphasis id="B958EAA8F72E932DFCBB1AFA1526F82D" bold="true" box="[814,831,2011,2032]" pageId="11" pageNumber="44">L</emphasis>
|
|||
|
|
) (red colour capitals).
|
|||
|
|
</paragraph>
|
|||
|
|
</caption>
|
|||
|
|
<paragraph id="8B9336BAF729932AFF501DB9143DFE71" blockId="12.[151,1437,151,1760]" pageId="12" pageNumber="45">
|
|||
|
|
<taxonomicName id="4C2C4D39F729932AFF501DB914A3FF72" authority="Serrano & Aguiar, 2004" authorityName="Serrano & Aguiar" authorityYear="2004" box="[197,698,151,176]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="species" species="quartaui">
|
|||
|
|
<emphasis id="B958EAA8F729932AFF501DB917B8FF6D" box="[197,417,152,176]" italics="true" pageId="12" pageNumber="45">Geocharis quartaui</emphasis>
|
|||
|
|
Serrano & Aguiar, 2004
|
|||
|
|
</taxonomicName>
|
|||
|
|
. Material examined: Serra de Sicó (Pombal) (U.T.M. coordinates: 29SNE3319),
|
|||
|
|
<date id="FF92107AF729932AFEAF1D9D17A2FF09" box="[314,443,188,212]" pageId="12" pageNumber="45" value="2008-03-31">31.III.2008</date>
|
|||
|
|
, 1Ƥ,
|
|||
|
|
<date id="FF92107AF729932AFE681D9D1462FF09" box="[509,635,188,212]" pageId="12" pageNumber="45" value="2008-04-15">15.IV.2008</date>
|
|||
|
|
, 13, 3Ƥ; Serra do Sicó (Pombal) (U.T.M. coordinates: 29SNE3419),
|
|||
|
|
<date id="FF92107AF729932AFF021DFE170CFF2A" box="[151,277,223,247]" pageId="12" pageNumber="45" value="2008-04-15">15.IV.2008</date>
|
|||
|
|
, 23 3,
|
|||
|
|
<specimenCount id="9D2AFD33F729932AFEFA1DFE17A8FF2A" box="[367,433,223,247]" pageId="12" pageNumber="45" type="generic">10 Ƥ.</specimenCount>
|
|||
|
|
This species was described on the basis of several specimens collected near Alcobaça (Carvalhal), a locality close to Serra de Aire e Candeeiros and recorded later in Serra de Montejunto (
|
|||
|
|
<bibRefCitation id="EFBD4B4BF729932AFAB11C251733FEE2" author="Serrano" pageId="12" pageNumber="45" refString="Serrano, A. R. M. & Aguiar, C. A. S. (2008) A new species of the genus Geocharis Ehlers, 1883 and some faunistic data on endogean carabids from Portugal (Coleoptera: Carabidae). Revue Suisse de Zoologie, 115, 779 - 788." type="journal article" year="2008">Serrano & Aguiar 2008</bibRefCitation>
|
|||
|
|
). This new localization in Serra do Sicó (almost
|
|||
|
|
<quantity id="4CD49B5FF729932AFCCC1C0615B8FEE2" box="[857,929,295,319]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="12" pageNumber="45" unit="km" value="40.0">40 km</quantity>
|
|||
|
|
north of Serra de Montejunto), indicates for this species a wider distribution than the previously thought (
|
|||
|
|
<figureCitation id="13172A3FF729932AFCC61C6D1580FEB9" box="[851,921,332,356]" captionStart="FIGURE 7" captionStartId="11.[151,250,1975,1998]" captionTargetBox="[359,1229,221,1922]" captionTargetId="figure@11.[337,1249,193,1954]" captionTargetPageId="11" captionText="FIGURE 7. Geographical distribution in Portugal (U. T. M. 10 x 10 km coordinates) of G. barcorabelo n. sp. (B), G. a n t h e ro i n. sp. (A), G. olisipensis (O), G. quartaui (Q), and H. lusitanicus (L) (red colour capitals)." httpUri="https://zenodo.org/record/207167/files/figure.png" pageId="12" pageNumber="45">Fig. 7</figureCitation>
