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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.232.103649" ID-Pensoft-Pub="1314-2003-232-89" ID-Pensoft-UUID="DCC32AA52960584B8DFADEC0D4FE66F6" ModsDocID="1314-2003-232-89" checkinTime="1694651764838" checkinUser="pensoft" docAuthor="Julius, Avelinah, Siti-Munirah, Mat Yunoh &amp; Utteridge, Timothy M. A." docDate="2023" docId="10F8D44E3CFE595FB97F5B4B7A41E235" docLanguage="en" docName="PhytoKeys 232: 89-98" docOrigin="PhytoKeys 232" docPubDate="2023-09-13" docSource="http://dx.doi.org/10.3897/phytokeys.232.103649" docTitle="Ardisia recurvipetala Julius, Siti-Munirah &amp; Utteridge, sp. nov. (§ Crispardisia)" docType="treatment" docVersion="1" id="DCC32AA52960584B8DFADEC0D4FE66F6" lastPageNumber="89" masterDocId="DCC32AA52960584B8DFADEC0D4FE66F6" masterDocTitle="Ardisia recurvipetala (Primulaceae - Myrsinoideae), a new species from northern Peninsular Malaysia" masterLastPageNumber="98" masterPageNumber="89" pageNumber="89" updateTime="1694651764838" updateUser="pensoft">
<mods:mods id="212AD4737EBFF7C2384B92CCE068A710" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A38C86C8F64DE232F062D3F8C93F0F2F">
<mods:title id="AC41D29B088733178A1D63EC00D72286">Ardisia recurvipetala (Primulaceae - Myrsinoideae), a new species from northern Peninsular Malaysia</mods:title>
</mods:titleInfo>
<mods:name id="8C90C8D5F2C4E9879FB23F767FE40EE0" type="personal">
<mods:role id="2D926B6B42840E7C61CC5847B64A3437">
<mods:roleTerm id="88FFF98287B11955D967277287FBC69C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E77EC1EAB221A0A295F05F4B67652E68">Julius, Avelinah</mods:namePart>
<mods:nameIdentifier id="C8498E15C5BBC1BEED027FFB634C7748" type="ORCID">https://orcid.org/0000-0003-1991-1915</mods:nameIdentifier>
<mods:affiliation id="53120C0FEFC6B7F182626A5D1154D119">Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Jalan UMS, 88400 Kota Kinabalu, Sabah, Malaysia</mods:affiliation>
<mods:nameIdentifier id="9A613A120F1CB22348062CEF798CDFA5" type="email">plagiovel80@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:name id="C1F5BABE705C145D87552EF88AB758A8" type="personal">
<mods:role id="51CF2516D65EF8A2CDE08E1F2A2C39C8">
<mods:roleTerm id="9BEDD7C3E675F0D5C94AFB1BA58AB252">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1FE9933BB745C1BC3721A23C2795B6EC">Siti-Munirah, Mat Yunoh</mods:namePart>
<mods:nameIdentifier id="44F822D990380D590ED3B4F0D4FC1799" type="ORCID">https://orcid.org/0000-0002-5062-9988</mods:nameIdentifier>
<mods:affiliation id="980AA8FBDEE8A91BF50FF98460DDDAAC">Forest Research Institute Malaysia, Kepong, Selangor, 52109, Malaysia</mods:affiliation>
</mods:name>
<mods:name id="F8F3B6B106F2AAB2D92E5EC23C409192" type="personal">
<mods:role id="C6EF339DF80C0C7D5684F135FEED569C">
<mods:roleTerm id="29E13EFE656828D614ADB82C4B4A2512">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0B209F1F10A231C8EB2AF0C33A1F4752">Utteridge, Timothy M. A.</mods:namePart>
<mods:nameIdentifier id="2DB80884D9D438E3FDFCD5964E86A8C1" type="ORCID">https://orcid.org/0000-0003-2823-0337</mods:nameIdentifier>
<mods:affiliation id="1B6982AA2D86513B6201A1D60D49F3C3">Herbarium, Royal Botanic Gardens, Kew, Richmond, Surrey, TW 9 3 AE, UK</mods:affiliation>
</mods:name>
<mods:typeOfResource id="48CE7429F89AD3CB1935C6F096838611">text</mods:typeOfResource>
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<mods:titleInfo id="61CF961364B2669B57C6A6B41C8EE61D">
<mods:title id="4661D6987BFF5D8F0D873D50DBE4C4BB">PhytoKeys</mods:title>
</mods:titleInfo>
<mods:part id="EC5609BCA2C3E569CA30AA25C9460B60">
<mods:date id="8B946C552642607C5268A881255E7B21">2023</mods:date>
<mods:detail id="D70644172BF2B5E67A960BE0C21DD961" type="pubDate">
<mods:number id="91710B51810D6B0F06402219E72BB088">2023-09-13</mods:number>
</mods:detail>
<mods:detail id="8A49ECB21569A2974D26E9A262D4CB32" type="volume">
<mods:number id="C445273BDAD461438965C5308B5D106B">232</mods:number>
</mods:detail>
<mods:extent id="449B06DCE18D47A9034C6E5BC7EBD9E3" unit="page">
<mods:start id="2F3C7A685194459788AAC9003365573B">89</mods:start>
<mods:end id="6168F3D34FEBAA78BEC3E1830EFD13CA">98</mods:end>
</mods:extent>
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<mods:location id="B16B2BE7F6005C42E735D03F054A3E7E">
<mods:url id="577E96F140DBECA7A17574B2B218E5BA">http://dx.doi.org/10.3897/phytokeys.232.103649</mods:url>
</mods:location>
<mods:classification id="C8ECD40844D2BA850BADFB3B7E5FF354">journal article</mods:classification>
<mods:identifier id="7C1A24E401CE4BED672EE87CA3075718" type="DOI">http://dx.doi.org/10.3897/phytokeys.232.103649</mods:identifier>
<mods:identifier id="A10F087CEB6493240DB3F5B210637865" type="Pensoft-Pub">1314-2003-232-89</mods:identifier>
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<treatment id="10F8D44E3CFE595FB97F5B4B7A41E235" LSID="urn:lsid:plazi:treatment:10F8D44E3CFE595FB97F5B4B7A41E235" httpUri="http://treatment.plazi.org/id/10F8D44E3CFE595FB97F5B4B7A41E235" lastPageNumber="89" pageId="0" pageNumber="89">
<subSubSection id="28ABD0F9BC2CE5B6B2FE14C9FD438069" pageId="0" pageNumber="89" type="nomenclature">
<paragraph id="6C594CEA6AA68A487AC7697958D74620" pageId="0" pageNumber="89">
