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<mods:title id= "E842700203858509A4C04C1FA95AB9ED" > Taxonomic and nomenclatural notes on the genera Themus Motschulsky and Lycocerus Gorham (Coleoptera, Cantharidae)</mods:title>
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<mods:namePart id= "52E21657D7E27653FC3D1CB4838DB1E2" > Yang, Yuxia</mods:namePart>
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<mods:namePart id= "0E95E4A57AD8B423AF3E8EA0108E20AD" > Kopetz, Andreas</mods:namePart>
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<mods:namePart id= "66546668AD6B57ED403544BE3BDD569F" > Yang, Xingke</mods:namePart>
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<treatment id= "E2E6CB619FDB2F64B9401EAF495E6EFC" ID-GBIF-Taxon= "152048637" LSID= "urn:lsid:plazi:treatment:E2E6CB619FDB2F64B9401EAF495E6EFC" httpUri= "http://treatment.plazi.org/id/E2E6CB619FDB2F64B9401EAF495E6EFC" lastPageId= "5" lastPageNumber= "6" pageId= "4" pageNumber= "5" scope_family= "Cantharidae" scope_order= "Coleoptera" >
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<paragraph id= "1314D864F1C4EECEBFAF58F1D6D63876" pageId= "4" pageNumber= "5" >
<taxonomicName id= "AD598D146DA559D2AC1C4C85C9ED051F" LSID= "http://species-id.net/wiki/Lycocerus_bilineatus" authority= "Wittmer, 1995" authorityName= "Wittmer" authorityYear= "1995" genus= "Lycocerus" lsidName= "Lycocerus bilineatus" pageId= "4" pageNumber= "5" rank= "species" species= "bilineatus" > Lycocerus bilineatus (Wittmer, 1995)</taxonomicName>
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Figs 7-8
</paragraph>
</subSubSection>
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<subSubSection id= "8765285A7267DE7A5EA5CAFC60D3CA76" pageId= "4" pageNumber= "5" type= "reference_group" >
<paragraph id= "10C96FD1D6C580AACCEDD3457A9A7294" pageId= "4" pageNumber= "5" >
<taxonomicName id= "6D081FEFE9D1D958227C5F71183A8CA2" genus= "Athemus" lsidName= "Athemus (Isathemus) bilineatus" pageId= "4" pageNumber= "5" rank= "species" species= "bilineatus" subGenus= "Isathemus" > Athemus (Isathemus) bilineatus</taxonomicName>
<bibRefCitation id= "E38FF94F278A0BC3E38F679493B54495" author= "Wittmer, W" journalOrPublisher= "Entomologica Basiliensia" pageId= "13" pageNumber= "14" pagination= "171 - 286" title= "Zur Kenntnis Gattung Athemus Lewis (Col. Cantharidae)." volume= "18" year= "1995" > Wittmer 1995</bibRefCitation>
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: 275, figs. 140, 141.
</paragraph>
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<paragraph id= "0EB6AA70F3C4D8B9DA044D121A063F5A" pageId= "4" pageNumber= "5" >
<taxonomicName id= "B6B3FF6711CD26851FFBE1995420E4AC" genus= "Athemus" lsidName= "Athemus" pageId= "4" pageNumber= "5" rank= "genus" > Athemus</taxonomicName>
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(s.str.) amplus
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<bibRefCitation id= "89399F5D90079FF869BF62748E620E7D" author= "Wittmer, W" journalOrPublisher= "Entomologica Basiliensia" pageId= "13" pageNumber= "14" pagination= "171 - 286" title= "Zur Kenntnis Gattung Athemus Lewis (Col. Cantharidae)." volume= "18" year= "1995" > Wittmer 1995</bibRefCitation>
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: 278, figs. 146, 147, 203.