|
|||
|
|
). Interesting also is the fact that the Serra de Sicó male specimens exhibit a stronger median tooth on the internal margin of the hind femora than that observed in specimens of the other localities.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
<subSubSection id="C3366531F729932AFF501C9917ACF93D" pageId="12" pageNumber="45" type="reference_group">
|
|||
|
|
<paragraph id="8B9336BAF729932AFF501C991303FD32" blockId="12.[151,1437,151,1760]" pageId="12" pageNumber="45">
|
|||
|
|
<taxonomicName id="4C2C4D39F729932AFF501C9914EDFE12" ID-CoL="3NZYL" authority="Serrano & Aguiar, 2004" authorityName="Serrano & Aguiar" authorityYear="2004" box="[197,756,439,464]" class="Insecta" family="Carabidae" genus="Hypotyphlus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="species" species="lusitanicus">
|
|||
|
|
<emphasis id="B958EAA8F729932AFF501C9917CDFE0D" box="[197,468,440,464]" italics="true" pageId="12" pageNumber="45">Hypotyphlus lusitanicus</emphasis>
|
|||
|
|
Serrano & Aguiar, 2004
|
|||
|
|
</taxonomicName>
|
|||
|
|
. Material examined: Aldeia do Catarredor (Serra da Lousã) (U.T.M. coordinates: 29TNE6636),
|
|||
|
|
<date id="FF92107AF729932AFDD31CFD14AEFE29" box="[582,695,476,500]" pageId="12" pageNumber="45" value="2008-04-02">2.IV.2008</date>
|
|||
|
|
, 2 3; Fundeira (Pampilhosa da Serra) (U.T.M. coordinates: 29TNE8731),
|
|||
|
|
<date id="FF92107AF729932AFEAF1CDE17B3FDCA" box="[314,426,511,535]" pageId="12" pageNumber="45" value="2008-04-03">3.IV.2008</date>
|
|||
|
|
, 4 3, 3 Ƥ; Silvares (Fundão) (U.T.M. coordinates: 29TPE1141),
|
|||
|
|
<date id="FF92107AF729932AFB191CDE12E5FDCA" box="[1164,1276,511,535]" pageId="12" pageNumber="45" value="2008-04-04">4.IV.2008</date>
|
|||
|
|
, 1 Ƥ; Folgosa (Peso da Régua) (U.T.M. coordinates: 29TPF1357),
|
|||
|
|
<date id="FF92107AF729932AFD661F05156BFDE1" box="[755,882,548,572]" pageId="12" pageNumber="45" value="2009-04-29">29.IV.2009</date>
|
|||
|
|
,
|
|||
|
|
<specimenCount id="9D2AFD33F729932AFC151F0515AAFDE1" box="[896,947,548,572]" pageId="12" pageNumber="45" type="generic">1 Ƥ.</specimenCount>
|
|||
|
|
One male and
|
|||
|
|
<specimenCount id="9D2AFD33F729932AFBFD1F0512DAFDE1" box="[1128,1219,548,572]" pageId="12" pageNumber="45" type="female">1female</specimenCount>
|
|||
|
|
from Fundeira are deposited in the collection of Vicente Ortuño; all the remaining specimens are deposited in the collection of the first author, Departamento de Biologia Animal (Faculdade de Ciências da Universidade de Lisboa). This remarkable species was found in the centre of
|
|||
|
|
<collectingCountry id="F33B762AF729932AFDDB1FAE14B4FD7A" box="[590,685,655,679]" name="Portugal" pageId="12" pageNumber="45">Portugal</collectingCountry>
|
|||
|
|
(Aldeia do Mato near Tomar) (
|
|||
|
|
