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161 lines
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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.67.6703" ID-PMC="PMC4972245" ID-Pensoft-Pub="1314-2003-67-1" ID-Pensoft-UUID="D846EB3F7746FFFE4A469751FFEF3B22" ID-PubMed="27536193" ID-Zenodo-Dep="133270" ModsDocID="1314-2003-67-1" checkinTime="1469054247116" checkinUser="pensoft" docAuthor="Kriebel, Ricardo" docDate="2016" docId="332AD8C0D91C50DF926E7075CD346608" docLanguage="en" docName="PhytoKeys 67: 1-326" docOrigin="PhytoKeys 67" docPubDate="2016-07-20" docSource="http://dx.doi.org/10.3897/phytokeys.67.6703" docTitle="Conostegia jefensis Kriebel 2016, comb. nov." docType="treatment" docVersion="3" id="D846EB3F7746FFFE4A469751FFEF3B22" lastPageNumber="197" masterDocId="D846EB3F7746FFFE4A469751FFEF3B22" masterDocTitle="A Monograph of Conostegia (Melastomataceae, Miconieae)" masterLastPageNumber="326" masterPageNumber="1" pageNumber="193" updateTime="1668141747722" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A Monograph of Conostegia (Melastomataceae, Miconieae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kriebel, Ricardo</mods:namePart>
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<mods:affiliation>Department of Botany, University of Wisconsin-Madison, 430 Lincoln Drive Madison, Wisconsin 53706, USA</mods:affiliation>
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<mods:nameIdentifier type="email">kriebelr@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2016</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2016-07-20</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>67</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>326</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.67.6703</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.67.6703</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-67-1</mods:identifier>
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<mods:identifier type="Pensoft-UUID">D846EB3F7746FFFE4A469751FFEF3B22</mods:identifier>
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<mods:identifier type="Zenodo-Dep">133270</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="182226951" LSID="urn:lsid:plazi:treatment:332AD8C0D91C50DF926E7075CD346608" httpUri="http://treatment.plazi.org/id/332AD8C0D91C50DF926E7075CD346608" lastPageId="196" lastPageNumber="197" pageId="192" pageNumber="193">
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<subSubSection pageId="192" pageNumber="193" type="nomenclature">
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<paragraph pageId="192" pageNumber="193">
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<taxonomicName LSID="332AD8C0-D91C-50DF-926E-7075CD346608" authority="(Almeda) Kriebel" authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Kriebel" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia jefensis" order="Myrtales" pageId="192" pageNumber="193" phylum="Tracheophyta" rank="species" species="jefensis" status="comb. nov.">Conostegia jefensis (Almeda) Kriebel</taxonomicName>
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<taxonomicNameLabel pageId="192" pageNumber="193">comb. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 172" captionStartId="F172" captionText="Figure 172. Conostegia jefensis. A Branch with infructescence B Abaxial leaf surface C Apex of a branch. Note orange pubescence D Base of the abaxial leaf surface A-B from specimen vouchered R. Kriebel and J. Burke 5659 C-D from specimen vouchered R. Kriebel and J. Burke 5680." figureDoi="10.3897/phytokeys.67.6703.figure172" httpUri="https://binary.pensoft.net/fig/96530" pageId="192" pageNumber="193">Fig. 172</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="194" lastPageNumber="195" pageId="193" pageNumber="194" type="reference_group">
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<paragraph lastPageId="194" lastPageNumber="195" pageId="193" pageNumber="194">
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<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Kriebel" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia jefensis" order="Myrtales" pageId="193" pageNumber="194" phylum="Tracheophyta" rank="species" species="jefensis" status="comb. nov.">
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<pageBreakToken pageId="193" pageNumber="194" start="start">Conostegia</pageBreakToken>
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jefensis
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</taxonomicName>
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(Almeda) Kriebel. Basionym:
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<taxonomicName authorityName="Kriebel" authorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Miconia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Miconia jefensis" order="Myrtales" pageId="193" pageNumber="194" phylum="Tracheophyta" rank="species" species="jefensis" status="comb. nov.">Miconia jefensis</taxonomicName>
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Almeda, Proc. Calif. Acad. Sci. 52(4): 43. 2000. Type: Panama. Cerro Jefe, along summit road and
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<pageBreakToken pageId="194" pageNumber="195" start="start">along</pageBreakToken>
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trail into the Chagres Valley, ca. 900 m, 19 February 1988, F. Almeda, Daniel, T. & G. McPherson 5826 (holotype: CAS!; isotypes: MO!, NY!, PMA!).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="195" lastPageNumber="196" pageId="194" pageNumber="195" type="description">
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<paragraph pageId="194" pageNumber="195">Description.</paragraph>
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<paragraph lastPageId="195" lastPageNumber="196" pageId="194" pageNumber="195">
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Small tree 2-4 m tall with somewhat tetragonal and ridged stems in newer branches that are moderately to densely brown scurfy with stalked stellulate hairs; the nodal line present but inconspicuous. Petiole 0.3-4 cm. Leaf blade 17-43
