<documentID-CLB-Dataset="25374"ID-DOI="http://dx.doi.org/10.3897/nl.42.38341"ID-GBIF-Dataset="ca6f9005-fd01-41a8-85f6-48f0e4484b12"ID-Pensoft-Pub="2367-5365-2-129"ID-Pensoft-UUID="674D3B944E2C5DA5A1B5D05CBEEDA1B6"ID-ZooBank="18830D74393542779EA778C30A8FED48"ModsDocID="2367-5365-2-129"checkinTime="1569654109261"checkinUser="pensoft"docAuthor="Vargas, Hector A."docDate="2019"docId="3799519097FC5A85A8E07DC5D97E1567"docLanguage="en"docName="NotaLepidopt 42(2): 129-136"docOrigin="Nota Lepidopterologica 42 (2)"docSource="http://dx.doi.org/10.3897/nl.42.38341"docTitle="Crocidosema nitsugai Vargas, 2019, sp. nov."docType="treatment"docUuid="ABAD0B64-6794-4666-AE38-6471B431407A"docUuidSource="ZooBank"docVersion="6"id="674D3B944E2C5DA5A1B5D05CBEEDA1B6"lastPageNumber="129"masterDocId="674D3B944E2C5DA5A1B5D05CBEEDA1B6"masterDocTitle="A new species of Crocidosema Zeller (Lepidoptera, Tortricidae) from the Andes of northern Chile"masterLastPageNumber="136"masterPageNumber="129"pageNumber="129"updateTime="1732940382887"updateUser="ExternalLinkService">
<mods:titleid="C3F8560171BD9E1186D3CDA7E5701E87">A new species of Crocidosema Zeller (Lepidoptera, Tortricidae) from the Andes of northern Chile</mods:title>
<figureCitationid="588BDB00BA2AE52E65E2C721CFC31605"captionStart="Figures 1–5"captionStartId="F1"captionText="Figures 1 - 5. Adult stage of Crocidosema nitsugai sp. nov. 1. Holotype male in dorsal view. 2. Male genitalia in ventral view, phallus removed; upper right rectangle showing longitudinal carina on the neck of the right valva. 3. Phallus in lateral view; upper left rectangle showing base of cornuti. 4. Tergum (left) and sternum (right) of male abdominal segment VIII. 5. Female genitalia in ventral view. Scale bars: 2 mm (1), 0.3 mm (2 - 4), 0.25 mm (5)."figureDoi="DOI 10.3897/nl.42.38341.figures1-5"httpUri="https://binary.pensoft.net/fig/342934"pageId="0"pageNumber="129"tableDoi="DOI 10.3897/nl.42.38341.figures1-5">Figs 1-5</figureCitation>
<paragraphid="6E8F4F66E6DE29A4D2FFB6E9AFEC4B01"pageId="0"pageNumber="129">Paratypes, CHILE. One male, two females, same data as holotype, genitalia slides HAV-1261, 1270, 1271 (MNNC); four males, one female, same data as holotype, genitalia slides HAV-1259, 1275, 1276, 1277, 1279 (IDEA).</paragraph>
have a parallel-sided antrum with the dorsal wall posteriorly projected and the posterior margin of sternum VII widely notched at the middle. In contrast, those of
the neck of the valva is almost uniform in height throughout its length with a narrow longitudinal carina, and dorsal and ventral lobes of the cucullus are similar in size. In contrast, in
the neck of the valva is broadened on basal half and lacks carina, and the dorsal lobe of the cucullus is conspicuously smaller than the ventral lobe. The female genitalia of
Razowski & Becker, 2014, described from Carchi, Ecuador. However, the parallel-sided antrum, cingulum longer than the larger signum and two longitudinal stripes along the middle of sternum VII of
<figureCitationid="FFA60A410C6A20FBEF28F1B9C7E20690"captionStart="Figures 1–5"captionStartId="F1"captionText="Figures 1 - 5. Adult stage of Crocidosema nitsugai sp. nov. 1. Holotype male in dorsal view. 2. Male genitalia in ventral view, phallus removed; upper right rectangle showing longitudinal carina on the neck of the right valva. 3. Phallus in lateral view; upper left rectangle showing base of cornuti. 4. Tergum (left) and sternum (right) of male abdominal segment VIII. 5. Female genitalia in ventral view. Scale bars: 2 mm (1), 0.3 mm (2 - 4), 0.25 mm (5)."figureDoi="DOI 10.3897/nl.42.38341.figures1-5"httpUri="https://binary.pensoft.net/fig/342934"pageId="0"pageNumber="129"tableDoi="DOI 10.3897/nl.42.38341.figures1-5">Figs 1-5</figureCitation>
. Vertex and frons mainly whitish gray with a few scattered grayish brown scales. Labial palpus mainly grayish brown with a few scattered whitish gray scales. Antenna whitish gray.