<bibRefCitation id="EFBD4B4BF729932AFB9F1FAE1332FD7A" author="Serrano" box="[1034,1323,655,679]" pageId="12" pageNumber="45" refString="Serrano, A. R. M. & Aguiar, C. A. S. (2004 b) A remarkable new endogean species of the genus Hypotyphlus Jeannel (Coleoptera: Carabidae) from Portugal. The Coleopterists Bulletin, 58, 111 - 117." type="journal article" year="2004" yearSuffix="b">Serrano & Aguiar, 2004b</bibRefCitation>
|
|||
|
|
). Lately it was found in a new locality (Barragem do Cabril, Pedrógão Grande) almost
|
|||
|
|
<quantity id="4CD49B5FF729932AFC7F1F951228FD11" box="[1002,1073,692,716]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="45" unit="km" value="60.0">60 km</quantity>
|
|||
|
|
farther north (
|
|||
|
|
<bibRefCitation id="EFBD4B4BF729932AFB471F9516CDFD32" author="Serrano" pageId="12" pageNumber="45" refString="Serrano, A. R. M. & Aguiar, C. A. S. (2008) A new species of the genus Geocharis Ehlers, 1883 and some faunistic data on endogean carabids from Portugal (Coleoptera: Carabidae). Revue Suisse de Zoologie, 115, 779 - 788." type="journal article" year="2008">Serrano & Aguiar 2008</bibRefCitation>
|
|||
|
|
). The new localities notably increase east- and northwards the distribution of this species (
|
|||
|
|
<figureCitation id="13172A3FF729932AFB5D1FF61315FD32" box="[1224,1292,727,751]" captionStart="FIGURE 7" captionStartId="11.[151,250,1975,1998]" captionTargetBox="[359,1229,221,1922]" captionTargetId="figure@11.[337,1249,193,1954]" captionTargetPageId="11" captionText="FIGURE 7. Geographical distribution in Portugal (U. T. M. 10 x 10 km coordinates) of G. barcorabelo n. sp. (B), G. a n t h e ro i n. sp. (A), G. olisipensis (O), G. quartaui (Q), and H. lusitanicus (L) (red colour capitals)." httpUri="https://zenodo.org/record/207167/files/figure.png" pageId="12" pageNumber="45">Fig. 7</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B9336BAF729932AFF521FDA134CFB8A" blockId="12.[151,1437,151,1760]" pageId="12" pageNumber="45">
|
|||
|
|
<emphasis id="B958EAA8F729932AFF521FDA14CCFCC9" bold="true" box="[199,725,763,788]" pageId="12" pageNumber="45">Ecological and geographical considerations.</emphasis>
|
|||
|
|
The two new species of
|
|||
|
|
<taxonomicName id="4C2C4D39F729932AFC671FDD127EFCC9" ID-CoL="8KX23" box="[1010,1127,764,788]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B958EAA8F729932AFC671FDD127EFCC9" box="[1010,1127,764,788]" italics="true" pageId="12" pageNumber="45">Geocharis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
inhabit the endogean environment in the same way than other congeneric taxa. This means that they live in the soil, usually at different depths of the B-horizon. They are found under sunken stones laying at different depths, from superficial (epigean) to well-buried (edaphic or endogean) environments. Individuals of endogean carabids are more easily found close to the superficial horizon layers after heavy rains because then the soil reaches higher percentage of humidity (saturation or close to saturation), pushing the beetles upwards. References to this behaviour are not abundant (e.g.