<taxonomicName id="DCC9BA8EF8F5DFFBBC9A780A57103D2C" LSID="10F8D44E-3CFE-595F-B97F-5B4B7A41E235" authority="Julius, Siti-Munirah &amp; Utteridge, sp. nov. (§ Crispardisia)" authorityName="Julius, Siti-Munirah &amp; Utteridge, sp. nov. (§ Crispardisia)" class="Magnoliopsida" family="Primulaceae" genus="Ardisia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ardisia recurvipetala" order="Ericales" pageId="0" pageNumber="89" phylum="Tracheophyta" rank="species" species="recurvipetala">Ardisia recurvipetala Julius, Siti-Munirah &amp; Utteridge, sp. nov. (§ Crispardisia)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="02E95DF2121FF8D7E673814F74E59D89" pageId="0" pageNumber="89" type="description">
<paragraph id="39C0CADD274F4F6A9A1A93D3D5FD7CD9" pageId="0" pageNumber="89">
<figureCitation id="874FB7FBF88F8878E7939BCAE273D5E4" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Ardisia recurvipetala A habitat (inset: Ai habit Aii inflorescence position on lateral shoot and main stem) B lateral branch with flowers and young fruits C, D flowers." figureDoi="10.3897/phytokeys.232.103649.figure1" httpUri="https://binary.pensoft.net/fig/905304" pageId="0" pageNumber="89">Figs 1</figureCitation>
<figureCitation id="DEC3DBBF790030C6E452987A92296F11" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Leaf margin showing vascularised glands (yellow arrow) on the incision between crenation and the pustule-like structures (red arrow) along the crenation." figureDoi="10.3897/phytokeys.232.103649.figure2" httpUri="https://binary.pensoft.net/fig/905305" pageId="0" pageNumber="89">, 2</figureCitation>
<figureCitation id="71CEB4705CABC8095DD5EA31EDB405CE" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ardisia recurvipetala A flowering branches B flower bud (i), mature flower (ii) and flower with corolla removed (iii) C flower (spread) showing the stamen arrangement D adaxial view of sepal E ventral (left) and dorsal surfaces (right) of anther F anterior (left) and lateral views (right) of ovary G leaf margin with venation details and bacterial nodule H pustule-like structure on leaf margin crenation. Illustration by Mohd Aidil Nordin." figureDoi="10.3897/phytokeys.232.103649.figure3" httpUri="https://binary.pensoft.net/fig/905306" pageId="0" pageNumber="89">, 3</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="DE7F023ED1F0AD4B1352B4B5031A7FC2" pageId="0" pageNumber="89" type="diagnosis">
<paragraph id="133D37A6AB8A0FF1BD86EBCADCCF4E40" pageId="0" pageNumber="89">Diagnosis.</paragraph>
<paragraph id="A4AB246F2E91D3D3BAED6F52550D8027" pageId="0" pageNumber="89">
<taxonomicName id="A1243109900B5F951F5FBBB070BF7A29" class="Magnoliopsida" family="Myrsinoideae" genus="Ardisia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ardisia recurvipetala" order="Ericales" pageId="0" pageNumber="89" phylum="Tracheophyta" rank="species" species="recurvipetala">
<emphasis id="48CEABBB6B82601DB1AF782C8C62444C" italics="true" pageId="0" pageNumber="89">Ardisia recurvipetala</emphasis>
</taxonomicName>
is the only
<taxonomicName id="8B60D0E8F7A669EBB8DB57A895501244" class="Magnoliopsida" family="Myrsinoideae" genus="Ardisia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ardisia" order="Ericales" pageId="0" pageNumber="89" phylum="Tracheophyta" rank="genus">
<emphasis id="3EE8CF6EFBB82656C81B6A1F40F743E8" italics="true" pageId="0" pageNumber="89">Ardisia</emphasis>
</taxonomicName>
species with the following combination of characters: leaf margins with both vascularised glands and pustule-like structures, lamina raised between the venation giving a somewhat bullate appearance, inflorescences on the main shoot and specialised lateral branches, white flowers with recurved petals with a pink patch at the base of the corolla lobes.
</paragraph>
<caption id="18E0DF8EE97C24F5EE492FE909E3DCCE" doi="10.3897/phytokeys.232.103649.figure1" httpUri="https://binary.pensoft.net/fig/905304" pageId="0" pageNumber="89" start="Figure 1" startId="F1">
<paragraph id="36F9A3F5CC59F7AE2E90102F1FEC1AF7" pageId="0" pageNumber="89">
<emphasis id="F1F7BFFAD2DC815AE6078731D2C80E7C" bold="true" pageId="0" pageNumber="89">Figure 1.</emphasis>
<taxonomicName id="8C9A35070203B5CDF07384E271FCB382" class="Magnoliopsida" family="Myrsinoideae" genus="Ardisia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ardisia recurvipetala" order="Ericales" pageId="0" pageNumber="89" phylum="Tracheophyta" rank="species" species="recurvipetala">
<emphasis id="0808ED7523A88BFBB0D5B39D49950BD1" italics="true" pageId="0" pageNumber="89">Ardisia recurvipetala</emphasis>
</taxonomicName>
<emphasis id="E98B6A8A5FEDD16CEF0162FE14E07B76" bold="true" pageId="0" pageNumber="89">A</emphasis>
habitat (inset:
<emphasis id="F9549E51A508F225B0A9AEB18705305F" bold="true" pageId="0" pageNumber="89">Ai</emphasis>
habit
<emphasis id="E68E4B65CFFBD5459389E07B39B3A7AB" bold="true" pageId="0" pageNumber="89">Aii</emphasis>
inflorescence position on lateral shoot and main stem)
<emphasis id="022D02DFE945BFE901E2FADD9CB75212" bold="true" pageId="0" pageNumber="89">B</emphasis>
lateral branch with flowers and young fruits
<emphasis id="67097C21A299DAE081BDEDE0904F86B3" bold="true" pageId="0" pageNumber="89">C, D</emphasis>
flowers.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="5FA2157F93FA9428230FEC9ED347FF27" pageId="0" pageNumber="89" type="materials_examined">
<paragraph id="F17A2EA9F16E3E14A05324A552545D8F" pageId="0" pageNumber="89">
<typeStatus id="D218F689B87FD018F321BD4C2EFBA091">Type</typeStatus>
.