</paragraph>
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<paragraph id= "42D774F00C735774C3CF2529E37EAAFB" pageId= "4" pageNumber= "5" >
<taxonomicName id= "1EFF1E5C77369B203A2CF2742FD41E20" genus= "Lycocerus" lsidName= "Lycocerus amplus" pageId= "4" pageNumber= "5" rank= "species" species= "amplus" > Lycocerus amplus</taxonomicName>
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:
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<bibRefCitation id= "A3C70279B80C67201DCE4C8FD2416FD9" author= "Kazantsev, SV" editor= "Loebl, I" journalOrPublisher= "Apollo Books, Stenstrup" pageId= "12" pageNumber= "13" pagination= "234 - 298" title= "Cantharidae." year= "2007" > Kazantsev and Brancucci 2007</bibRefCitation>
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: 249. syn. n.
</paragraph>
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<paragraph id= "1E61E797DBAF22E7DD531DD37F6493C5" pageId= "4" pageNumber= "5" >
<taxonomicName id= "653D2920AE3C969FA5E48CA7AC515784" genus= "Lycocerus" lsidName= "Lycocerus bilineatus" pageId= "4" pageNumber= "5" rank= "species" species= "bilineatus" > Lycocerus bilineatus</taxonomicName>
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:
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<bibRefCitation id= "B4EEE917E77945A24149F1CB55C4C499" author= "Kazantsev, SV" editor= "Loebl, I" journalOrPublisher= "Apollo Books, Stenstrup" pageId= "12" pageNumber= "13" pagination= "234 - 298" title= "Cantharidae." year= "2007" > Kazantsev and Brancucci 2007</bibRefCitation>
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: 249.
</paragraph>
</subSubSection>
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<subSubSection id= "BCB1DA6C4E269B08E6342DF228B802E4" lastPageId= "5" lastPageNumber= "6" pageId= "4" pageNumber= "5" type= "type material examined" >
<paragraph id= "2FB7811C772E9547EFC1DBAF518008AD" pageId= "4" pageNumber= "5" > Type material examined.</paragraph>
<paragraph id= "701E052C05D8DDEAB1231AB9764AA541" pageId= "5" pageNumber= "6" >
<taxonomicName id= "2898F57691E67A5EDA92F181A220B32D" genus= "Athemus" lsidName= "Athemus (Isathemus) bilineatus" pageId= "5" pageNumber= "6" rank= "species" species= "bilineatus" subGenus= "Isathemus" >
<pageBreakToken id= "E9D0974B98AF82CA245D821E25527EFD" pageId= "5" pageNumber= "6" start= "start" > Athemus</pageBreakToken>
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(Isathemus) bilineatus
</taxonomicName>
: Holotype ♂ (NHMB): [p]" Nordwestal. China \ Chinkiang \ Col. Reitter" , [h]" REM \ 95 \ 11.9" , [h]" A. (Isathemus) \ bilineatus \ Wittm., \ det. W. Wittmer" , [p]
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<normalizedToken id= "33938C7EFAB8B82B320CEC7385615D74" originalValue= "“HOLOTYPUS”" > " HOLOTYPUS" </normalizedToken>
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, [p]" CANTHARIDAE \ CANTH00001927" .
</paragraph>
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<paragraph id= "207E9FE31E218C0435418F8BB05AAF9E" pageId= "5" pageNumber= "6" >
<taxonomicName id= "F92D74ADE2C6B198F8E4103C2E373D4D" genus= "Athemus" lsidName= "Athemus" pageId= "5" pageNumber= "6" rank= "genus" > Athemus</taxonomicName>
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(s.str.) amplus: Holotype ♂ (MNHN): [h]" V.1917 \ China \ Shanghai \ J.