|
|||
|
|
<bibRefCitation id="EFBD4B4BF729932AFF021EF51722FC31" author="Zaballos" box="[151,315,980,1004]" pageId="12" pageNumber="45" refString="Zaballos, J. P. (2005) Los Geocharis Ehlers, 1883 de Marruecos y Cadiz (Espana) (Coleoptera, Carabidae, Trechinae, Anillini). Graellsia, 61, 61 - 81." type="journal article" year="2005">Zaballos 2005</bibRefCitation>
|
|||
|
|
;
|
|||
|
|
<bibRefCitation id="EFBD4B4BF729932AFED21EF514DAFC31" author="Zaballos" box="[327,707,980,1004]" pageId="12" pageNumber="45" refString="Zaballos, J. P. & Perez-Gonzalez, S. (2011) A new species of Typhlocharis Dieck, 1869 (Coleoptera: Carabidae: Anillini) from South Spain, with notes on the phylogenetic value of sexually related characters and the presence of stridulatory organ (pars stridens) in the genus. Zootaxa, 2786, 42 - 50." type="journal article" year="2011">Zaballos & Pérez-González 2011</bibRefCitation>
|
|||
|
|
), though we sampled several specimens of
|
|||
|
|
<taxonomicName id="4C2C4D39F729932AFB251EF4171BFBD2" ID-CoL="6KBW4" authority="Serrano & Aguiar" authorityName="Serrano & Aguiar" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="species" species="sacarraoi">
|
|||
|
|
<emphasis id="B958EAA8F729932AFB251EF41324FC31" box="[1200,1341,981,1004]" italics="true" pageId="12" pageNumber="45">G. sacarraoi</emphasis>
|
|||
|
|
Serrano & Aguiar
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C2C4D39F729932AFEAF1ED91527FBD2" ID-CoL="59VKD" authority="Serrano & Aguiar, 2005" authorityName="Serrano & Aguiar" authorityYear="2005" box="[314,830,1015,1040]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="species" species="passosi">
|
|||
|
|
<emphasis id="B958EAA8F729932AFEAF1ED9143EFBCD" box="[314,551,1016,1040]" italics="true" pageId="12" pageNumber="45">Typhlocharis passosi</emphasis>
|
|||
|
|
Serrano & Aguiar, 2005
|
|||
|
|
</taxonomicName>
|
|||
|
|
after rain in these conditions. However, the soil horizons do not always present the same depth and many times the bed rock is closer to the epigean environment. In this case endogean individuals may occur for instance between plates of sunken schistose rocks (e.g.
|
|||
|
|
<figureCitation id="13172A3FF729932AFA96191E135EFB8A" box="[1283,1351,1087,1111]" captionStart="FIGURE 2" captionStartId="2.[151,250,1164,1187]" captionTargetBox="[151,1435,186,1142]" captionTargetId="figure@2.[151,1436,179,1143]" captionTargetPageId="2" captionText="FIGURE 2. The habitat of G. a n t h e ro i n. sp. in Carção (Vimioso region)." httpUri="https://zenodo.org/record/207162/files/figure.png" pageId="12" pageNumber="45">Fig. 2</figureCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B9336BAF729932AFF50194515CAFB19" blockId="12.[151,1437,151,1760]" pageId="12" pageNumber="45">
|
|||
|
|
Though only one species of
|
|||
|
|
<taxonomicName id="4C2C4D39F729932AFD8619451491FBA1" ID-CoL="8KX23" box="[531,648,1124,1148]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B958EAA8F729932AFD8619451491FBA1" box="[531,648,1124,1148]" italics="true" pageId="12" pageNumber="45">Geocharis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
occurs at a given locality, there are some recorded exceptions (e.g.
|
|||
|
|
<bibRefCitation id="EFBD4B4BF729932AFF0219A91720FB7D" author="Zaballos" box="[151,313,1160,1184]" pageId="12" pageNumber="45" refString="Zaballos, J. P. (2005) Los Geocharis Ehlers, 1883 de Marruecos y Cadiz (Espana) (Coleoptera, Carabidae, Trechinae, Anillini). Graellsia, 61, 61 - 81." type="journal article" year="2005">Zaballos 2005</bibRefCitation>
|
|||
|
|
; Serrano & Aguiar 2001, 2006, 2008). Sometimes
|
|||
|
|
<taxonomicName id="4C2C4D39F729932AFCE319A915F2FB7D" ID-CoL="8KX23" box="[886,1003,1160,1184]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B958EAA8F729932AFCE319A915F2FB7D" box="[886,1003,1160,1184]" italics="true" pageId="12" pageNumber="45">Geocharis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
species can also be syntopic with species of the genus
|
|||
|
|
<taxonomicName id="4C2C4D39F729932AFECC198D17F3FB19" ID-CoL="8KZDV" box="[345,490,1196,1220]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B958EAA8F729932AFECC198D17F3FB19" box="[345,490,1196,1220]" italics="true" pageId="12" pageNumber="45">Typhlocharis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(e.g.