</paragraph>
<paragraph id="789635AA4FF267C4FAC63A2D7EEC1F41" pageId="0" pageNumber="89">
<collectingCountry id="71D45459791B16FF5F4DAE6912111E35" name="Malaysia">Malaysia</collectingCountry>
. Peninsular
<materialsCitation id="206145E1D1216BAD38A6F0CAE21289E7" accessionNumber="FRI 98670" collectingDate="2022-06-15" country="Malaysia" elevation="201" latitude="5.0168056" location="Taman Negeri Kenyir" longLatPrecision="1" longitude="102.54164" municipality="Hulu" specimenCount="1" stateProvince="Terengganu" typeStatus="holotype">
<collectingCountry id="179B331DF8951BD8A44BA1600A3E023E" name="Malaysia">Malaysia</collectingCountry>
:
<collectingRegion id="899AE1FCC82012AD74F076F1E7F98EC4" country="Malaysia" name="Terengganu">Terengganu</collectingRegion>
,
<collectingMunicipality id="E1F15C6CC688ACFA6831E95690075F7C">Hulu</collectingMunicipality>
<collectingRegion id="023E861FCDFEC81BF67CA1FF8CBC3EA1" country="Malaysia" name="Terengganu">Terengganu Distr.</collectingRegion>
,
<location id="2BFEA97925E70F1DC7712AD92174631D" LSID="urn:lsid:plazi:treatment:10F8D44E3CFE595FB97F5B4B7A41E235:2BFEA97925E70F1DC7712AD92174631D" country="Malaysia" latitude="5.0168056" longLatPrecision="1" longitude="102.54164" municipality="Hulu" name="Taman Negeri Kenyir" stateProvince="Terengganu">Taman Negeri Kenyir</location>
,
<geoCoordinate id="67C6688D7C89266910CC2CE3DC6CD785" degrees="5" direction="north" minutes="01" orientation="latitude" precision="1" seconds="00.5" value="5.0168056">5°01'00.5&quot;N</geoCoordinate>
,
<geoCoordinate id="2AC1C085591A3327BE0F25030F325A93" degrees="102" direction="east" minutes="32" orientation="longitude" precision="1" seconds="29.9" value="102.54164">102°32'29.9&quot;E</geoCoordinate>
,
<quantity id="9F664053B6CC9E8B31F477466CE94C52" metricMagnitude="2" metricUnit="m" metricValue="2.01" unit="m" value="201.0">
<elevation id="D7AACF8F054B570BCE1C541217417847" metricMagnitude="2" metricUnit="m" metricValue="2.01" unit="m" value="201.0">201 m</elevation>
</quantity>
elev.,
<collectingDate id="E47924ACA8EC50BC248272933EFFBA48" value="2022-06-15">15 June 2022</collectingDate>
(fl. &amp; fr.),
<emphasis id="DAE0D32C313191633F53D5AC4D0D168A" italics="true" pageId="0" pageNumber="89">
Siti Munirah
<accessionNumber id="B286A36C614AD7999EA703FC65B573A0" httpUri="http://www.ncbi.nlm.nih.gov/protein/FRI98670">FRI 98670</accessionNumber>
</emphasis>
(
<typeStatus id="469B6A7E1FD85D8753EDBF3C04DF2A26">holotype</typeStatus>
KEP!; isotype BORH!)
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.
</paragraph>
<caption id="3384429963E45CD69F961CA9FF7B6113" doi="10.3897/phytokeys.232.103649.figure2" httpUri="https://binary.pensoft.net/fig/905305" pageId="0" pageNumber="89" start="Figure 2" startId="F2">
<paragraph id="CEC7CDF652126BA705A4B42EC23910FE" pageId="0" pageNumber="89">
<emphasis id="3A8CEB657AF9F0F3C9C387A2C491D3C9" bold="true" pageId="0" pageNumber="89">Figure 2.</emphasis>
Leaf margin showing vascularised glands (yellow arrow) on the incision between crenation and the pustule-like structures (red arrow) along the crenation.
</paragraph>
</caption>
<paragraph id="F391D263B9248DCA2FE02E838EA809BB" pageId="0" pageNumber="89">
<emphasis id="1122325FEF0C654A8BFFC6E5DB0782E5" bold="true" italics="true" pageId="0" pageNumber="89">Woody shrub</emphasis>
ca.
<quantity id="B32F58FE02F6BB141351D29C5A32B554" metricMagnitude="0" metricUnit="m" metricValue="1.0" unit="m" value="1.0">1 m</quantity>
tall.
<emphasis id="9C1056E7C8192C8C25DF3C5AF9BF09AA" bold="true" italics="true" pageId="0" pageNumber="89">Branches</emphasis>
<quantity id="477E2B03BD5030D75939377600F6E999" metricMagnitude="-2" metricUnit="m" metricValue="8.75" metricValueMax="12.5" metricValueMin="5.0" unit="cm" value="8.75" valueMax="12.5" valueMin="5.0">5-12.5 cm</quantity>
long.