. Mainly yellowish brown dorsally with a few scattered dark gray scales, whitish gray latero-ventrally; patagium grayish brown. Foreleg with anterior face grayish brown, posterior face whitish gray. Midleg similar to foreleg in coloration, tibial spurs whitish gray. Hindleg whitish gray, including tibial spurs. Forewing mainly yellowish brown with abundant whitish gray and grayish brown scales intermixed outside the discal cell, a few scattered dark gray scales near external margin; fringe grayish brown. Hindwing mainly grayish brown, scattered whitish gray scales, fringe whitish gray.
<figureCitationid="7894189E3F16BACA4D1F42EF736E66FC"captionStart="Figures 1–5"captionStartId="F1"captionText="Figures 1 - 5. Adult stage of Crocidosema nitsugai sp. nov. 1. Holotype male in dorsal view. 2. Male genitalia in ventral view, phallus removed; upper right rectangle showing longitudinal carina on the neck of the right valva. 3. Phallus in lateral view; upper left rectangle showing base of cornuti. 4. Tergum (left) and sternum (right) of male abdominal segment VIII. 5. Female genitalia in ventral view. Scale bars: 2 mm (1), 0.3 mm (2 - 4), 0.25 mm (5)."figureDoi="DOI 10.3897/nl.42.38341.figures1-5"httpUri="https://binary.pensoft.net/fig/342934"pageId="0"pageNumber="129"tableDoi="DOI 10.3897/nl.42.38341.figures1-5">Fig. 4</figureCitation>
) somewhat T-shaped; anterior margin straight; lateral margin widely excavated on anterior half; posterior margin about 1/3 the width of anterior margin. Sternum VIII somewhat square-shaped, posterior margin widely excavated in the middle.
<figureCitationid="4CB4F25583BA5D5D5CD1B63BBDDA6EB1"captionStart="Figures 1–5"captionStartId="F1"captionText="Figures 1 - 5. Adult stage of Crocidosema nitsugai sp. nov. 1. Holotype male in dorsal view. 2. Male genitalia in ventral view, phallus removed; upper right rectangle showing longitudinal carina on the neck of the right valva. 3. Phallus in lateral view; upper left rectangle showing base of cornuti. 4. Tergum (left) and sternum (right) of male abdominal segment VIII. 5. Female genitalia in ventral view. Scale bars: 2 mm (1), 0.3 mm (2 - 4), 0.25 mm (5)."figureDoi="DOI 10.3897/nl.42.38341.figures1-5"httpUri="https://binary.pensoft.net/fig/342934"pageId="0"pageNumber="129"tableDoi="DOI 10.3897/nl.42.38341.figures1-5">Figs 2</figureCitation>
<figureCitationid="FA577B5E2E84809109C711270A1C0C0C"captionStart="Figures 1–5"captionStartId="F1"captionText="Figures 1 - 5. Adult stage of Crocidosema nitsugai sp. nov. 1. Holotype male in dorsal view. 2. Male genitalia in ventral view, phallus removed; upper right rectangle showing longitudinal carina on the neck of the right valva. 3. Phallus in lateral view; upper left rectangle showing base of cornuti. 4. Tergum (left) and sternum (right) of male abdominal segment VIII. 5. Female genitalia in ventral view. Scale bars: 2 mm (1), 0.3 mm (2 - 4), 0.25 mm (5)."figureDoi="DOI 10.3897/nl.42.38341.figures1-5"httpUri="https://binary.pensoft.net/fig/342934"pageId="0"pageNumber="129"tableDoi="DOI 10.3897/nl.42.38341.figures1-5">3</figureCitation>
). Tegumen with anterior and posterior margins mainly parallel. Uncus cylindrical, slightly sclerotized, broadened basally, apex rounded, covered with hair-like setae. Socius slightly longer than uncus, narrow, tapering apically. Juxta semicircular, dorsal margin straight. Henion (sclerite between anellus and gnathos) narrow, elongated, well sclerotized, slightly longer than socius. Valvae symmetrical, wide incision on ventral margin; base of valva triangular; sacculus narrow; neck of valva almost uniform in height throughout its length, a narrow longitudinal carina on medial face; cucullus densely covered with hair-like scales, dorsal and ventral lobes similar in size. Phallus cylindrical, slightly longer than sacculus, slightly broadened basally; vesica with several spine-shaped cornuti.