|
|||
|
|
<bibRefCitation id="EFBD4B4BF729932AFDBE198D14A9FB19" author="Dieck" box="[555,688,1196,1220]" pageId="12" pageNumber="45" refString="Dieck, G. (1869) Beitrage zur subterranen Kaferfauna Sudeuropas und Maroccos. Berliner Entomologische Zeitschrift, 13, 337 - 360." type="journal article" year="1869">Dieck 1869</bibRefCitation>
|
|||
|
|
,
|
|||
|
|
<bibRefCitation id="EFBD4B4BF729932AFD2E198D15DEFB19" author="Serrano" box="[699,967,1196,1220]" pageId="12" pageNumber="45" refString="Serrano, A. R. M. & Aguiar, C. A. S. (2008) A new species of the genus Geocharis Ehlers, 1883 and some faunistic data on endogean carabids from Portugal (Coleoptera: Carabidae). Revue Suisse de Zoologie, 115, 779 - 788." type="journal article" year="2008">Serrano & Aguiar 2008</bibRefCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B9336BAF729932AFF5019F0140EF9F1" blockId="12.[151,1437,151,1760]" pageId="12" pageNumber="45">
|
|||
|
|
To date, all the
|
|||
|
|
<taxonomicName id="4C2C4D39F729932AFEE519F117FCFB35" ID-CoL="8KX23" box="[368,485,1232,1256]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B958EAA8F729932AFEE519F117FCFB35" box="[368,485,1232,1256]" italics="true" pageId="12" pageNumber="45">Geocharis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
species located north of Tejo River in
|
|||
|
|
<collectingCountry id="F33B762AF729932AFC0619F115EBFB35" box="[915,1010,1232,1256]" name="Portugal" pageId="12" pageNumber="45">Portugal</collectingCountry>
|
|||
|
|
were recorded from localities ranging between the Lisbon region and Serra de Aire e Candeeiros. Interestingly, despite several field surveys to the north of Tejo River, only those conducted in the western region north of Lisbon led to the discovery of
|
|||
|
|
<taxonomicName id="4C2C4D39F729932AFB451839135CFAED" ID-CoL="8KX23" box="[1232,1349,1304,1328]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B958EAA8F729932AFB451839135CFAED" box="[1232,1349,1304,1328]" italics="true" pageId="12" pageNumber="45">Geocharis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
species until now.
|
|||
|
|
<taxonomicName id="4C2C4D39F729932AFE9B181D17F3FA89" ID-CoL="3FP27" box="[270,490,1340,1364]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="species" species="quartaui">
|
|||
|
|
<emphasis id="B958EAA8F729932AFE9B181D17F3FA89" box="[270,490,1340,1364]" italics="true" pageId="12" pageNumber="45">Geocharis quartaui</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
(also collected in this study) was found in Serra de Sicó, a mountain also located in the western region. The finding of the two new species here reported from surveys near the Douro River and Vimioso has increased the distribution of this genus
|
|||
|
|
<quantity id="4CD49B5FF729932AFD3118A514E0FA41" box="[676,761,1412,1436]" metricMagnitude="5" metricUnit="m" metricValue="2.4" pageId="12" pageNumber="45" unit="km" value="240.0">240 km</quantity>
|
|||
|
|
farther north of Serra de Sicó. Yet, more remarkable was the discovery of
|
|||
|
|
<taxonomicName id="4C2C4D39F729932AFEBE18891420FA1D" ID-CoL="3NZYL" box="[299,569,1448,1472]" class="Insecta" family="Carabidae" genus="Hypotyphlus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="species" species="lusitanicus">
|
|||
|
|
<emphasis id="B958EAA8F729932AFEBE18891420FA1D" box="[299,569,1448,1472]" italics="true" pageId="12" pageNumber="45">Hypotyphlus lusitanicus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
in Folgosa in syntopy with
|
|||
|
|
<taxonomicName id="4C2C4D39F729932AFCED18891238FA1D" ID-CoL="3FNZ5" box="[888,1057,1448,1472]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="species" species="barcorabelo" status="sp. nov.">
|
|||
|
|
<emphasis id="B958EAA8F729932AFCED18891238FA1D" box="[888,1057,1448,1472]" italics="true" pageId="12" pageNumber="45">G. barcorabelo</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A26B57D3F729932AFBBC18891278FA1D" box="[1065,1121,1448,1472]" pageId="12" pageNumber="45" rank="species">
|
|||
|
|
<emphasis id="B958EAA8F729932AFBBC18891243FA1D" bold="true" box="[1065,1114,1448,1472]" pageId="12" pageNumber="45">n.sp</emphasis>
|
|||
|
|
.