<emphasis id="8209CD73C11663FF9E12AE7082ECEA6F" bold="true" italics="true" pageId="0" pageNumber="89">Indumentum</emphasis>
absent except for short, white or brown, simple eglandular and glandular hairs on reproductive parts.
<emphasis id="D1A4801DE3AD3D43C282FA05103C0499" bold="true" italics="true" pageId="0" pageNumber="89">Leaves</emphasis>
spirally arranged; petioles (0.5-)
<quantity id="54E2DD838FD7C774B9881DD85F0DB69C" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" unit="cm" value="1.25" valueMax="1.5" valueMin="1.0">1-1.5 cm</quantity>
long, winged by the decurrent leaf base, glabrous; lamina chartaceous, with sparse black gland dots throughout abaxially, raised between the venation; usually broadly elliptic, occasionally oblanceolate, (12.5-)17.5-20.5
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<quantity id="5AD9AE8C6ABD7A0537197D2B54436068" metricMagnitude="-2" metricUnit="m" metricValue="6.0" metricValueMax="7.5" metricValueMin="4.5" unit="cm" value="6.0" valueMax="7.5" valueMin="4.5">4.5-7.5 cm</quantity>
(excluding reduced leaves subtending inflorescences on specialised lateral branches); young leaves light green, mature leaves dark green above, pale green beneath; base cuneate; margin crenate with vascularised glands (bacterial nodules) at the incision between crenatures and with pustule-like structures along the crenations from projecting venation; apex acute to obtuse; glabrous on both surfaces; mid-rib flat above, raised below; lateral veins 9-14 pairs, irregularly spaced, joining at the marginal vein, distinct on the adaxial surface, prominent on abaxial side; intersecondary veins sometimes present; intercostal veins obscure.
<emphasis id="6F4E78FEEE1C4D15A4D9D02DA5D146B6" bold="true" italics="true" pageId="0" pageNumber="89">Inflorescences</emphasis>
subsessile, terminal on main shoot and on relatively short specialised lateral branches with 1-2(-3) subtending leaves, condensed racemose, ca. 2
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<quantity id="DACD4D52F8BCF4F23B3CC9F5819F0824" metricMagnitude="-2" metricUnit="m" metricValue="3.0" unit="cm" value="3.0">3 cm</quantity>
, unbranched, 10-18-flowered.
<emphasis id="1B6812A92E6646D07E73B861DBD65598" bold="true" italics="true" pageId="0" pageNumber="89">Flowers</emphasis>
5-merous; pedicels brownish-red,
<quantity id="01E665F4E69E5A0568F37AC4A6421FC1" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" unit="cm" value="1.25" valueMax="1.5" valueMin="1.0">1-1.5 cm</quantity>
long, slender, densely glandular hairy with globular tips, covered with dense, brown gland dots; sepals yellowish-green, pale pink at base, not overlapping, covered with dense brown gland dots, triangular-ovate, 1.5-2
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<quantity id="E69B0EF1D663364CA9A8B10D787A358C" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1-1.5 mm</quantity>
, sparsely glandular hairy abaxially, glabrous adaxially, margin ciliate with laxly spaced, simple, pale brown hairs, apex obtuse; corolla tube ca.
<quantity id="D696E00D31CE3F10554E20F129EC5E74" metricMagnitude="-4" metricUnit="m" metricValue="5.0" unit="mm" value="0.5">0.5 mm</quantity>
long, lobes 5, white except the creamy-white apex and pinkish base, covered with dense, brown gland dots, lanceolate, 6-7
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<quantity id="C916ADE9C4C44D9CF12233D5D081BB4A" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3-4 mm</quantity>
, glabrous on both surfaces, apex acute, strongly recurved at anthesis; stamens 5, yellowish, subsessile, anthers narrowly lanceolate-mucronate, 5-6
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<quantity id="FA0011D9EB550A676AABE71CD5DED6D1" metricMagnitude="-3" metricUnit="m" metricValue="1.5" unit="mm" value="1.5">1.5 mm</quantity>
, glabrous throughout, gland dotted abaxially, thecae not locellate, dehiscent by longitudinal slits; ovary subglobose, ca. 2
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<quantity id="63611580541AA1CB27B1FF0E436F8AC8" metricMagnitude="-3" metricUnit="m" metricValue="1.5" unit="mm" value="1.5">1.5 mm</quantity>
, glabrous, ovules ca. 9 arranged in 1-series, style and stigma slender, ca.
<quantity id="76B7ACE49CA340D2DC3A0A2F5E37062E" metricMagnitude="-3" metricUnit="m" metricValue="5.3" unit="mm" value="5.3">5.3 mm</quantity>
long, glabrous.
<emphasis id="85A47FCCCFDADDB6C0CF3813CB9C7174" bold="true" italics="true" pageId="0" pageNumber="89">Young fruits</emphasis>
globose, with dense gland dots, 4-6
<normalizedToken id="B624C4CB3AF111A43C47084727FFE71C" originalValue="×">x</normalizedToken>
<quantity id="B417A0137AA5F6EC20077D200E0D5D54" metricMagnitude="-3" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" unit="mm" value="5.0" valueMax="6.0" valueMin="4.0">4-6 mm</quantity>
, green, glabrous; pedicels becoming thickened and obconically flared,
<quantity id="925CB4E82B32E45C960410B5A712DEAC" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" unit="cm" value="1.9" valueMax="2.0" valueMin="1.8">1.8-2 cm</quantity>
long. Mature fruits not observed.