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<normalizedToken originalValue="×">x</normalizedToken>
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8-14 cm, 5(-7) plinerved, with the innermost pair of veins usually diverging from the midvein 2.5-6 cm above the base in opposite to alternate fashion, elliptic, the base gradually tapering to decurrent on the petiole or less commonly rounded to slightly cordate, the apex acuminate to long acuminate, the margin entire to undulate, the adaxial surface sparsely scurfy-pulverulent to glabrous, the abaxial surface moderately and deciduously scurfy-pulverulent on the secondary and higher order veins. Inflorescence a terminal panicle 4.4-11.2 cm long branched at the base, accessory branches absent, brown scurfy on the branches, subulate, bracteoles 0.3-0.75
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<normalizedToken originalValue="×">x</normalizedToken>
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0.2-0.5 mm, persistent, fused and forming a nodal collar or ridge. Pedicel 0.5-2 mm. 1.25-1.75
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<normalizedToken originalValue="×">x</normalizedToken>
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1.25-1.75 mm, hypanthium campanulate, not constricted, deciduously scurfy-pulverulent. Flowers 5-merous, not calyptrate, flower buds ca. 0.3-0.5
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<normalizedToken originalValue="×">x</normalizedToken>
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0.1-0.2 mm, calyx tube 0.2-0.3 mm long, calyx lobes depressed-triangular to undulate, 0.5 mm long but concealed and barely exceeded by the exterior subulate teeth, androecial fringe present and beset with glandular hairs. Petals 2.75-3.25
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<normalizedToken originalValue="×">x</normalizedToken>
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0.75-1 mm, magenta, linear-oblong, spread
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<pageBreakToken pageId="195" pageNumber="196" start="start">ing</pageBreakToken>
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an anthesis, the apex acute, glabrous, spreading an anthesis, the margin entire. Stamens 10, 2.25-2.75 mm, radially arranged around the style, the filaments 1.25-1.5 mm long, with a geniculation near the apex, white, anthers 1-1.25 0.25-0.4 mm, oblong, yellow, the pore ca. 0.1 mm wide, somewhat dorsally inclined. Ovary 5 locular, 3/4 to 4/5 inferior, glabrous apically. Style 3.5-4.25 mm long, curving towards the apex, distance from the anther pore to the stigma 1-1.5 mm, stigma truncate, ca. 0.35 mm wide. Berry 3-4
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<normalizedToken originalValue="×">x</normalizedToken>
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3-4 mm, purple-black. Seeds 0.35-0.5 mm, pyramidate, smooth with verruculose angles.
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</paragraph>
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<caption ID-Zenodo-Dep="973999" doi="10.3897/phytokeys.67.6703.figure172" httpUri="https://binary.pensoft.net/fig/96530" pageId="195" pageNumber="196" start="Figure 172" startId="F172">
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<paragraph pageId="195" pageNumber="196">
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<emphasis bold="true" pageId="195" pageNumber="196">Figure 172.</emphasis>
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<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Kriebel" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia jefensis" order="Myrtales" pageId="195" pageNumber="196" phylum="Tracheophyta" rank="species" species="jefensis">
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<emphasis italics="true" pageId="195" pageNumber="196">Conostegia jefensis</emphasis>
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</taxonomicName>
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.
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<emphasis bold="true" pageId="195" pageNumber="196">A</emphasis>
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Branch with infructescence
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<emphasis bold="true" pageId="195" pageNumber="196">B</emphasis>
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Abaxial leaf surface
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<emphasis bold="true" pageId="195" pageNumber="196">C</emphasis>
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Apex of a branch. Note orange pubescence
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<emphasis bold="true" pageId="195" pageNumber="196">D</emphasis>
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Base of the abaxial leaf surface
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<emphasis bold="true" pageId="195" pageNumber="196">A-B</emphasis>
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from specimen vouchered
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<emphasis italics="true" pageId="195" pageNumber="196">R. Kriebel and J. Burke 5659</emphasis>
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<emphasis bold="true" pageId="195" pageNumber="196">C-D</emphasis>
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from specimen vouchered
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<emphasis italics="true" pageId="195" pageNumber="196">R. Kriebel and J. Burke 5680</emphasis>
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.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection lastPageId="196" lastPageNumber="197" pageId="195" pageNumber="196" type="distribution">
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<paragraph pageId="195" pageNumber="196">Distribution</paragraph>
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<paragraph pageId="195" pageNumber="196">
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(Fig.