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
The presence of the former species north and south of Serra da Estrela (
|
|||
|
|
<figureCitation id="13172A3FF729932AFD0D18ED14C6FA39" box="[664,735,1484,1508]" captionStart="FIGURE 7" captionStartId="11.[151,250,1975,1998]" captionTargetBox="[359,1229,221,1922]" captionTargetId="figure@11.[337,1249,193,1954]" captionTargetPageId="11" captionText="FIGURE 7. Geographical distribution in Portugal (U. T. M. 10 x 10 km coordinates) of G. barcorabelo n. sp. (B), G. a n t h e ro i n. sp. (A), G. olisipensis (O), G. quartaui (Q), and H. lusitanicus (L) (red colour capitals)." httpUri="https://zenodo.org/record/207167/files/figure.png" pageId="12" pageNumber="45">Fig. 7</figureCitation>
|
|||
|
|
) corroborates the hypothesis that
|
|||
|
|
<taxonomicName id="4C2C4D39F729932AFBF218EC131CFA39" ID-CoL="3NZYL" box="[1127,1285,1484,1508]" class="Insecta" family="Carabidae" genus="Hypotyphlus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="species" species="lusitanicus">
|
|||
|
|
<emphasis id="B958EAA8F729932AFBF218EC131CFA39" box="[1127,1285,1484,1508]" italics="true" pageId="12" pageNumber="45">H. lusitanicus</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
is a zoogeographical relict of a lineage that once must have extended westward from the Lionigurian massif to the Lusitanian massif (
|
|||
|
|
<bibRefCitation id="EFBD4B4BF729932AFF7A1B351413F9F1" author="Serrano" box="[239,522,1556,1580]" pageId="12" pageNumber="45" refString="Serrano, A. R. M. & Aguiar, C. A. S. (2004 b) A remarkable new endogean species of the genus Hypotyphlus Jeannel (Coleoptera: Carabidae) from Portugal. The Coleopterists Bulletin, 58, 111 - 117." type="journal article" year="2004" yearSuffix="b">Serrano & Aguiar 2004b</bibRefCitation>
|
|||
|
|
).
|
|||
|
|
</paragraph>
|
|||
|
|
<paragraph id="8B9336BAF729932AFF501B1917ACF93D" blockId="12.[151,1437,151,1760]" pageId="12" pageNumber="45">
|
|||
|
|
Finally we would like point out that
|
|||
|
|
<taxonomicName id="4C2C4D39F729932AFDC81B19151CF98D" ID-CoL="3FNZ5" box="[605,773,1592,1616]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="species" species="barcorabelo" status="sp. nov.">
|
|||
|
|
<emphasis id="B958EAA8F729932AFDC81B19151CF98D" box="[605,773,1592,1616]" italics="true" pageId="12" pageNumber="45">G. barcorabelo</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<taxonomicNameLabel id="A26B57D3F729932AFC991B191552F98D" box="[780,843,1592,1616]" pageId="12" pageNumber="45" rank="species">
|
|||
|
|
<emphasis id="B958EAA8F729932AFC991B19155DF98D" bold="true" box="[780,836,1592,1616]" pageId="12" pageNumber="45">n. sp</emphasis>
|
|||
|
|
.