</paragraph>
<caption id="47C4E98DD794C4CFC40FF994F0F667F9" doi="10.3897/phytokeys.232.103649.figure3" httpUri="https://binary.pensoft.net/fig/905306" pageId="0" pageNumber="89" start="Figure 3" startId="F3">
<paragraph id="2285BAF2D7EDC4F55769409ACD562205" pageId="0" pageNumber="89">
<emphasis id="B82A2F6D4F6D077C60144496536472B5" bold="true" pageId="0" pageNumber="89">Figure 3.</emphasis>
<taxonomicName id="D733AC6054FE3089DEB995A57032C7C9" class="Magnoliopsida" family="Myrsinoideae" genus="Ardisia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ardisia recurvipetala" order="Ericales" pageId="0" pageNumber="89" phylum="Tracheophyta" rank="species" species="recurvipetala">
<emphasis id="D010BB3B06E89D7D57306E1CBA0500EB" italics="true" pageId="0" pageNumber="89">Ardisia recurvipetala</emphasis>
</taxonomicName>
<emphasis id="3C945869BC0F0E10F5F4BB5A3B11BC00" bold="true" pageId="0" pageNumber="89">A</emphasis>
flowering branches
<emphasis id="1BC4709EEA82958B4AFB35A84D4B828B" bold="true" pageId="0" pageNumber="89">B</emphasis>
flower bud (
<emphasis id="378CCE7B666C9B96C117139670ACB4A2" bold="true" pageId="0" pageNumber="89">i</emphasis>
), mature flower (
<emphasis id="6694F6B2122C7570F54A17FD1FA5BDC2" bold="true" pageId="0" pageNumber="89">ii</emphasis>
) and flower with corolla removed (
<emphasis id="85DCA673AF1F3CEBC041EE6EFF0F583A" bold="true" pageId="0" pageNumber="89">iii</emphasis>
)
<emphasis id="BCAC7FE2CDA99A39A6B6B45BB2A891E1" bold="true" pageId="0" pageNumber="89">C</emphasis>
flower (spread) showing the stamen arrangement
<emphasis id="F8B171A804B8ED29DAC8D3C8C080A1CC" bold="true" pageId="0" pageNumber="89">D</emphasis>
adaxial view of sepal
<emphasis id="02A6550FD300921AA935A12771587E86" bold="true" pageId="0" pageNumber="89">E</emphasis>
ventral (left) and dorsal surfaces (right) of anther
<emphasis id="8B046C79BDA07744E0930046A124E07D" bold="true" pageId="0" pageNumber="89">F</emphasis>
anterior (left) and lateral views (right) of ovary
<emphasis id="359122285905771C9C0B2E67E02B6084" bold="true" pageId="0" pageNumber="89">G</emphasis>
leaf margin with venation details and bacterial nodule
<emphasis id="F9C3A5A3EA19F4F0DE29B1EE1A86AEBA" bold="true" pageId="0" pageNumber="89">H</emphasis>
pustule-like structure on leaf margin crenation. Illustration by Mohd Aidil Nordin.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="2558CC32372B090793B75A4038F17227" pageId="0" pageNumber="89" type="distribution">
<paragraph id="EDA2E73BF11921D6E9C82CB5D3283DDE" pageId="0" pageNumber="89">Distribution.</paragraph>
<paragraph id="C34E1977D6A2601C5D5F15CD19532C81" pageId="0" pageNumber="89">
Endemic to Peninsular Malaysia, Terengganu (Taman Negeri Kenyir); currently known only from the type locality (Fig.
<figureCitation id="8BB1124E54F51C058E5F37EC8F2D1F63" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Map showing the type locality of Ardisia recurvipetala in Peninsular Malaysia." figureDoi="10.3897/phytokeys.232.103649.figure4" httpUri="https://binary.pensoft.net/fig/905307" pageId="0" pageNumber="89">4</figureCitation>
).
</paragraph>
<caption id="8194DE1F3CE46AAE39871595DF1CF492" doi="10.3897/phytokeys.232.103649.figure4" httpUri="https://binary.pensoft.net/fig/905307" pageId="0" pageNumber="89" start="Figure 4" startId="F4">
<paragraph id="79F6FCD1F1D2F186AAA6BB6610EC52BB" pageId="0" pageNumber="89">
<emphasis id="42FF95245F22DE962B02D7411BEDB068" bold="true" pageId="0" pageNumber="89">Figure 4.</emphasis>
Map showing the type locality of
<taxonomicName id="707231F0641AC7C2D77AA37F575853DB" class="Magnoliopsida" family="Myrsinoideae" genus="Ardisia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ardisia recurvipetala" order="Ericales" pageId="0" pageNumber="89" phylum="Tracheophyta" rank="species" species="recurvipetala">
<emphasis id="E27B2C11A054641F516E349A39ABADB9" italics="true" pageId="0" pageNumber="89">Ardisia recurvipetala</emphasis>
</taxonomicName>
in Peninsular Malaysia.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="F875765B613CC2E8BE26646C98F8081A" pageId="0" pageNumber="89" type="ecology">
<paragraph id="7FEFB1A24BC620519876B31EEC0C3D50" pageId="0" pageNumber="89">Ecology.</paragraph>
<paragraph id="620FB020222F600452B983FA11981B20" pageId="0" pageNumber="89">Growing in a lowland dipterocarp forest at 201 m elev. in a shaded area not far from a small stream.</paragraph>
</subSubSection>
<subSubSection id="8340ABF6C9D520039F1C501263AA3709" pageId="0" pageNumber="89" type="etymology">
<paragraph id="DE922F26390C8E48670A1311EDCB70C0" pageId="0" pageNumber="89">Etymology.</paragraph>
<paragraph id="764C96CCC6CD8CC87ED5C05F93E2C6C4" pageId="0" pageNumber="89">The species epithet is derived from its recurved corolla lobes.</paragraph>
</subSubSection>
<subSubSection id="B5D3CA50464105AA192BA585184B8D8E" pageId="0" pageNumber="89" type="conservation status">
<paragraph id="2CAD5396E41EF98554D274E23A911E46" pageId="0" pageNumber="89">Conservation status.</paragraph>
<paragraph id="F9B2E375DCD479BE623A36141D91FE38" pageId="0" pageNumber="89">
Data Deficient (DD).