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<normalizedToken id= "D84230855FF806552F3343579B7D7431" originalValue= "Hervé-Bdzin”" > Herve-Bdzin" </normalizedToken>
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, [h]" Athemus s.str. \ amplus \ Wittm. \ det. W. Wittmer" , [p]
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<normalizedToken id= "DAB616BC3CB75D078D46DF158B00D5AB" originalValue= "“HOLOTYPUS”" > " HOLOTYPUS" </normalizedToken>
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. Paratypes: 3♂♂, 2♀♀ (MNHN): [h]
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<normalizedToken id= "A28B6E07199773E4EA6CF6C649B65838" originalValue= "“Shanghai”" > " Shanghai" </normalizedToken>
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, [p]
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<normalizedToken id= "66FB7D82F384822D73084A7B97F51337" originalValue= "“PARATYPUS”" > " PARATYPUS" </normalizedToken>
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; 1♂ (NHMB): [p]" Shanghai \ V.1917 \ J.
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<normalizedToken id= "EE0E2EF71C3430592F936E8386DB13FE" originalValue= "Hervé-Bdzin”" > Herve-Bdzin" </normalizedToken>
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, [p]
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<normalizedToken id= "43368199890B019B8A2723DF7F59A6EB" originalValue= "“PARATYPUS”" > " PARATYPUS" </normalizedToken>
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, " CANTHARIDAE \ CANTH00001340" ; 1♀ (NHMB): [p]" Shanghai \ V \ 1917, J.
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<normalizedToken id= "20E1A59903D3395B9655DD75DDE51530" originalValue= "Hervé-Bdzin”" > Herve-Bdzin" </normalizedToken>
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, [p]
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<normalizedToken id= "35316E7B713B1C32BEC6036DC0A3E421" originalValue= "“PARATYPUS”" > " PARATYPUS" </normalizedToken>
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, [p]" CANTHARIDAE \ CANTH00002072" ; 1♂ (NHMB): [p]" Zi-ka-wei \ 20.IV.1924" , [p]
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<normalizedToken id= "5E34A94576B7E57B60BC3BF943A5E8D4" originalValue= "“PARATYPUS”" > " PARATYPUS" </normalizedToken>
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, [p]" CANTHARIDAE \ CANTH00001986" .
</paragraph>
</subSubSection>
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<subSubSection id= "06B94F6553B06CDFDB8ECAFF23354417" pageId= "5" pageNumber= "6" type= "additional material examined" >
<paragraph id= "C07DE30A9961A62FA0163B3E952DBEF6" pageId= "5" pageNumber= "6" > Additional material examined.</paragraph>
<paragraph id= "2D3D5E9F6AA1AD1AC4FA398AC2397879" pageId= "5" pageNumber= "6" >
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CHINA: Shanghai: 1♂, 2♀♀ (IZAS): " Shanghai, 19.VI.1947, Marist Brothers" ; 1♂ (IZAS):
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<normalizedToken id= "D59DD45F3DCA3545B3B337BD580C0C34" originalValue= "“Zi-ka-wei”" > " Zi-ka-wei" </normalizedToken>
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, 20.IV.1924. Jiangxi: 2♂♂, 1♀ (IZAS): Jiangxi.Hubei: 2♂♂ (IZAS): Xingshan, Xiakou, 140m, 2.V.1994, leg. Xing-Ke Yang; 2♀♀ (IZAS): Zigui, Jiulingtou, 110m, 1.V.1994, leg. Wen-Zhu Li.