|
|||
|
|
</taxonomicNameLabel>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C2C4D39F729932AFC171B1915E7F98D" ID-CoL="3FNZ3" box="[898,1022,1592,1616]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="species" species="antheroi" status="sp. nov.">
|
|||
|
|
<emphasis id="B958EAA8F729932AFC171B1915E7F98D" box="[898,1022,1592,1616]" italics="true" pageId="12" pageNumber="45">G. antheroi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
<emphasis id="B958EAA8F729932AFB901B191224F98D" bold="true" box="[1029,1085,1592,1616]" pageId="12" pageNumber="45">
|
|||
|
|
<taxonomicNameLabel id="A26B57D3F729932AFB901B191224F98D" box="[1029,1085,1592,1616]" pageId="12" pageNumber="45" rank="species">n.sp.</taxonomicNameLabel>
|
|||
|
|
</emphasis>
|
|||
|
|
were not found within the limits of any Natural Park or protected area and as such are not currently under any conservation protection. From the 20
|
|||
|
|
<taxonomicName id="4C2C4D39F729932AFF2F1BA11736F945" ID-CoL="8KX23" box="[186,303,1664,1688]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="genus">
|
|||
|
|
<emphasis id="B958EAA8F729932AFF2F1BA11736F945" box="[186,303,1664,1688]" italics="true" pageId="12" pageNumber="45">Geocharis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
species known in
|
|||
|
|
<collectingCountry id="F33B762AF729932AFE6A1BA11447F945" box="[511,606,1664,1688]" name="Portugal" pageId="12" pageNumber="45">Portugal</collectingCountry>
|
|||
|
|
(including the new ones) (Table 1), only 7 (
|
|||
|
|
<emphasis id="B958EAA8F729932AFBDB1BA112B5F945" box="[1102,1196,1664,1688]" italics="true" pageId="12" pageNumber="45">G. b i v a r i</emphasis>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C2C4D39F729932AFB2B1BA1132CF945" ID-CoL="3FNZ8" box="[1214,1333,1664,1688]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="species" species="boieiroi">
|
|||
|
|
<emphasis id="B958EAA8F729932AFB2B1BA1132CF945" box="[1214,1333,1664,1688]" italics="true" pageId="12" pageNumber="45">G. boieiroi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C2C4D39F729932AFAD11BA116EEF961" ID-CoL="3FNZW" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="species" species="monfortensis">
|
|||
|
|
<emphasis id="B958EAA8F729932AFAD11BA116EEF961" italics="true" pageId="12" pageNumber="45">G. monfortensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<emphasis id="B958EAA8F729932AFE921B8417B8F961" box="[263,417,1700,1724]" italics="true" pageId="12" pageNumber="45">G. m o s c a t e l u s</emphasis>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C2C4D39F729932AFE201B84146FF961" ID-CoL="3FP26" box="[437,630,1700,1724]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="species" species="portalegrensis">
|
|||
|
|
<emphasis id="B958EAA8F729932AFE201B84146FF961" box="[437,630,1700,1724]" italics="true" pageId="12" pageNumber="45">G. portalegrensis</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
,
|
|||
|
|
<taxonomicName id="4C2C4D39F729932AFD131B841511F961" ID-CoL="3FP27" box="[646,776,1701,1724]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="species" species="quartaui">
|
|||
|
|
<emphasis id="B958EAA8F729932AFD131B841511F961" box="[646,776,1701,1724]" italics="true" pageId="12" pageNumber="45">G. quartaui</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
and
|
|||
|
|
<taxonomicName id="4C2C4D39F729932AFCD71B8415CEF961" ID-CoL="6KBVS" box="[834,983,1700,1724]" class="Insecta" family="Carabidae" genus="Geocharis" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="45" phylum="Arthropoda" rank="species" species="rodriguesi">
|
|||
|
|
<emphasis id="B958EAA8F729932AFCD71B8415CEF961" box="[834,983,1700,1724]" italics="true" pageId="12" pageNumber="45">G. rodriguesi</emphasis>
|
|||
|
|
</taxonomicName>
|
|||
|
|
) were recorded inside the geographical limits of a protected area.
|
|||
|
|
</paragraph>
|
|||
|
|
</subSubSection>
|
|||
|
|
</treatment>
|
|||
|
|
</document>
|