<taxonomicName id="63F727C9467F568CCFFB10D21F37FF5B" class="Magnoliopsida" family="Myrsinoideae" genus="Ardisia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ardisia recurvipetala" order="Ericales" pageId="0" pageNumber="89" phylum="Tracheophyta" rank="species" species="recurvipetala">
<emphasis id="54ACB3E82BF7314E09F0E3CD80F2B477" italics="true" pageId="0" pageNumber="89">Ardisia recurvipetala</emphasis>
</taxonomicName>
is known from fewer than five individuals in flower and fruit collected from intact forest patches within the Taman Negeri Kenyir. The sites where the taxon was found were previously part of the Tembat Forest Reserve before it was gazetted into Taman Negeri Kenyir in 2018. Further surveys are needed to understand the threats at the type locality and if the species is distributed outside the current area and, until these data are obtained, the species is assessed as Data Deficient (DD) (
<bibRefCitation id="3902FEC638768B2B5F6D9E437FCA34F0" author="Chua, LSL" journalOrPublisher="Science Press, Beijing, and Missouri Botanical Garden Press, St Louis" pageId="0" pageNumber="89" refId="B2" refString="Chua, LSL, Saw, LG, 2006. Malaysia Plant Red List: Guide for Contributors. Forest Research Institute Malaysia, 28 pp." title="Malaysia Plant Red List: Guide for Contributors. Forest Research Institute Malaysia, 28 pp." year="2006">Chua and Saw 2006</bibRefCitation>
;
<bibRefCitation id="EF941B45C63B6DAF25FEBFD8EE61D192" author="Hu, CM" journalOrPublisher="Museum National d'Histoire Naturelle, Paris" pageId="0" pageNumber="89" refId="B5" refString="IUCN Standards and Petitions Committee (2022) Guidelines for Using the IUCN Red ListCategories and Criteria. Version 15. Prepared by the Standards and Petitions Committee. [Accessed 22 February 2023]" year="2004">IUCN Standards and Petitions Committee 2022</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="ECDCB914908C445B158938B2A5F5B1D9" pageId="0" pageNumber="89" type="notes">
<paragraph id="F0C8D7F75E1A39B39490EF8AD016F10C" pageId="0" pageNumber="89">Notes.</paragraph>
<paragraph id="D497054C1281E85B606597CF49ABFFAE" pageId="0" pageNumber="89">
Peninsular Malaysian members of §
<taxonomicName id="DD16B0300B04C9CDD548DBC790C1531B" genus="Crispardisia" lsidName="Crispardisia" pageId="0" pageNumber="89" rank="genus">
<emphasis id="5DC75D2C5EBB7F5BA44C126317D5E30A" italics="true" pageId="0" pageNumber="89">Crispardisia</emphasis>
</taxonomicName>
can be placed into two informal groups, based on inflorescence position (
<bibRefCitation id="EF9BEB3AA0EC10219B8D77937BC1B4DC" author="Stone, BC" journalOrPublisher="Academy of Natural Sciences of Philadelphia" pageId="0" pageNumber="89" pagination="263 - 306" refId="B28" refString="Stone, BC, 1989. New and noteworthy Malesian Myrsinaceae, III. On the genus Ardisia Sw. in Borneo. Proceedings. Academy of Natural Sciences of Philadelphia 141: 263 - 306" title="New and noteworthy Malesian Myrsinaceae, III. On the genus Ardisia Sw. in Borneo. Proceedings." volume="141" year="1989">Stone 1989</bibRefCitation>
;
<bibRefCitation id="E07A3F9B901141E9FBB20033DC1F745A" DOI="https://doi.org/10.11646/phytotaxa.480.1.7" author="Julius, A" journalOrPublisher="Phytotaxa" pageId="0" pageNumber="89" pagination="79 - 84" refId="B8" refString="Julius, A, Utteridge, TMA, 2021. Ardisia recurvisepala (Primulaceae - Myrsinoideae), a new species from Peninsular Malaysia. Phytotaxa 480 (1): 79 - 84, DOI: https://doi.org/10.11646/phytotaxa.480.1.7" title="Ardisia recurvisepala (Primulaceae - Myrsinoideae), a new species from Peninsular Malaysia." url="https://doi.org/10.11646/phytotaxa.480.1.7" volume="480" year="2021">Julius and Utteridge 2021</bibRefCitation>
). The first group has inflorescences strictly terminal on specialised lateral leafy branches, while the second group has inflorescences lateral (axillary) and/or terminal on the main stem and/or lateral branches.
<taxonomicName id="FEFC4B3239C83773FE373D9026535B26" class="Magnoliopsida" family="Myrsinoideae" genus="Ardisia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ardisia recurvipetala" order="Ericales" pageId="0" pageNumber="89" phylum="Tracheophyta" rank="species" species="recurvipetala">
<emphasis id="AB941F0A439F3E858F490419A5AFBB4F" italics="true" pageId="0" pageNumber="89">Ardisia recurvipetala</emphasis>
</taxonomicName>
falls into the second group, in which the majority of species in §
<taxonomicName id="ED8644713F30B5610F5319431992D712" genus="Crispardisia" lsidName="Crispardisia" pageId="0" pageNumber="89" rank="genus">
<emphasis id="ACEB0CA6FF4F79F35730EDDE7F49CC3E" italics="true" pageId="0" pageNumber="89">Crispardisia</emphasis>
</taxonomicName>
belong, including six species from Peninsular Malaysia. Other than characters mentioned in the diagnosis, the new species can be easily recognised by its unbranched and subsessile inflorescences terminal on the main shoot and on relatively short specialised lateral branches with 1 to 2(-3) subtending leaves, tiny sepals (1.5-2
<normalizedToken id="121A937EB13642E06C098A85163B97AA" originalValue="×">x</normalizedToken>
1-1.5 mm) that are yellowish-green and pinkish at base and corolla lobes that are strongly recurved at anthesis (Figs
<figureCitation id="86CA0608979FE95EFADF4035DDD30045" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Ardisia recurvipetala A habitat (inset: Ai habit Aii inflorescence position on lateral shoot and main stem) B lateral branch with flowers and young fruits C, D flowers." figureDoi="10.3897/phytokeys.232.103649.figure1" httpUri="https://binary.pensoft.net/fig/905304" pageId="0" pageNumber="89">1B</figureCitation>
, 3Bii).