<figureCitationid="1738BBDC7EEC910CD4B98E33C4406ADE"captionStart="Figures 1–5"captionStartId="F1"captionText="Figures 1 - 5. Adult stage of Crocidosema nitsugai sp. nov. 1. Holotype male in dorsal view. 2. Male genitalia in ventral view, phallus removed; upper right rectangle showing longitudinal carina on the neck of the right valva. 3. Phallus in lateral view; upper left rectangle showing base of cornuti. 4. Tergum (left) and sternum (right) of male abdominal segment VIII. 5. Female genitalia in ventral view. Scale bars: 2 mm (1), 0.3 mm (2 - 4), 0.25 mm (5)."figureDoi="DOI 10.3897/nl.42.38341.figures1-5"httpUri="https://binary.pensoft.net/fig/342934"pageId="0"pageNumber="129"tableDoi="DOI 10.3897/nl.42.38341.figures1-5">Fig. 5</figureCitation>
). Papillae analis narrow, elongated, slightly sclerotized, with hair-like setae. Posterior apophysis spine-shaped, slightly broadened basally, about twice length of papillae analis. Anterior apophysis similar to posterior apophysis in shape and length. Tergum VIII with U-shaped notch on anterior margin. Sternum VII with anterior margin straight; lateral margin widely excavated close anterior margin; posterior margin bilobed, widely notched in middle; two mainly parallel longitudinal stripes along the middle diverging close the anterior margin. Antrum parallel-sided, dorsal wall posteriorly projected. Ductus bursae slightly curved, membranous basally and apically; a well-developed cingulum with apex exceeding anterior margin of sternum VII. Corpus bursae membranous, pear-shaped, about 1.5 times length of ductus bursae; two slightly curved saw-like signa laterally.
<figureCitationid="4039193EFC692D749BE358DDF194C18C"captionStart="Figures 6–9"captionStartId="F2"captionText="Figures 6 - 9. Habitat and host plant of Crocidosema nitsugai sp. nov. 6. Habitat of C. nitsugai in the type locality, near Putre, Parinacota Province, at 3670 m elevation on the Andes northern Chile. 7. The host plant Lupinus oreophilus. 8. Inflorescence of L. oreophilus. 9. Fruits of L. oreophilus."figureDoi="DOI 10.3897/nl.42.38341.figures6-9"httpUri="https://binary.pensoft.net/fig/342935"pageId="0"pageNumber="129"tableDoi="DOI 10.3897/nl.42.38341.figures6-9">Fig. 6</figureCitation>
<figureCitationid="011CF325BEBB9C40E432F8203E57CACB"captionStart="Figures 6–9"captionStartId="F2"captionText="Figures 6 - 9. Habitat and host plant of Crocidosema nitsugai sp. nov. 6. Habitat of C. nitsugai in the type locality, near Putre, Parinacota Province, at 3670 m elevation on the Andes northern Chile. 7. The host plant Lupinus oreophilus. 8. Inflorescence of L. oreophilus. 9. Fruits of L. oreophilus."figureDoi="DOI 10.3897/nl.42.38341.figures6-9"httpUri="https://binary.pensoft.net/fig/342935"pageId="0"pageNumber="129"tableDoi="DOI 10.3897/nl.42.38341.figures6-9">Figs 7-9</figureCitation>