</paragraph>
</subSubSection>
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<subSubSection id= "4FB7EAC6885004C9ADBA257C2ADABE05" pageId= "5" pageNumber= "6" type= "distribution" >
<paragraph id= "29F4FF2B0726315F05558DF2B494A2AA" pageId= "5" pageNumber= "6" > Distribution.</paragraph>
<paragraph id= "6C472CC9CC63F37D0F0A9492E9FFDE89" pageId= "5" pageNumber= "6" > China (Jiangsu, Shanghai, Jiangxi, Hubei).</paragraph>
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</subSubSection>
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<subSubSection id= "250339FAE2819ED0DA4489AF9D4897D0" pageId= "5" pageNumber= "6" type= "remarks" >
<paragraph id= "05DEAD608ACB78B98EBB7D797227A24D" pageId= "5" pageNumber= "6" > Remarks.</paragraph>
<paragraph id= "141524A730F04C0445CDC3861450028B" pageId= "5" pageNumber= "6" >
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Both
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<taxonomicName id= "D8351410A5899EB92607A5A81F4DC193" genus= "Lycocerus" lsidName= "Lycocerus amplus" pageId= "5" pageNumber= "6" rank= "species" species= "amplus" > Lycocerus amplus</taxonomicName>
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(Wittmer, 1995) and
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<taxonomicName id= "62A6CBD8449236FDB5E3C321FCD25E2E" genus= "Lycocerus" lsidName= "Lycocerus bilineatus" pageId= "5" pageNumber= "6" rank= "species" species= "bilineatus" > Lycocerus bilineatus</taxonomicName>
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(Wittmer, 1995) were originally described in
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<taxonomicName id= "B9DDE072ECD7DC049ABAD9243E8768CD" genus= "Athemus" lsidName= "Athemus" pageId= "5" pageNumber= "6" rank= "genus" > Athemus</taxonomicName>
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and assigned to different subgenera. In the original manuscript (
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<bibRefCitation id= "874D5292ED56802338EBF1AA384116F8" author= "Wittmer, W" journalOrPublisher= "Entomologica Basiliensia" pageId= "13" pageNumber= "14" pagination= "171 - 286" title= "Zur Kenntnis Gattung Athemus Lewis (Col. Cantharidae)." volume= "18" year= "1995" > Wittmer 1995</bibRefCitation>
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),
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<taxonomicName id= "1804140E4D9140E5EA3835A1DFB1DB57" genus= "Lycocerus" lsidName= "Lycocerus bilineatus" pageId= "5" pageNumber= "6" rank= "species" species= "bilineatus" > Lycocerus bilineatus</taxonomicName>
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was described on a single male type, so it made no sense that it was attributed to the subgenus
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<taxonomicName id= "0604C69615DFEA41BF4DDF66368B4F50" genus= "Isathemus" lsidName= "Isathemus" pageId= "5" pageNumber= "6" rank= "genus" > Isathemus</taxonomicName>
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because of no female available. Furthermore, having examined the holotypes of both nominal species and some paratypes, as well as a series of additional specimens, we could not find any difference justifying their separation, even in the tarsal claws, which is the character to distinguish the subgenera
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<taxonomicName id= "1F45D1377CA0E9E2018A0A9CC416FB33" genus= "Athemus" lsidName= "Athemus" pageId= "5" pageNumber= "6" rank= "genus" > Athemus</taxonomicName>
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and
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<taxonomicName id= "5F7529BD1D75DF7CE9F55E4EE341CA00" genus= "Isathemus" lsidName= "Isathemus" pageId= "5" pageNumber= "6" rank= "genus" > Isathemus</taxonomicName>
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(
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<bibRefCitation id= "116A6EE09F64373D4266CE6FEA1935BF" author= "Wittmer, W" journalOrPublisher= "Entomologica Basiliensia" pageId= "13" pageNumber= "14" pagination= "171 - 286" title= "Zur Kenntnis Gattung Athemus Lewis (Col. Cantharidae)." volume= "18" year= "1995" > Wittmer 1995</bibRefCitation>
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). Consequently, we synonymized
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<taxonomicName id= "6D42304C6EEC7B4EE5E5F1A76FC4796C" genus= "Lycocerus" lsidName= "Lycocerus amplus" pageId= "5" pageNumber= "6" rank= "species" species= "amplus" > Lycocerus amplus</taxonomicName>
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under
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<taxonomicName id= "483F8178711C4967553CA87435E25573" genus= "Lycocerus" lsidName= "Lycocerus bilineatus" pageId= "5" pageNumber= "6" rank= "species" species= "bilineatus" > Lycocerus bilineatus</taxonomicName>
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.
</paragraph>
</subSubSection>
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