</paragraph>
<paragraph id="364CCFA6DA87B0991A8745799FF683F7" pageId="0" pageNumber="89">
Unlike other members of §
<taxonomicName id="9CA7D3B74F9C5ABD73086736557C2C44" genus="Crispardisia" lsidName="Crispardisia" pageId="0" pageNumber="89" rank="genus">
<emphasis id="5EBA2DAD9E2EFD6C849C7B3FE801C09D" italics="true" pageId="0" pageNumber="89">Crispardisia</emphasis>
</taxonomicName>
, the leaf margins of the new species have both vascularised glands at the incisions between the crenatures and pustule-like structures along the crenations (Fig.
<figureCitation id="FE8E5B8D7A4FB81FC86AC6D55AAF3004" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Leaf margin showing vascularised glands (yellow arrow) on the incision between crenation and the pustule-like structures (red arrow) along the crenation." figureDoi="10.3897/phytokeys.232.103649.figure2" httpUri="https://binary.pensoft.net/fig/905305" pageId="0" pageNumber="89">2</figureCitation>
); these result from the venation projecting from the leaf margin and should not be mistaken for bacterial nodules. This combination of vascularised glands and pustule-like structures is not observed in other Peninsular Malaysian members of §
<taxonomicName id="24CAFB615239B2F92E3450A1FF684783" genus="Crispardisia" lsidName="Crispardisia" pageId="0" pageNumber="89" rank="genus">
<emphasis id="7AF76F71C7849B17EF1B9BD86F62DCFD" italics="true" pageId="0" pageNumber="89">Crispardisia</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="CC3857035265BC64962F35CAA0C32199" pageId="0" pageNumber="89">
The new species is similar to
<taxonomicName id="F66CA4840CAB4734A326B001691AEB5B" class="Magnoliopsida" family="Myrsinoideae" genus="Ardisia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ardisia sphenobasis" order="Ericales" pageId="0" pageNumber="89" phylum="Tracheophyta" rank="species" species="sphenobasis">
<emphasis id="FCD72965EE77C60836C6364EFE046D49" italics="true" pageId="0" pageNumber="89">Ardisia sphenobasis</emphasis>
</taxonomicName>
in having 17.5-23 cm long leaves (excluding reduced leaves subtending inflorescences on specialised lateral branches), but it has inflorescences that are terminal on the main shoot and specialised lateral branches (vs. strictly lateral on main stem). The whitish corollas of the new species are somewhat similar to those of
<taxonomicName id="055F3EE6B11F4254A7BD5479E7043D93" class="Dicotyledones" family="Primulaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. villosa" order="Ericales" pageId="0" pageNumber="89" phylum="Angiospermae" rank="species" species="villosa">
<emphasis id="437A77CF6D27F85C476A647462B197E9" italics="true" pageId="0" pageNumber="89">A. villosa</emphasis>
</taxonomicName>
and
<taxonomicName id="611EBC75E39340808A9BAF71801A0676" class="Dicotyledones" family="Primulaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. crenata" order="Ericales" pageId="0" pageNumber="89" phylum="Angiospermae" rank="species" species="crenata">
<emphasis id="51CEA2105DE815F67A97F34C81B0730A" italics="true" pageId="0" pageNumber="89">A. crenata</emphasis>
</taxonomicName>
. However,
<taxonomicName id="2056D709B9747FB8ED5174720D08188E" class="Dicotyledones" family="Primulaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. recurvipetala" order="Ericales" pageId="0" pageNumber="89" phylum="Angiospermae" rank="species" species="recurvipetala">
<emphasis id="3AD8B230E9A0A4D53E88B0ADEEBF300E" italics="true" pageId="0" pageNumber="89">A. recurvipetala</emphasis>
</taxonomicName>
(Fig.
<figureCitation id="7B01F558CB1D96637A0D31E20C06E801" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Ardisia recurvipetala A flowering branches B flower bud (i), mature flower (ii) and flower with corolla removed (iii) C flower (spread) showing the stamen arrangement D adaxial view of sepal E ventral (left) and dorsal surfaces (right) of anther F anterior (left) and lateral views (right) of ovary G leaf margin with venation details and bacterial nodule H pustule-like structure on leaf margin crenation. Illustration by Mohd Aidil Nordin." figureDoi="10.3897/phytokeys.232.103649.figure3" httpUri="https://binary.pensoft.net/fig/905306" pageId="0" pageNumber="89">3</figureCitation>
) differs in its largely glabrous flowers, except for short, simple eglandular and glandular hairs on the pedicels and calyx (vs. hairy throughout with long, villous hairs in
<taxonomicName id="5D33F24649A4DE0DE7CB545D8B10FBB1" class="Dicotyledones" family="Primulaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. villosa" order="Ericales" pageId="0" pageNumber="89" phylum="Angiospermae" rank="species" species="villosa">
<emphasis id="D9725F278E36C165FB87D20D7C737950" italics="true" pageId="0" pageNumber="89">A. villosa</emphasis>
</taxonomicName>
). Compared to
<taxonomicName id="530336DF79D3339912A16A526C752315" class="Dicotyledones" family="Primulaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. crenata" order="Ericales" pageId="0" pageNumber="89" phylum="Angiospermae" rank="species" species="crenata">
<emphasis id="282413C02CF15CE87F29C3BDBC2DCACF" italics="true" pageId="0" pageNumber="89">A. crenata</emphasis>
</taxonomicName>
, a widespread species with a whitish to pinkish corolla, branched and unbranched inflorescences and a flat and coriaceous lamina, the new species has corolla lobes with a creamy-white apex and a pink patch at the base, strictly unbranched inflorescences and chartaceous leaves with the lamina raised between the venation.