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<figureCitation captionStart="Figure 173" captionStartId="F173" captionText="Figure 173. Distribution of Conostegia jefensis." figureDoi="10.3897/phytokeys.67.6703.figure173" httpUri="https://binary.pensoft.net/fig/96531" pageId="195" pageNumber="196">173</figureCitation>
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). Endemic to Cerro Jefe in Panama from 750-1000 m elevation.
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</paragraph>
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<caption ID-Zenodo-Dep="974001" doi="10.3897/phytokeys.67.6703.figure173" httpUri="https://binary.pensoft.net/fig/96531" pageId="195" pageNumber="196" start="Figure 173" startId="F173">
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<paragraph pageId="195" pageNumber="196">
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<emphasis bold="true" pageId="195" pageNumber="196">Figure 173.</emphasis>
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Distribution of
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<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Kriebel" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia jefensis" order="Myrtales" pageId="195" pageNumber="196" phylum="Tracheophyta" rank="species" species="jefensis">
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<emphasis italics="true" pageId="195" pageNumber="196">Conostegia jefensis</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</caption>
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<paragraph lastPageId="196" lastPageNumber="197" pageId="195" pageNumber="196">
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<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Kriebel" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia jefensis" order="Myrtales" pageId="195" pageNumber="196" phylum="Tracheophyta" rank="species" species="jefensis">
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<emphasis italics="true" pageId="195" pageNumber="196">Conostegia jefensis</emphasis>
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</taxonomicName>
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can be recognized by its reddish scurfy pubescence, strongly plinerved leaves and magenta petals. This is a puzzling species because it includes two morphotypes endemic to Cerro Jefe. The common morphotype resembles
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<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Gleason" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia consimilis" order="Myrtales" pageId="195" pageNumber="196" phylum="Tracheophyta" rank="species" species="consimilis">
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<emphasis italics="true" pageId="195" pageNumber="196">Conostegia consimilis</emphasis>
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</taxonomicName>
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in the decurrent leaf base but differs from it in its coarse habit, somewhat coriaceous leaves and magenta petals. The second morphotype which corresponds to the mor
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<pageBreakToken pageId="196" pageNumber="197" start="start">phology</pageBreakToken>
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of the holotype is apparently much less common and does not have the decurrent leaf base but rather a rounded to cordate leaf base. This morphotype also has the typical magenta petals of the other morphotype. Unexpectedly, the two morphotypes did not form a clade in the molecular phylogeny. The holotype matching morphotype formed a strong sister relationship to another Cerro Jefe endemic,
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<taxonomicName authorityName="Kriebel" authorityYear="2016" baseAuthorityName="Kriebel" baseAuthorityYear="2016" class="Magnoliopsida" family="Melastomataceae" genus="Conostegia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Conostegia peltata" order="Myrtales" pageId="196" pageNumber="197" phylum="Tracheophyta" rank="species" species="peltata">
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<emphasis italics="true" pageId="196" pageNumber="197">Conostegia peltata</emphasis>
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</taxonomicName>
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. Lack of resolution and more samples of each morphotype are needed at this time to further confirm the possibility that they represent different taxa.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="196" pageNumber="197" type="materials_examined">
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<paragraph pageId="196" pageNumber="197">Specimens examined.</paragraph>
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<paragraph pageId="196" pageNumber="197">
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<emphasis bold="true" pageId="196" pageNumber="197">
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<collectingCountry name="Panama">PANAMA</collectingCountry>
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.
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<normalizedToken originalValue="Panamá">
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<collectingRegion country="Panama" name="Panama">Panama</collectingRegion>
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</normalizedToken>
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</emphasis>
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: Una milla
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<normalizedToken originalValue="después">despues</normalizedToken>
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de la Eneida,
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<normalizedToken originalValue="Región">Region</normalizedToken>
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de Cerro Jefe, Correa and Dressler 946 (MO, NY, PMA); P. N. Chagres, sendero hacia Cerro Jefe que sale de la
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<normalizedToken originalValue="urbanización">urbanizacion</normalizedToken>
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Altos de Cerro Azul, Kriebel and Burke 5659, 5669 (PMA, NY).
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</paragraph>
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</subSubSection>
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</treatment>
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</document>
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