</paragraph>
<paragraph id="BFEE82F6CEF80331518DFCC3C7747670" pageId="0" pageNumber="89">
This latest addition brings the number of §
<taxonomicName id="0806B3053B8B4085670F7BEFA5C2CE58" genus="Crispardisia" lsidName="Crispardisia" pageId="0" pageNumber="89" rank="genus">
<emphasis id="D20AAE6BD4E7EC66AC4150DA39893575" italics="true" pageId="0" pageNumber="89">Crispardisia</emphasis>
</taxonomicName>
species native to Peninsular Malaysia to ten. Of these, four species, including the new one, are endemic to Peninsular Malaysia:
<taxonomicName id="AB5F83FEEAF373AE140DF1899A1DE81D" class="Dicotyledones" family="Primulaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. lankawiensis" order="Ericales" pageId="0" pageNumber="89" phylum="Angiospermae" rank="species" species="lankawiensis">
<emphasis id="8BAE73DD6F74B34DA9A1EC105A25189A" italics="true" pageId="0" pageNumber="89">A. lankawiensis</emphasis>
</taxonomicName>
King &amp; Gamble,
<taxonomicName id="882AB37A04EB5468AE751CA185B4E05E" class="Dicotyledones" family="Primulaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. minor" order="Ericales" pageId="0" pageNumber="89" phylum="Angiospermae" rank="species" species="minor">
<emphasis id="5A26593DF3FF5FFBC7135920C8FFAD6A" italics="true" pageId="0" pageNumber="89">A. minor</emphasis>
</taxonomicName>
King &amp; Gamble,
<taxonomicName id="8A6BFB8EFB7C901A886859356DFF6588" class="Dicotyledones" family="Primulaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. recurvipetala" order="Ericales" pageId="0" pageNumber="89" phylum="Angiospermae" rank="species" species="recurvipetala">
<emphasis id="97874D52AA3BC63B7A7919E27ED5FE62" italics="true" pageId="0" pageNumber="89">A. recurvipetala</emphasis>
</taxonomicName>
and
<taxonomicName id="1AC7F8BA82E3C6ABF7BC6E4B2E56F3D6" class="Dicotyledones" family="Primulaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. recurvisepala" order="Ericales" pageId="0" pageNumber="89" phylum="Angiospermae" rank="species" species="recurvisepala">
<emphasis id="93FD16939A70F4E10A8161959ECDD11F" italics="true" pageId="0" pageNumber="89">A. recurvisepala</emphasis>
</taxonomicName>
Julius &amp; Utteridge.
</paragraph>
<paragraph id="D735B56E2F25052AF61428AD4B975705" pageId="0" pageNumber="89">
We consider species of
<taxonomicName id="05FB194EA11064B0CFAF2A5387939C49" class="Magnoliopsida" family="Myrsinoideae" genus="Ardisia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ardisia" order="Ericales" pageId="0" pageNumber="89" phylum="Tracheophyta" rank="genus">
<emphasis id="634FE94F6BCCAF34B89824FD40343063" italics="true" pageId="0" pageNumber="89">Ardisia</emphasis>
</taxonomicName>
as important indicators of tropical and subtropical forest quality. Whilst some species are found in a range of habitats (e.g.
<taxonomicName id="F74D9A74592861D607DCC39BB4B1BE1B" class="Dicotyledones" family="Primulaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. elliptica" order="Ericales" pageId="0" pageNumber="89" phylum="Angiospermae" rank="species" species="elliptica">
<emphasis id="00E8FB10B1B5844A13852DB413AB5DFA" italics="true" pageId="0" pageNumber="89">A. elliptica</emphasis>
</taxonomicName>
Thunb.), many species have restricted distributions and habitat requirements, including several species of §
<taxonomicName id="280BFE256B0AF2863A8FC1D172F873AE" genus="Crispardisia" lsidName="Crispardisia" pageId="0" pageNumber="89" rank="genus">
<emphasis id="B668A91F1BF10E008CB80F2B027DC20E" italics="true" pageId="0" pageNumber="89">Crispardisia</emphasis>
</taxonomicName>
. For example, in Peninsular Malaysia,
<taxonomicName id="1D279A87BEB69826384EFE71D6BCA035" class="Dicotyledones" family="Primulaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. lankawiensis" order="Ericales" pageId="0" pageNumber="89" phylum="Angiospermae" rank="species" species="lankawiensis">
<emphasis id="C1C1ABFF7F0AF7AD18A3B7A7799009E0" italics="true" pageId="0" pageNumber="89">A. lankawiensis</emphasis>
</taxonomicName>
is restricted to limestone habitats on Langkawi Islands and, in Thailand,
<taxonomicName id="624275313D83EF45CFCE8B9CDD8BC9F3" class="Dicotyledones" family="Primulaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. pilosa" order="Ericales" pageId="0" pageNumber="89" phylum="Angiospermae" rank="species" species="pilosa">
<emphasis id="D088B5D98E1C836C6D9508865EB3D1B6" italics="true" pageId="0" pageNumber="89">A. pilosa</emphasis>
</taxonomicName>
H.R.Fletcher is restricted to the subtropical forests found on only Phu Kradeung mountain in north-east Thailand. That this new taxon is currently only found in unlogged areas within the Taman Negeri Kenyir suggests that
<taxonomicName id="CF0C64CC5F1DE42A82CF0A64F2B9FA14" class="Dicotyledones" family="Primulaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="A. recurvipetala" order="Ericales" pageId="0" pageNumber="89" phylum="Angiospermae" rank="species" species="recurvipetala">
<emphasis id="CD8B1B4586C3E3549B397DFCA84D7D5D" italics="true" pageId="0" pageNumber="89">A. recurvipetala</emphasis>
</taxonomicName>
is a useful indicator of primary forest within the forest reserve boundary. The discovery and description of new species is important. It contributes to naming and documenting our local biodiversity as part of a revision of the
<taxonomicName id="87805F6D70ABC474172978E49A08BCF3" class="Dicotyledones" family="Primulaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="" order="Ericales" pageId="0" pageNumber="89" phylum="Angiospermae" rank="family">Primulaceae</taxonomicName>
for the FPM, provides a better understanding of botanical distribution patterns and results in information about forest quality in Peninsular Malaysia.
</paragraph>
</subSubSection>
</treatment>
</document>
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